Cape mountain zebra habitat utilization and diet in the dystrophic fynbos habitat of the Bontebok National Park was found to be highly selective. Mountain zebra concentrated in specific sectors and habitat types in the park on a seasonal basis, preferring recently burnt habitat with a veld age younger than one year in all seasons, except during the warm, dry summer. Proteoid Fynbos with a veld age between one and five years was preferred in the cool winter, while Drainage Lines and the fringes of Inland Pans were preferred during summer. Asteraceous Fynbos was avoided, irrespective of veld age, as well as all other habitats with a veld age greater than five years. Within habitat types with a veld age greater than one year, specific sites were selected and avoided on a seasonal basis, which were found to differ in terms of habitat suitability, based on the availability of dietary plant species. The annual diet consisted of 72.6% grass, 11.8% restio, 5.9% sedge, 8.8% geophyte and less than 1% forb and shrub species. Three grass species formed the bulk of the annual diet, Themeda triandra, Cymbopogon marginatus and Eragrostis curvula, for which leaf use was greater than stem use. Themeda trianda was preferred throughout the year, but contributed to the diet in greatest proportion in the warm, dry summer, when it was available at greatest leaf height and diameter. Cymbopogon marginatus was preferred during the cool winter, when diet composition and greenness was also greater than in other seasons. Preference of Cymbopogon marginatus decreased as leaf height and diameter increased. Eragrostis curvula was preferred in the warm autumn, when it composed the largest proportion of the diet, and selection of this species at feeding sites was based on both greenness and volume. In summer mountain zebra also preferred grass stems and inflorescences of Aristida diffusa, Stipagrostis zeyheri and Briza maxima. During the cool spring the diet included stems and inflorescences of sedges and restios, primarily Ischyrolepis capensis, and in autumn, dry bulbs of the geophyte Moraea collina were utilized. Habitat utilization, as well as grass height and greenness surveys in the Recently Burnt Area, and the nutritional status of mountain zebra was found to be in line with the Summer Nutritional Stress Hypothesis. The hypothesis proposes that the harsh climatic conditions of the area during summer are linked to the low availability of C4 grass, on which grazers would depend in summer. This is supported by the avoidance of the Recently Burnt Area in summer, and the preference of species like Themeda triandra during summer despite low greenness levels. Faecal nitrogen and phosphorus for mountain zebra in Bontebok National Park and De Hoop Nature Reserve were at minimum levels during the warm seasons. Faecal nitrogen was below the threshold for dietary deficiency in all seasons except spring, and faecal phosphorus was above the threshold for deficiency during two seasons only. The findings of this study are in line with other recent work on mountain zebra in the Baviaanskloof suggesting that, due to a high required rate of forage intake, mountain zebra are limited by both poor resource quantity and quality in dystrophic fynbos ecosystems.

Global declines in amphibian species have directed research towards investigating why this is happening. One of the major causes of these declines is the fragmentation and loss of habitat. This study examined the effect of land use on the species composition of frogs within North Eastern KwaZulu-Natal, and the use of buffer zones to facilitate the protection of these species. Three land use types were investigated: sugar cane (Saccharum officinarum), gum (Eucalyptus sp.) plantations and conservation areas. The average number of frog species differed between areas: conservation 13.2 ± 6.6; plantations 3.8 ± 1.3; and sugar cane 2.8 ± 1.4. Sugar and gum plantation were found to be lacking the wetland and grassland/woodland habitats. In addition to this, the frog species that were not present on these land use types were those that are totally dependent on water as well as those that are not dependent on a water source. Two species were highlighted as possible indicator species of land use: Amietophrynus gutturalis and Hyperolius marmoratus. To mitigate the effect of these land use types, the use of buffer zones was explored in a desktop study. A range of buffer zones were applied to wetlands in a sample study area, using a range of distances including the distances of 290 m and 159 m recommended by Semlitsch and Bodie (2003), and the recommended distances for wetlands in South Africa of 10-20 m. The application of a 290 m and 159 m buffer zone on a conglomerate of wetlands connected by a 100m buffer was the most feasible as it incorporated a percentage of the total study (6.4% and 4.3%) area similar to the percentage occupied by the recommended 20m buffer zone (5.5%) around all wetlands, and still incorporated the range under protection put forward by Semlitsch and Bodie (2003). Management implications of these findings are discussed. / Thesis (M.Sc.)-University of KwaZulu-Natal, Pietermaritzburg, 2010.

