A phytosociological study covering approximately 12 000 km2 was completed within Southeastern Mpumalanga high altitude grasslands. The towns of Belfast, Barberton, Piet Retief and Wakkerstroom represented the four corners of the area. The study formed part of the Grassland Biome Project sponsored by the Department of Environmental Affairs and Tourism. The Braun-Blanquet approach was applied. Grassland research results from the western side of the country, determined that soil patterns and vegetation distribution are significantly correlated. Based on these results, it was hypothesised that the same correlation would exist in the east. The eight soil patterns used during the survey as homogenous units, were Ab, Ac, Ad, Ae, Ea, Fa, Fb and lb. A pro rata, randomly stratified sample size of 405 plots were used. An in-depth literature study of the environmental and other factors related to vegetation distribution, indicated on local scale that a significant correlation exists between landform and plant distribution. Two geographic information systems, Idrisi and Arc-Info/Niew, aided in the modelling and extraction of environmental attributes from existing databases. A TWINSPAN classification of the complete floristic data set falsified the null hypothesis based on the soil patterns but verified the null hypothesis based on landforms. The TWINSPAN dendrogram revealed clusters associated with three landforms (mountains, hills and lowlands, and plains) in two ecosystems, the terrestrial and the wetlands or aquatic. The three landforms represented three mapping units: Southeastern Mpumalanga Mountain Vegetation Type, Southeastern Mpumalanga Hills and Lowland Vegetation Type and Southeastern Mpumalanga Plain Vegetation Type. A second TWINSPAN classification was executed on each of the vegetation types. The resulting clusters were tested for uniqueness and informational value using a set of criteria. Those clusters which qualified were arranged in a Braun-Blanquet table to determine communities, sub-communities and variations using constancy and fidelity values. The plant communities within the vegetation types were described in terms of floristic composition and environmental attributes. The indirect gradient analysis ordination program DECORANA was used to determine environmental trends and was confirmed by using multiple regression. Soil water availability was the most significant environmental trend between the two ecosystems and within the communities of the ecosystems in each vegetation type. Soil water availability is influenced by numerous factors, whose significance differs from one vegetation type to another. lt was concluded from this study that, in high rainfall areas, soil patterns and vegetation distribution are not significantly correlated but that landforms and vegetation distribution are significantly correlated. Landscape slope configuration or landform influences soil water availability through soil characteristics (rockiness, texture and depth) and local climatic conditions (aspect, perpendicular insolation and precipitation). lt is suggested that the current Grassland - Savanna Biome boundary be changed towards the west using the boundary between the covered and exposed shields. This suggestion is supported by the presence of Hyparrhenia hirta/Hyperthelia dissoluta - Acacia sieberiana communities to the east of the study area in the different vegetation types which correlate significantly with the thorntree-tall grass savanna formation on a global scale. AFRIKAANS : 'n Fitososiologiese studie is gedoen van die Suid-oostelike Mpumalanga hoe liggende grasveld tussen die dorpe Belfast, Barberton, Piet Retief en Wakkerstroom. 'n Area van ongeveer 12 000 km2 is bestudeer. Die studie is deel van die Grasveld Bioom Projek gefinansier deur die Departement van Omgewingsake en Toerisme. Die Braun-Blanquet benadering is gebruik. Resultate van grasveldnavorsing in die westelike deel van die grasveld bioom, het aangedui dat 'n betekenisvolle verband tussen grondpatrone en plantegroei verspreiding bestaan. Na aanleiding van hierdie resultate, is die hipotese gemaak dat dieselfde verband in die ooste sal voorkom. Agt grondpatrone, Ab, Ac, Ad, Ae, Fa, Fb, en lb is as homogene eenhede beskou. 'n Eweredige ewekansige steekproefneming van 405 punte is gebruik. 'n Deeglike literatuurstudie van die omgewing en faktore wat verband hou met plantegroei verspreiding, het aangedui op lokale skale, dat 'n verband tussen landvorm en plantegroei verspreiding bestaan. Twee geografiese inligtingstelsels, ldrisi en Arc-lnfo/View, is gebruik in die modelering en verkryging van omgewings data vanuit bestaande databasisse. 'n TWINSPAN klassifikasie gebaseer op die volledige spesiesamestelling, het die nul hipotese oor die grondpatrone ongeldig verklaar maar die nul hipotese oor die landvorme bevestig. Die TWINSPAN dendrogram het groeperings bevat wat verband hou met drie landvorme (berge, heuwels en laaglande en vlaktes) in twee ekosisteme, terrestrieel en vleiland van akwatiese omgewings. Die drie landvorme is beskou as drie karteerbare eenhede: Suid-oostelike Mpumalanga Berg Plantegroeitipe, Suidoostelike Mpumalanga Heuwels en Laagland Plantegroeitipe en Suid-oostelike Mpumalanga Vlakte Plantegroeitipe. 'n Tweede TWINSPAN klassifikasie is op elk van die plantegroeitipes toegepas. Elke plantegroeitipe se groeperings is met behulp van bepaalde voorwaardes vir uniekheid en inligtingswaarde getoets. Die groeperings wat gekwalifiseer het, is met behulp van konstantheids en getrouheids waardes in 'n Braun-Blanquet tabel gerangskik in gemeenskappe, sub-gemeenskappe en variasies. Elke plantegroei tipe se gemeenskappe is beskryf in terme van floristies spesiesamestelling en omgewingsfaktore. 'n indirekte gradientanalise ordeningsprogram DECORANA is gebruik vir die bepaling van omgewings tendense en is bevestig met behulp van veelvuldige regressies. Grondwaterbeskikbaarheid was die mees betekenisvolle omgewingstendens tussen die ekosisteme en die gemeenskappe van die plantegroei tipes. Grondwaterbeskikbaarheid word deur 'n aantal faktore beïnvloed, waarvan die belangrikheid van plantegroei tipe tot plantegroeitipe wissel. 'n Gevolgtrekking van die studie is, dat in hoë reënvalomgewings daar nie 'n betekenisvolle verband is tussen grondpatrone en plantegroei verspreiding nie maar wel tussen landvorme en plantegroeiverspreiding. Landskap hellingsamestelling of landvorme beïnvloed grondwaterbeskikbaarheid deur middel van grondeienskappe (klipperigheid, tekstuur en diepte) en lokale klimaatstoestande (aspek, loodregte bestraling en presipitasie). Daar word voorgestel dat die huidige Grasveld- Savanna Bioom grens weswaarts geskuif word om ooreen te stem met die grens tussen die bedekte en blootgestelde plate. Die voorstel word ondersteun deur die teenwoordigheid van Hyparrhenia hirta/Hyperthelia dissoluta - Acacia sieberana plantgemeenskappe in die ooste van die plantegroeitipes wat betekenisvol ooreenstem met die langgras-doring boom savanna formasie op globale skaal. / Dissertation (MSc)--University of Pretoria, 1999. / Plant Science / unrestricted

