This study was undertaken to determine the identity of the causative agents of the filariases reported from northwestern tetraonids (Fowle, 1946; Babero, 1953), the incidence of filarial infections in the Tetraonidae of British Columbia, and the means by which these birds acquire their infections. More than 400 wild gallinaceous birds of 10 species and over 200 adult non-gallinaceous birds from 27 families were examined. Adults of the following 4 species of filariae (Onchocercidae: Splendidofilariinae) were collected from the Tetraonidae of B.C. and/or Alaska and are described herein: 1) Skriabinocta flexivaqinalis (Jones, 1961) n. comb.; 2) Splendidofilaria pectoralis new species; 3) Splendidofilaria "species A"; and 4) Splendidof il aria papil locerca (Lubimov, 1946;) Anderson and Chabaud, 1959. Descriptions are provided also for the microfilariae of S. flexivaqinalis. S. pectoralis and S. papillocerca, and for Microfilaria laqopodis (Haaland, 1928),Brinkmann, 1950, and Microfilaria "species B" from tetraonid hosts. Recent systems of filarioid classification are evaluated and some modifications proposed, including revival of the genus Skriabinocta Chertkova, 1946 to comprise S. petrowi Chertkova, 1946, S. flexivaginalis. S. chitwoodae (Anderson, 1961) n. comb,, S. striatospicula (Hibler, 1964) n. comb., and provisionally S. lienalis (Orloff, 1947) n. comb. On Vancouver Island, Microfilaria sp. B is hyperenzootic in blue grouse, with a significantly higher prevalence in adult males than in adult hens or in yearlings or chicks. Mf. sp. B is enzootic in Vancouver Island ruffed grouse. S. flexivaginalis is sporadic in British Columbia ruffed grouse. Splendidofilaria pectoralis is sporadic to enzootic in tetraonids from central B. C. and Alaska. Mf. lagopodis is.enzootic in willow ptarmigan of northern B. C. Microfilariae closely resembling Mf. flexivaginalis occasionally parasitize non-gallinaceous birds of Vancouver Island, but Mf. sp. B and S. pectoralis seem to be restricted totetraonid hosts. The larvae of S. flexivaginalis develop in the thoracic muscles of the ceratopogonid, Culicoides unicolor (Coq.) group. Those of Mf. sp. B develop in abdominal fat bodies of the simuliids, Simulium aureum Fries and Cnephia minus (D. and S.); the former being considerably more efficient as an intermediate host. Development of Mf. sp. B fails at temperatures which do not range above 62°F. Larval development is described and on the basis of larval characters some features of the adults of Mf. sp. B are predicted. Knowledge acquired on the biology of the tetraonid hosts, the incidence of the filariases, and the seasonal abundance of the vectors and the effects of various factors on their feeding activities, is integrated in a discussion of the epizootiology of Skri abinoct a flexivaqinalis and Microfilaria sp. B. / Science, Faculty of / Zoology, Department of / Graduate

The greater sage-grouse (Centrocercus urophasianus; sage-grouse) has been a species of conservation concern since the early 20th century due to range-wide population declines. To contribute to knowledge of the ecology of sage-grouse populations that inhabit the Box Elder Sage Grouse Management Area (SGMA) in northwestern Utah and quantify their responses to landscape scale habitat manipulations, I monitored vital rates and habitat selection of 45 female sage-grouse from 2014 to 2015. Using telemetry locations of female sage-grouse with known nest and brood fates, I created Generalized Linear Mixed Models to estimate the influence of proximity to pinyon (Pinus spp.) and juniper (Juniperus spp.; conifer) encroachment, and removal projects may have on sagegrouse reproductive fitness in the Box Elder SGMA. The best fit model suggested that for every 1 km a nest was located away from a conifer removal area, probability of nest success was reduced by 9.1% (β = -0.096, P < 0.05). Similarly, for every 1 unit increase in the log-odds of selection for distance to treatment, probability of brood success declined by 52.6% (P = 0.09). The probability of brood success declined by 77.2% (P < 0.05) as selection for conifer canopy cover increased. To evaluate sage-grouse habitat use, I used fecal pellet surveys to estimate relative pellet density in conifer encroachment, removal, and undisturbed sagebrush areas. Sage-grouse pellet densities were estimated at 4.6 pellets/ha (95% CI = 1.2, 10.9), 8.6 pellets/ha (95% CI = 3.8, 15.2), and 50.6 pellets/ha (95% CI = 36.8, 69.6), in conifer encroachment, removal, and undisturbed sagebrush areas respectively. Density estimates did not statistically differ between conifer encroachment and removal areas. To determine if vegetation micro-site characteristics at sage-grouse use sites influenced nest or brood fate, I recorded standard vegetation measurements for all radio-marked sage-grouse nests and a stratified random sample of brood-use sites from 2014- 2015 and compared them to random sites. Micro-site vegetation characteristics measured did not differ for successful and unsuccessful nests. Many characteristics differed between micro-sites used by successful broods and those used by unsuccessful broods. Sites used by successful broods also differed from random sites.