Thesis (MSc)--University of Stellenbosch, 2006. / ENGLISH ABSTRACT: The conservation of biodiversity is becoming increasingly challenging as habitats are disturbed, fragmented or destroyed. Although nature reserves now cover more than 10 % of the earths’ surface it has become clear that more will have to be done to ensure the long-term survival of species. Therefore, focus is increasingly shifting towards conserving biodiversity in natural and semi-natural remnants in human-influenced areas. This study aimed to determine the contribution of remnants in human-influenced areas to the conservation of biodiversity in the Cape Floristic Region (CFR) lowlands, using ground-dwelling arthropods, specifically ants, as the focal taxon. Initially, base-line information of arthropods and in particular ants was obtained. Sampling arthropods generally involves a large sample effort. Therefore maximizing sampling effort for ants in the CFR was investigated by trapping ground-dwelling ants at a single locality. Doubling the number of grids of pitfall traps was found to be more effective in trapping a greater number of species than doubling the duration of sampling. Therefore increasing spatial sampling intensity rather than sampling duration maximizes sample effort for CFR ants. Also, the seasonal changes of ground-dwelling arthropods, including ants, were determined by sampling four times during the year at a single locality. Overall arthropod abundance was found to peak in summer while dropping to a minimum in winter. This pattern was mirrored by that of the ants, indicating that ant results have a broader relevance than to ants only. The ground-dwelling fauna was dominated by ants emphasizing their importance in the CFR lowlands, and demonstrating that ants are an appropriate flagship taxon for epigaeic arthropod diversity in the CFR. Finally the contribution of remnants in human-influenced areas to the conservation of the CFR was investigated. A nested hierarchical approach was used, where five localities were selected across the CFR, each containing one reserve site and one site with natural remnants. Ants were sampled, along with environmental variables, namely weather, vegetation and soil. Overall, remnants were found to support similar ant assemblages to those of reserves. However for individual localities some remnants were significantly different to their reserve counterparts. Differences in ant assemblages were found to be greater between localities than between reserves and remnants. The relatively high heterogeneity of ants found in this study emphasizes the conservation significance of invertebrates along with that of plants in the CFR. Remnants clearly show the potential to conserve ant assemblages, however correct management is needed for these areas to maximize their potential. Disturbances such as the presence of the invasive Argentine ant and increasing soil nutrients by fertilization, pose a distinct threat to the ability of remnants to conserve ant assemblages. This study has shown that remnants currently support ant assemblages representative of those present in the CFR today. Therefore, some remnant patches of habitat in agricultural areas currently do contribute highly to the conservation of a functional important taxon in this global biodiversity hotspot, and if managed correctly, may continue to do so in the future. / AFRIKAANSE OPSOMMING: Die vernietiging en fragmentering van habitatte maak die bewaring van biodiversiteit al hoe meer van ‘n uitdaging. Alhoewel natuur reservate reeds meer as 10 % van die aarde se oppervlak beslaan is dit duidelik dat meer gedoen sal moet word vir die lang-termyn voortbestaan van spesies. Dus word die fokus van biodiversiteit-bewaring toenemend gerig op bewaring van natuurlike en semi-natuurlike fragmente in menslik-beinvloede gebiede. Die doel van hierdie studie was om te bepaal wat die bydrae van fragmente van natuurlike veld in menslik-beinvloede gebiede is tot die bewaring van die streek. Dit is gedoen deur van grond-lewende geleedpotiges en spesifiek, miere in die Kaapse floraryk (CFR) gebruik te maak. Aanvanklik is kennis ingewin oor die geleedpotiges en spesifiek miere in die omgewing. Omdat die versameling van geleedpotige diere gewoonlik baie moeite vereis is ‘n maksimum steekproef gedoen by ‘n enkele lokaliteit. Daar is gevind dat ‘n verdubbling van die aantal ruitsteekproefnemings met vanggate meer effektief is om miere te vang as ‘n verdubbling in die tydperiode wat vanggate oop is. Dus, is ‘n hoër ruimtelike steekproef intensiteit meer effektief in vergelyking met ‘n langer tydsduur vir miere in die CFR. Die seisoenale veranderinge van grond-lewende geleedpotiges, sowel as miere, was ook bepaal. Dit was gedoen deur vier seisoenale steekproewe te doen by ‘n enkele lokaliteit. Die totale geleedpotige-talrykheid was die meeste gedurende die somer en die minste in die winter. Die miertalrykheid het ook hierdie patroon weerspieël. Dit