Hieracium pilosella is an invasive weed of New Zealand's short tussock grasslands. Since the 1960s, the abundance of H. pilosella has dramatically increased; it is now thought to occur in 6 million hectares of New Zealand (Espie, 2001), predominantly in grasslands. It is at least common in 42% of this area (Espie, 2001). Ecology is inherently spatial and as plants closely interact with their direct neighbours, the spatial arrangement of plants is vital to their functioning. A handful of recently published articles have implicated spatial structure of plant communities in theories of plant competition, resource use and the invasion of plant communities. The aims of this thesis were to: 1) determine if there are consistent spatial patterns in New Zealand's short tussock grasslands at relatively small scales (i.e. spatial relationships between individuals); 2) investigate how the invasion of H. pilosella may be altering these spatial patterns; and 3) establish if the spatial patterns of species, life-forms and root systems are being altered in different ways. Spatial patterns of both tussock and inter-tussock species, life-forms and root functional groups were evaluated at a range of short tussock grassland sites across a gradient of H. pilosella invasion levels in Canterbury, using both join-count statistics and Ripley's K-function. A classification system for the root functional groups of vascular species in these communities was developed and applied. It was found that species, life-forms and root functional groups in short tussock grasslands had generally consistent spatial patterns across sites both within and between species. These patterns were variable between significantly different levels of H. pilosella ground cover. The type of spatial pattern exhibited, and the way it was altered differed between species, life-form and root functional groups. For example, tussocks exhibited increased regularity up to scales of 160 cm and increasing aggregation at scales up to 500 cm, with increases in H. pilosella abundance. In contrast, both Agrostis capillaris and herbaceous chamaephytes had increased aggregation across scales up to 160 cm. These differences in spatial patterns along the gradient of invasion are a strong indication that H. pilosella is structurally fragmenting New Zealand's short tussock grasslands. This fragmentation is likely to have far reaching effects including the disturbance of invertebrate communities and the disruption of ecosystem services including pollination, vegetation regeneration, and nutrient cycling.