The ruffed grouse (Bonasa umbellus incana) inhabits much of the deciduous woodland of northern and central Utah. Its distribution is confined mainly to the Wasatch and Uinta mountain ranges. It has been observed in willows (Salix spp.) at 5, 000 feet elevation and in Douglas-fir (Pseudotsuga menziesii) at 9,000 feet.

Greater sage-grouse (Centrocercus urophasianus) populations have declined throughout the western United States over the past 3 decades. Habitat loss within the sagebrush steppe ecosystem is a major factor leading to sage-grouse population decline. Hen sage-grouse were captured, marked, and tracked during the summer of 2012 in southwestern and south-central Wyoming. I performed vegetation surveys, and avian point counts were performed at 1 early-season brood location, 1 late-season brood location, and an accompanying random location for each marked hen regardless of reproductive status. Multinomial models were run to determine what habitat variables were most informative in predicting site selection by hen sage-grouse. During early-brood season, hen sage-grouse with chicks selected sites that had high total shrub cover density; these areas also exhibited high densities of American kestrels (Falco sparverius). They did not avoid areas with common ravens (Corvus corax). Hen sage-grouse not accompanied by a brood selected sites with high total shrub cover and low densities of common ravens and American kestrels. During late-brood season, hen sage-grouse that were accompanied by a brood selected sites with high shrub cover and low densities of small avian predators, such as black-billed magpies (Pica hudsonia) and American kestrels as well as medium-sized predators, such as common ravens, buteo hawks (Buteo spp.), and northern harriers (Circus cyaneus). Hens that were not accompanied by broods were more often found in sites with high total shrub cover and low densities of small avian predators, but selected sites with higher densities of medium-sized predators. Hen sage-grouse select areas with high total shrub cover during early and late-brood season regardless of their reproductive status. By avoiding predators and selecting areas with cover, hens with broods can reduce the risk of their chicks being depredated.

The particular purpose of this study was to determine the effects of large-scale sagebrush reseeding projects on sage grouse populations and whether such effects were partly or entirely beneficial, neutral, or detrimental to the survival of such populations. During two seasons of field work, the studies initiated on a short-time basis had the following specific objectives: To compare the utilization by sage grouse of reseeded and non-reseeded lands for the seasonal activities of mating, nesting, raising a brood, fall coveying, and wintering. To compare the utilization by sage grouse of reseeded and non-reseeded lands for daily activities of feeding, watering, resting, hiding, and roosting. To determine fall and winter movements of the grouse in relation to reseeded lands. To determine the food and cover available to grouse on randomly selected sample plots To arrive at an index to food preferences through comparison of stomach analysis and food availability studies. To determine the effect of livestock grazing of reseeded lands on sage grouse. To determine the effect of plant succession on availability of food and cover within reseeded lands.

No description available.

Feeding trials.showed that ruffed grouse (Bonasa umbellus) performed well on diets containing 20% Christmas hollyfem (Polystichum acrostichoides) or mountain laurel (Kalmia latifolia), but diets containing 40% of these forages resulted in lower protein and energy intake and the Christmas hollyfem diet caused a loss of body mass. Grouse were not able to maintain themselves solely on evergreen leaves. Glucuronide excretion was greatest for the 40% mountain laurel diet. Ornithine conjugate excretion was greatest for a diet with 40% deciduous leaves. Sulfate excretion did not vary among diets. Intake rate of leaves was an asymptotic function of bite size when the density of bites did not limit intake. Intake rate of leaves decreased at plants densities < 322 plants/m2. The maximum intake rate of leaves was 25% of the intake rate of aspen buds observed in wild grouse (Huempfuer and Tester 1988). Intake rate of raisins was an asymptotic function of bite size and was 20 times greater than the intake rate of leaves. Ruffed grouse in the Southeast must forage for> 100 min/day under ideal conditions to satisfy energy requirements. / Ph. D.