dui daarop dat veranderinge in mier versamelings van breër belang is vir alle grondlewende geleedpotiges. Miere was die dominante grond-lewende geleedpotiges en beklemtoon die belangrikheid van miere in die CFR, sowel as hulle toepaslikheid as vlagskip taksa vir grond-lewende geleedpotige diversiteit in die CFR. Laastens was die bydrae van gefragmenteerde natuurlike veld in menslik–beinvloede gebiede tot die bewaring van die CFR ondersoek. ’n Krimpende/ genestelde hiërargies benadering is gebruik in vyf geselekteerde lokaliteite, elk het bestaan uit ‘n area in ‘n natuur reservaat en ‘n area in ‘n naasliggende fragment. Miere was versamel saam met ‘n verskeidenheid omgewings veranderlike, naamlik weer, plantegroei en grond. In die algemeen is gevind dat fragmente en reservate gelyksoortige mier versamelings het. Daar was wel gevind dat party fragmente aansienlik verskillend was van die reservaat teenstuk. Verskille in mier versamelings tussen lokaliteite was groter as verskille tussen reservate en fragmente. Die relatief hoë heterogeniteit van miere beklemtoon die bewaringsbelang van invertebrate saam met dié van plante in die CFR. Dit is duidelik dat fragmente wel ‘n potensiale bydrae kan maak om die mier versamelinge te bewaar, maar gepaste bestuur is nodig om hierdie potentiaal te maksimaliseer. Versteurings soos die teenwoordigheid van die indringer Argentynse mier en toenemende grondvoedingstofkonsentrasie as gevolg van bemesting is ‘n groot bedreiging tot die vermoë van fragmente om mier versamelings te bewaar. Hierdie studie wys dat mier versamelings in gefragmenterde areas verteenwordigend is van die algemene mier versamlings wat op die oomblik in die CFR is. Dus lewer party fragmente in landbou gebiede op die oomblik ‘n wesenlike bydrae tot die bewaring van ‘n funksioneel belangrike takson in hierdie globale biodiversiteitsbrandpunt en die bydra sal volhoubaar wees met korekte bestuur.

Thesis (MPhil)--Stellenbosch University, 2003. / ENGLISH ABSTRACT: South Africa, like so many other developing countries, is confronted with a very complex situation: although large-scale environmental degradation is threatening the future of the country, environmental matters are not regarded as of sufficient importance amongst the general population for them to contribute to environmental improvements. Most of the country's populations, many argue, are simply too poor and too focused on fulfilling short-term survival needs to give priority to the environment. Employing the 2001 World Values Survey, this study analyzes the attitudes of the general public of South Africa towards environmental matters. These attitudes are contrasted with those of the elites, who have a disproportionate influence over policy-making, by looking at the 2000 South African National Leadership Opinion Survey. This study finds that, as hypothesized, the general population of the country is quite unwilling (or unable) to contribute materially or financially to environmental improvements, especially in relation to the elites. In the final part of the study, these findings are employed in the development of a set of guidelines that policy-makers can use to increase the probability of developing successful and effective environmental policies. These guidelines take into account the nature and dimensions of popular attitudes, and consider ways in which environmental policies can foster the support of a public that is confronted with so many other serious issues. / AFRIKAANSE OPSOMMING: Suid-Afrika, soos soveel ander ontwikkelende lande, is gekonfronteer met 'n baie komplekse situasie: alhoewel grootskaalse degradasie van die omgewing die toekoms van die land bedreig, sien die algemene publiek nie omgewingskwessies as belangrik genoeg om 'n bydrae te lewer nie. Die argument is dat te veel van die land se bevolking net te arm of te gefokus is op die bevrediging van kort-termyn oorlewingsbehoeftes om prioriteit te gee aan omgewingskwessies. Deur gebruik te maak van die 2001 Wêreld Waardestudie, analiseer hierdie studie die houdings van die Suid-Afrikaanse publiek teenoor omgewingskwessies. Hierdie houdings word gekontrasteer met die van die elites, wat 'n disproporsionele invloed het oor die beleidsmakingsproses, deur gebruik te maak van die 2000 Suid-Afrikaanse Nasionale Leierskap Opinie Studie. Die studie vind dat, soos gehipotiseer, die publiek onwillig (of net nie in staat) is om bydrae te lewer tot die oplossing van omgewingskwessies, veral in vergelyking met die elites. In die finale afdeling van hierdie studie word die bevindinge gebruik om 'n stel riglyne te ontwikkel wat deur beleidmakers gebruik kan word om die waarskynlikheid van suksesvolle en effektiewe omgewingsbeleid te verbeter. Hierdie riglyne neem in ag die natuur en dimensies van publieke houdings, en bring na vore maniere waardeur omgewingsbeleide die ondersteuning van 'n publiek, wat gekonfronteer word met soveel ander ernstige kwessies, kan opbou.