This study set out to investigate the effects of level of nitrogen supply on the root dynamics, carbon (C) and nitrogen (N) return to soil from root turnover, decomposition, and stability of improved grasslands. Field studies involved a two-year field experiment, where plant species composition, above ground biomass production, root production, litter decomposition and short term soil organic matter (SOM) dynamics, as a response to varying levels of inorganic fertiliser nitrogen supply were studied. The results showed that root dynamics are strongly affected by N fertilisation, with an increase in root production and death with increasing N supply. Reduced water availability lead to a greater root disappearance rate and the production of roots with a shorter lifespan. Root decomposition was strongly influenced by the age of the grass ley, which also exerted a strong influence on the structure of the soil microbial communities (SMC). Rate of litter decomposition and SMC structure were also influenced by the level of N supply. Level of mineral N supply and age of the grass ley also influenced the formation and stability of water stable aggregates, the microbial community structure and microbial community function within the aggregates. Mineral N applications are likely to influence SOM and soil nutrient dynamics. The finding that climatic conditions, and in particular water availability, had the strongest impact on both above and below ground productivity, and strongly influenced the amount of C, N and root/shoot biomass returned to soils, is of great importance in helping to make more accurate predictions of the response of plant communities to projected changes in the global climate.

Doctor of Philosophy / Division of Biology / Jesse B. Nippert / Water availability is a key driver of many plant and ecosystem processes in tallgrass prairies, yet we have a limited understanding of how grassland plants utilize water through space and time. Considering that tallgrass prairies experience tremendous heterogeneity in soil resources, identifying spatiotemporal variation in plant ecohydrology is critical for understanding current drivers of plant responses to water and for predicting ecosystem responses to future changes in climate. Here, I investigated the patterns, drivers, and ecological consequences of plant water use (e.g., water uptake, water redistribution, and water loss) in a native tallgrass prairie located in northeastern Kansas, USA. Using a combination of leaf gas exchange, sap flow, and isotopic techniques, I addressed four main questions: 1) How does fire and grazing by bison impact use of water from different sources and niche overlap for common grasses, forbs, and shrubs? 2) Does hydraulic lift occur in grazed and ungrazed tallgrass prairie, and does this impact facilitation for water within grassland communities? 3) What are the patterns and drivers of nocturnal transpiration in common grassland species? 4) How does diel stem sap flow and canopy transpiration vary among common grassland species? I found that bison grazing increased the depth of water uptake by Andropogon gerardii and Rhus glabra, reducing niche overlap with co-occurring species. Conversely, grazing did not affect hydraulic lift, which was generally uncommon and likely limited by nocturnal transpiration. Further, leaf gas exchange measurements indicated that nocturnal transpiration occurred commonly in tallgrass prairie plants and was greatest among grasses and early in the growing season. Nocturnal transpiration was not driven by vapor pressure deficit or soil moisture, as commonly observed in other systems, but was regulated by nocturnal stomatal conductance in most species. Finally, I found that daytime sap flow rates were variable among species and functional types, with larger flux rates among woody species. Nocturnal sap flow rates were more consistent across species, which caused nighttime sap flow and transpiration to account for a larger proportion of daily flux rates in grasses than in forbs or shrubs. These results show that water uptake, water redistribution, and water loss are all influenced by different biotic and abiotic drivers and have varying ecological impacts across a heterogeneous landscape. Additionally, extensive differences in water flux exist among co-occurring species and plant functional groups, which likely reflect varying strategies to tolerate water limitation. These results suggest that shifts in the abundance of these species with future climate changes, or with ecosystem state changes, will likely impact ecosystem-level water balance.

No description available.

Thesis (M.S.)--Oregon State University, 2008. / Printout. Includes bibliographical references (leaves 86-93). Also available on the World Wide Web.

No description available.

This research has examined the historical vegetation of two Eastern Otago sites below the regional treeline, with the aim of addressing questions about the distribution and spread of native tussock grasslands prior to human arrival in New Zealand c. 800 yr BP. Pollen and phytolith (plant opaline silicate) proxies have been extracted from peat cores at Swampy Summit and Clarks Junction to provide a record of vegetation spanning the Holocene. Using multiple proxies and two sample sites has allowed for comparisons of the record of vegetation from within sites and between sites. A record of the modern pollen rain was also gathered from localised moss polsters at Swampy Summit in order to reconcile modern pollen assemblages and transport patterns with historical findings. It became clear from the research that the record of vegetation inferred from phytoliths was not analogous to the pollen-based records, which supported the hypothesis that vegetation reconstructions based solely on pollen may be unreliable. Good pollen preservation in the sediments allowed for the identification of over 50 taxa, although only Chionochloid forms were identifiable to a family level in the phytolith records. Poaceae pollen was abundant throughout the Clarks Junction record, suggesting grassland had persisted at this site during the Holocene, while Poaceae pollen at Swampy Summit was minimal and sporadic. Phytoliths at Swampy Summit show grasses have persisted at the site thoughout the Holocene, at times in much greater proportions than the pollen record would suggest, while Chionochloid phytoliths only become common near the top of the record, possibly reflecting increasing dominance of this taxa after human disturbance. In contrast to Swampy Summit, the Clarks Junction phytolith record reflects a more stable presence of grasses throughout the Holocene, with Chionochloid forms present throughout. Phytoliths appear to be a more reliable proxy for local vegetation, with both sites indicating a Holocene presence of grasses below the regional treeline prior to human arrival in New Zealand. In comparison, the pollen record appears to indicate a more regional pattern of vegetation, with the grassland pollen record complicated by pollen dispersal and deposition factors.