I addressed the effects of landscape characteristics on ruffed grouse (Bonasa umbellus) home range size and movement, and examined grouse selection for specific landscape characteristics and cover types. Grouse home ranges and movement patterns derived from telemetry data gathered from fall 1996 through fall 1998 were overlaid onto a GIS database of Clinch Mtn. Wildlife Management Area, VA. This database was developed from GPS data and LANDSAT thematic mapping imagery (30 m pixel scale) and was composed of 22 cover types. Landscape metrics were calculated using FRAGSTATS/ARC, and multiple regression was used to relate changes in home range size and movement to these metrics. I used Wilcoxon signed-rank tests to compare the values of landscape metrics calculated for each home range to those calculated for the area encompassed by the home range plus a surrounding 300 m buffer. I used Wilcoxon rank-sum tests to compare the values of landscape metrics for the home ranges to the metrics calculated for 50 33 ha random plots. I used compositional analysis to test for preferential use of cover types. I developed 2 regression models (P < 0.01) relating changes in home range size to landscape characteristics, 1 model (P = 0.09) relating the distance between seasonal home range centers to landscape characteristics, and 1 model (P = 0.03) relating average daily movement to landscape characteristics. Grouse home range size increased as patch shape became more irregular and patch size and the number of different cover types per hectare increased, and decreased as the amount of high contrast edge in the landscape increased. The distance between seasonal home range centers increased as Shannon's diversity index and the average distance between patches of similar cover types increased, and decreased as the amount of high contrast edge increased. Average daily movement increased as the average distance between patches of the same cover type increased and as the percent cover of a full (~75%) rhododendron and/or laurel understory within a grouse's home range increased, and decreased as the amount of high contrast edge in a bird's home range increased. Ruffed grouse were selecting areas with high densities of smaller than average patches that were of uniform size and regular shape and contained higher than average amounts of high contrast edge. Areas containing a greater diversity of cover types than what was available in the study area also were preferred. Within these areas, clearcuts and mesic deciduous stands with a rhododendron/laurel understory were the most preferred cover types. Creating and maintaining a landscape with high densities of small patches that are of uniform size and regular (square) shape would provide the highest quality ruffed grouse habitat in this region. Several of these patches should be early successional cover to provide an abundance of high contrast edge. Rhododendron and/or laurel thickets also may be beneficial as supplemental winter cover, and mesic stands of mature hardwoods should be well interspersed with these cover types to provide supplemental food sources. / Master of Science

This dissertation examines the operation of neoliberal environmentality through the instrumentalization of the Greater Sage-grouse (Centrocercus urophasianus) in Wyoming. It treats technological interventions within environmental construction as generating biotic-machinic entanglements termed technonature. I present the formation and operation of the Wyoming Conservation Exchange as a case study of technonatural territorialization connected to global trona and hydrocarbon commodity flows. The theoretical framework elaborates how "the environment" is constructed and governed through tactical instrumental deployments connected to technocratic management allowing economically powerful actors to inscribe their desires within Wyoming's landscape, politics and biota as a function of environmental security related to commodity development. The question motivating this work is "Whose environment is the Environmental Defense Fund defending?" The Greater Sage-grouse has become an object of U.S. Federal environmental governance since the late 1990's. It has experienced significant population declines due to anthropogenic disturbance and habitat loss through industrial action across its range. Wyoming's Sagebrush Steppe contains 37.5% of the remaining range wide population. The grouse was listed as a candidate species under the 1973 U.S. Endangered Species Act triggering responses from Federal, State, and international wildlife management agencies, as well as environmental non-governmental organizations. Wyoming could lose nearly a quarter of its surface should Federal regulations require the designation of critical sage-grouse habitat. Governor Dave Freudenthal signed Executive Order 2008-2 into law in response to the regulatory threat to Wyoming's hydrocarbon and mineral based economy. The grouse, in response was de-listed as a candidate species in 2015 by the U.S. Fish and Wildlife Service. EO 2008-2 established the Wyoming Core Area Strategy as a statewide conservation umbrella and laid the framework for a habitat mitigation economy allowing industrial activity to continue within sage-grouse habitat. This incentivized the Environmental Defense Fund (EDF) to test a market-based instrument – a habitat exchange – within Wyoming. The Greater Sage-grouse is a test species as it is highly sensitive to changes in its environment and this dissertation examines how the habitat mitigation economy advanced by EDF is drawing the grouse into global commodity networks as a territorialization process for global flows of hydrocarbons and minerals. At stake is the ability to write the history of the species, land, and the global environment as EDF develops conservation technologies prioritizing flows critical to the hydrocarbon environment through the technology of the Wyoming Conservation Exchange. / Doctor of Philosophy / The Greater Sage-grouse (Centrocercus urophasianus) entered Euro-American scientific study as early as the Lewis and Clark expedition as they explored the Intermountain region of Western North America. The first thorough scientific study of the sage-grouse in the 20th Century, The Sage Grouse in Wyoming, by Dr. Robert Lansing Patterson included the effects of anthropogenic disturbance on grouse populations. Since the 1952 publication of Patterson's study, Greater Sage-grouse numbers have been declining as the bird loses its home to encroachments such as urbanization, agriculture, grazing, mining, and fossil fuel extraction. The last stronghold of the grouse is the Sagebrush Steppe within Wyoming containing nearly 40% of the remaining population. Known for its flamboyant mating displays, the ground-dwelling avian species has become a political flashpoint in conservation, land management, and environmental circles as its numbers declined steadily since the 1990's due to an accelerating energy boom threatening its habitat. The bird became a threat to extractive industry in Wyoming at the turn of the Millennium as environmentally concerned groups petitioned the U.S. Fish and Wildlife Service (UFWS) to evaluate its populations under the Endangered Species Act (ESA). Nearly a quarter of Wyoming's surface would be strictly policed as critical habitat were the grouse listed as endangered or threatened under the ESA. Wyoming and its partners created the Wyoming Core Area Protection Strategy (CAP) as a wildlife management framework through Executive Order 2008-2. The Wyoming CAP includes the foundation of a habitat mitigation economy allowing industry to trade surface disturbances within critical sage-grouse habitat for modified land purportedly to the benefit of the species. The Nature Conservancy invited the Environmental Defense Fund to form the Wyoming Conservation Exchange – a market-based conservation instrument tailored to trading in habitat mitigation credits. This dissertation studies the Wyoming Conservation Exchange as an instrument connected to larger networks of wildlife management agencies, non-governmental organizations, and mining and fossil fuel interests. It evaluates the effects of the Wyoming Conservation Exchange and the economy it seeks to establish as changing how the environment is managed across the Sagebrush Steppe. Environmental Defense Fund's conservation instrument is reviewed through the economy created for and through the Greater Sage-grouse as an object of environmental governance. Habitat offsetting can, has and will change the physical, and political environment of Wyoming allowing powerful actors to write the rules of how the environment should be managed. As such, this dissertation questions whose environment the Environmental Defense Fund is defending as it explores sage-grouse management within the state.