Thesis (MScConsEcol (Conservation Ecology and Entomology))--University of Stellenbosch, 2009. / Habitat loss and fragmentation are two of the most pressing threats to biodiversity. Avifaunal diversity and integrity is under immense pressure from these two processes. We have made major advances in our understanding of avifaunal responses to habitat fragmentation, but mostly focus on either fragment scale and/or landscape scale influences of fragmentation on birds. A more comprehensive approach to assessing the impacts of fragmentation was used in this study. The avifaunas of two different geographical regions and bioregions were surveyed and a multiscale analysis of avifaunal responses to fragmentation was attempted. The study sites include the West Coast and East Coast Renosterveld Bioregions in the Cape Floristic Region, South Africa. Assemblage shifts, feeding guild compositional changes, species abundance variation and species persistence were examined at the three spatial scales. Time- and distance-restricted point counts were used to document birds that were directly dependent on the habitat fragments. Forty fragments were selected in each bioregion and a once-off snapshot of the avifaunal richness and diversity was obtained. Results indicate that the avifauna of the two bioregions responded differently to habitat fragmentation. In the East Coast Renosterveld Bioregion, the assemblages, guild composition and species abundances were most accurately predicted by landscape configuration. An assemblage shift occurred at 20 ha fragment area, compared to the 50 ha fragment area threshold of the West Coast Renosterveld Bioregion’s avifauna composition. In the West Coast Renosterveld Bioregion, fragment area was the better predictor of assemblage, guild composition and species abundances. However in both bioregions, the persistence of common species was equally sensitive to area and landscape scale effects.

M.Sc. (Aquatic Health) / Yellowfish and specifically Labeobarbus marequensis are a charismatic species targeted by anglers throughout South Africa. Their population are limited to the north-western parts of the country including the lower reaches of the Crocodile River that flows through the Kruger National Park (KNP). Despite conservation efforts the Crocodile River in the KNP is still highly impacted. The effect of these impacts on the ecosystem is largely unknown. The main aim of the study was to determine the influence of changing water quantity and quality in the Crocodile River on adult L. marequensis. This was achieved by evaluating altered flows (discharge) on the behaviour of adult L. marequensis in the Crocodile River using biotelemetry over a two year period. The influence of altered water quality was assessed using metal bioaccumulation as an indicator of metal exposure in L. marequensis, Clarias gariepinus and Hydrocynus vittatus in the Crocodile and Sabie Rivers during a high and low flow season. Biotelemetry was used on 16 L. marequensis and 12 H. vittatus to determine the habitat use and movement responses of the species. Fish were tagged with Advanced Telemetry Systems (ATS) and Wireless Wildlife (WW) tags and tracked remotely and manually. Home ranges were determined using Arc GIS ®, Habitat uses were analyzed using Windows Excel (© 2011, Microsoft inc.). Environment variables recorded were scored as primary and secondary and then combined with a weighting variable 2:1 ratio (primary variable: secondary variable). A mixed-model analysis of variance (ANOVA) approach with a random co-efficients model and Akaike’s information criteria (AIC) were used to test for significance. Analyses were conducted using SAS version 9 (SAS institute, Cary, NC)...