Grasslands are the most important ecosystem to humanity, as they are responsible for feeding that majority of the human population. These are also very large ecosystems; they cover approximately 40% of the surface of the earth (Loveland et al., 1998), making ground-based surveys for monitoring grassland health and productivity extremely time consuming. Remote sensing has the advantage of providing reliable and repeatable observations over large swaths of land; however, optical sensors exploiting the visible and near infrared regions of electromagnetic (EM) spectrum will be unable to collect information from the ground if clouds are present (Wang et al., 2009). Imaging radar sensors, the most common being synthetic aperture radar (SAR), have the advantage of being able to image the ground even during cloudy conditions. The longer wavelengths of EM energy used by the SAR sensor are able to penetrate clouds while shorter wavelength used by optical sensors are scattered. A grassland monitoring tool based on SAR imagery would have many advantages over an optical imagery system, especially when SAR data becomes widely available. To demonstrate the feasibility of grassland monitoring using SAR, this study experimented with a set of dual-polarimetric SAR imagery to extract several grassland biophysical parameters such as soil moisture, canopy moisture, and green grass biomass over the mixed grassland in southwestern Saskatchewan. Soil moisture was derived from these images using the simple Delta Index (Thoma et al., 2006) first developed for a sparsely vegetated landscape. The Delta Index was found to explain 80% of the variation in soil moisture, in this vegetated landscape. Canopy moisture was modeled using the water cloud model (Attema and Ulaby, 1978). This model has a similar explanatory power of R2 = 0.80. This study found that only the photosynthesizing green grass biomass had a significant relationship with the canopy moisture model. However, only about 40% of the variation in green grass biomass can be explained by canopy moisture alone. The cross-polarized ratio developed from the dual polarimetric images was found to reflect the plant form diversity of the grassland. Biophysical parameters extracted from optical satellite imagery, Landsat-5 in the case of this study, were compared to those derived from the SAR images. This comparison revealed that the SAR images were superior in sensitivity to soil and canopy moisture. Optical imagery was found to be more sensitive to green canopy cover. An approach combining the results from both sensors showed an improvement in green grass biomass estimation (Adjusted R2 = 0.71).

Hieracium pilosella is an invasive weed of New Zealand's short tussock grasslands. Since the 1960s, the abundance of H. pilosella has dramatically increased; it is now thought to occur in 6 million hectares of New Zealand (Espie, 2001), predominantly in grasslands. It is at least common in 42% of this area (Espie, 2001). Ecology is inherently spatial and as plants closely interact with their direct neighbours, the spatial arrangement of plants is vital to their functioning. A handful of recently published articles have implicated spatial structure of plant communities in theories of plant competition, resource use and the invasion of plant communities. The aims of this thesis were to: 1) determine if there are consistent spatial patterns in New Zealand's short tussock grasslands at relatively small scales (i.e. spatial relationships between individuals); 2) investigate how the invasion of H. pilosella may be altering these spatial patterns; and 3) establish if the spatial patterns of species, life-forms and root systems are being altered in different ways. Spatial patterns of both tussock and inter-tussock species, life-forms and root functional groups were evaluated at a range of short tussock grassland sites across a gradient of H. pilosella invasion levels in Canterbury, using both join-count statistics and Ripley's K-function. A classification system for the root functional groups of vascular species in these communities was developed and applied. It was found that species, life-forms and root functional groups in short tussock grasslands had generally consistent spatial patterns across sites both within and between species. These patterns were variable between significantly different levels of H. pilosella ground cover. The type of spatial pattern exhibited, and the way it was altered differed between species, life-form and root functional groups. For example, tussocks exhibited increased regularity up to scales of 160 cm and increasing aggregation at scales up to 500 cm, with increases in H. pilosella abundance. In contrast, both Agrostis capillaris and herbaceous chamaephytes had increased aggregation across scales up to 160 cm. These differences in spatial patterns along the gradient of invasion are a strong indication that H. pilosella is structurally fragmenting New Zealand's short tussock grasslands. This fragmentation is likely to have far reaching effects including the disturbance of invertebrate communities and the disruption of ecosystem services including pollination, vegetation regeneration, and nutrient cycling.