Natality characteristics were calculated for 3 regions in the southern Appalachians (Ridge and Valley, Alleghany Plateau, and Ohio River Valley). I report data collected in the first 2 years of a long term (> 6 years) study conducted by the Appalachian Cooperative Grouse Research Project (ACGRP). Nesting rate, pooled over all regions, was 83.6% in 1997 and 79.7% in 1998. In the 2-year period, the Alleghany Plateau reported the highest nesting rate (97.6%) while the Ohio River Valley reported the lowest rate (54.2%). Overall hen success rates were 81.5% in 1997 and 56.9% in 1998. Yearling hen success rates were as high or higher than adults. Adult hen success was 85.7% in 1997 and 48.5% in 1998; yearling hen success was 86.7% in 1997 and 82.3% in 1998. Additionally, I found a lower renest rate (8% over 2 years) in the southern Appalachians than previous studies have reported. The mean first-nest clutch size in the southern Appalachian region was considerably lower (9.5, years and regions pooled) than that reported for other portions of ruffed grouse range. Recommendations are given on how ACGRP natality data collection may be improved in upcoming years. Ruffed grouse chick survival estimates were calculated from data collected in the first 2 years of a long term ACGRP study as well as data collected separate from ACGRP protocol. First-week chick survival estimates ranged from 0.18 to 0.32 in 1997 and 0.45 to 0.48 in 1998. Late brood season survivorship values (0.11-0.13 at week 5, 0.07 at week 10) were considerably lower in the southern Appalachians than those reported from more northern portions of ruffed grouse range. Additionally, the mean number of chicks per brood in July was lower in the southern Appalachians than that reported in the Great Lakes region during July and August. Recommendations are given on how ACGRP chick count data collection may be improved in upcoming years. I compared micro-habitat characteristics at known brood locations with randomly selected locations to determine which characteristics are selected by ruffed grouse hens and broods in the southern Appalachians. In the first half of the brood season (weeks 1-6) hens and broods selected sites with tall, complete, vegetative ground cover. Additionally, broods selected forested sites with a well-developed canopy, rather than areas affected by large canopy gaps or openings. Higher ground cover at brood sites may have been due to a lack of midstory structure. The abundance of arthropods, fruit, and forage at brood flush sites was higher during the first few weeks of the brood season; this was possibly due to flush sites being located in open, mid-age or mature forest. Several authors have speculated that as the chicks' diet shifts from primarily arthropods to fruit and forage at approximately 3 weeks of age, the habitat selected by hens and their broods may change to accommodate this dietary shift. In my study, a change in habitat selection did not occur between weeks 3 and 4 as expected but after week 6 and may indicate the chicks' dietary shift occurs later than some have predicted. / Master of Science