Herbivores have a significant influence on their environment. Factors that influence herbivore distribution in a landscape are important for conservation. Artificial water provision is one such factor, with water sources being focal points of herbivore activity. Variation between herbivore utilisation of different waterhole types and habitat integrity surrounding the different waterholes is assessed in this study. Correlations are drawn between herbivore utilisation and habitat integrity to quantify the impact that artificial waterholes have on the landscape. A scoring system is devised to investigate the various factors affecting vegetation around artificial waterholes. Results show that there are significant variations between herbivore utilisation for different artificial waterhole types, as well as significant variation in habitat integrity surrounding the different waterhole types. Distance between waterholes and drainage lines, and utilisation by specific herbivore species have a significant impact on habitat integrity - specifically affecting veld condition and disturbance observed on woody plant species. A habitat score that was created by combining data from both the herbaceous and woody layers is not affected by waterhole type, distance from waterholes, or the different herbivore species utilising the different waterhole types. Earth dams have the greatest impact on surrounding vegetation and are the most utilised waterhole type. Closures of earth dams are not recommended due to their importance to herbivores. Less utilised waterhole types are also important, mitigating the impact of herbivore damage to vegetation at earth dams. / Environmental Sciences / M. Sc. (Nature Conservation)

Thesis (PhD)--Stellenbosch University, 2011. / ENGLISH ABSTRACT: Both taxonomic diversity and diversity of species interaction networks are experiencing declines as a result of agricultural intensification at habitat and landscape scales. Reversing this trend is a key conservation issue, particularly for important functional groups such as flower-visiting insects and the networks within which they interact. This is of great concern in regions of high conservation priority such as the Cape Floristic Region (CFR), known for its high level of floral and faunal endemism and exceptional species turnover. Holistic approach to conservation in agricultural landscapes involves both preservation of natural land and wildlife friendly management of the farm land to achieve conservation targets. The value of these extensive management approaches is yet to be fully assessed, especially in perennial systems such as vineyards. I examined here the effects of vineyard management and landscape context on species richness and abundance of flower-visiting insects and their species interaction networks. Possible taxon specific effects were verified. I also investigated whether vineyards under organic and conventional management homogenized insect-flower interaction networks and whether vineyards with different management practices vary in patterns of species turnover. I sampled flower-visiting insects and their interactions in organic and conventional vineyards, and in natural reference sites. Inclusion of natural reference sites enabled me to make management recommendations for patches of natural vegetation in CFR agricultural landscape. Statistical models showed taxon-specific benefit of organic farm management, and of landscape (distance to natural habitat). There was benefit to monkey beetles (Scarabaeidae) but not to bees (Apidae). Organic vineyards had a higher number of insect-flower interactions than conventional ones, but vineyards under the two types of management were similar in terms of other important network indices. However, networks of the vineyards were more nested than the natural sites, indicating that they may be potentially more stable to perturbation and random extinctions. Multivariate dispersion tests revealed insect-flower interaction networks were not homogenized by both organic and conventional vineyards across the landscapes. I also found, through additive partitioning, that organic and conventional vineyards were similar in terms of species turnover of bees and flowering plants. The findings of this study provide heuristic value to current debates on the value of vineyard habitats for insect conservation. Both organic and conventional vineyards that promote sustainable management of the non-crop floral vegetation between vineyard rows are potential solutions for conservation of flower-visiting insects and their interactions. Also, attention has to be paid to the quality and connectivity of the natural habitat patches that are within CFR agricultural landscape. Site specific management and assessment of the value of these landscape elements is important. Management approaches such as carefully controlled burning may be beneficial, as the CFR natural vegetation is a fire-driven community. / AFRIKAANSE OPSOMMING: Taksonomiese diversiteit en diversiteit van spesies-interaksie netwerke ondervind beide afnames as gevolg van landboukundige intensifikasie op habitat en landskap skaal. Om die neiging terug te swaai, is ’n sleutel bewaringsaangeleentheid, veral vir belangrike funksionele groepe soos blom-besoekende insekte en die netwerke waarbinne hulle op mekaar inwerk. Dit is van groot kommer in streke met hoë bewaringsprioriteite soos in die Kaapse Floristiese Streek (KFS), bekend vir sy hoë vlak van plant- en dierendemisme en buitengewone spesies kentering. ’n Holistiese benadering tot bewaring van landboukundige landskappe behels beide die bewaring van natuurlike land en natuurlewe-vriendelike bestuur van die plaasgrond om bewaringsdoelwitte te bereik. Die waarde van hierdie ekstensiewe bestuursbenaderings moet nog volledig bepaal word, veral in meerjarige sisteme soos wingerde. Ek het die uitwerkings van wingerdbestuur en landskapsamehang op spesiesrykheid en volopheid van blombesoekende insekte en hulle spesies interaksie netwerke ondersoek. Moontlike takson-spesifieke uitwerkings is nagegaan. Ek het ook ondersoek ingestel of wingerde onder organiese en gebruiklike bestuur ooreenstemmende insek-blom interaksie netwerke met wingerde met verskillende bestuurspraktyke in patroon van spesies kentering gewissel het. Ek het blom-besoekende insekte en hulle interaksies in organiese en konvensionele wingerde, asook in natuurlike verwysingsgebiede gemonster. Insluiting van natuurlike verwysingsgebiede het my in staat gestel om bestuursvoorstelle vir gebiede van natuurlike plantegroei in KFS landboulandskappe voor te stel. Statistiese modelle toon takson-spesifieke voordeel van organiese plaasbestuur en van die landskap (afstand van natuurlike habitat) self. Daar was voordeel vir bobbejaankewers (Scarabaeidae), maar nie vir bye (Apidae) nie. Organiese wingerde het ’n groter getal insek-blom interaksies as konvensionele wingerde gehad, maar wingerde onder beide tipes van bestuur was soortgelyk in terme van ander belangrike netwerk aanduiders. Netwerke van wingerde was egter meer geklomp dan natuurlike gebiede wat aandui dat hulle potensieel meer stabiel betreffende versteuring en lukrake uitsterwings is. Multivariate verspreidingstoetse het aangetoon dat insek-blom interaksie netwerke by beide organiese en konvensionele wingerde oor landskappe nie eenvormig was nie. Ek het ook bevind deur aanvullende verdeling dat organiese en konvensionele wingerde gelykwaardig was in terme van spesies kentering van bye en blomplante. Die bevindings van hierdie studie verskaf heuristise waarde tot huidige debatte oor die waarde van wingerdhabitatte vir insekbewaring. Beide organies en konvensionele wingerde wat volhoubare bestuur van die nie-gewas plantegroei binne wingerdrye bevorder, is moontlike oplossings vir die bewaring van blom-besoekende insekte en hulle wisselwerkings. Bykomend moet aandag gegee word aan die kwaliteit en verbindings van en tussen natuurlike habitat gebiede binne die KFS landboulandskap. Plekspesifieke (plaaslike) bestuur en bepaling van die waarde van hierdie landskapelemente is belangrik. Bestuursbenaderings, soos noukeurig beheerde brand, mag voordelig wees aangesien die KFS natuurlike plantegroei ’n vuurgedrewe gemeenskap is.

Thesis (MScConsEcol (Conservation Ecology and Entomology))--University of Stellenbosc, 2007. / Freshwater insect species and their host ecosystems are widely threatened, particularly within agricultural and urban landscapes of Mediterranean areas, including that of the Western Cape Province, South Africa. The study here determined the biodiversity value of nineteen artificial ponds (temporary and permanent) and two river margin sites in the Cape Floristic Region (CFR). The aim was to determine aquatic beetle and bug abundance and species richness in these ponds, a topic which has been hardly explored in South Africa. Sites were sampled from September 2005 to September 2006. A total of 18 677 aquatic beetle and bug individuals were sampled, representing 43 genera (28 beetle and 15 bug genera), 64 species (44 aquatic beetle and 20 bug species). Different pond types had distinct species assemblages. The associated physico-chemical characteristics of these sites were also investigated. The key environmental variable affecting aquatic beetle and bug distribution was naturalness (no human and environmental impact on waterbody). The results showed that the most important determinant of aquatic beetle and bug species richness across all sites was emergent macrophytes, such as Typha capensis and Persicaria decipiens. Artificial ponds in the CFR clearly increase the area of occupancy for insects, and therefore play a major role in conserving them. This is especially so when the ponds are at moderate temperatures (i.e. 19º C). Although all ponds contributed to the aquatic beetle and bug diversity in the region, they differed in relative value, depending on the combination of environmental variables affecting each pond.