Gabbroic rocks in sills commonly display well organized magnetic fabrics that are generally attributed to magma flow. Yet in a number of cases, magnetic fabrics that are considered proxies for flow fabrics, form at a high angle to the intrusive - host rock wall. A symmetrical oblique magnetic fabric observed in opposite margins of a dike has been interpreted as the result of viscous drag against the margin, a model referred to as fabric imbrication. In this study tested if a similar viscous drag model could also apply to sills. The Karoo Large Igneous Province (LIP) of South Africa offers unparalleled opportunities to investigate fabrics, internal zonation and magma flow in mafic sills of thicknesses ranging from 5 to 50 m. Exceptionally; Karoo sills can reach a 1000 m in thickness (e.g., Insizwa). Geochemical studies suggest that the Karoo gabbro sills have a consistent tholeiitic magma composition across the entire Karoo Basin, especially as far as major elements are concerned. It is also practically undeformed and upon initial inspection unweathered, making it a well suited natural laboratory to test fabric models in gabbro sills. Magnetic fabrics are generally considered a good proxy for magmatic fabrics. The anisotropy of magnetic susceptibility has been measured on oriented hand specimens collected across vertical profiles through two representative sills. Scalar parameters such as magnetic susceptibility, degree of anisotropy and shape factor show variations with height in the intrusion. The variations in these parameters can be interpreted in terms of magmatic dynamic and static processes such as, for example, thermal convection, gravitational crystal settling or magmatic mush compaction. Directional parameters provide additional constrains on the dynamic vs. static nature of fabric-forming processes. Petrographic studies indicate that the primary carrier of the magnetic properties of the Karoo gabbros is titanomagnetite. Accessorily, pyrrhotite and hematite are present, but not in sufficient quantities to account for more than 1% of the sampled intrusions magnetic properties. Hydrous phyllosilicates are present in one sample, suggesting some alteration has occurred at the upper contact of the Country Club Road sill. Magnetic fabrics of the studied sills reveal complex susceptibility and remanence fabrics as a function of stratigraphic height. The proposed fabric development model does not sufficiently describe the observations, thus these fabrics were formed by a combination of mechanisms, both early and late in the cooling history of the intrusions. The observed magnetic fabrics display properties consistent with both the fabric imbrication model and the proposed viscous shear model. It is possible that crystal settling, melt segregation and possibly convection processes also were likely integral to the formation of the magnetic fabrics of the studied Karoo sills, though further studies are necessary.

This study was undertaken in the Karoo, a semi-arid grazing land in South Africa, to elucidate the interaction between grasshopper assemblages and various aspects of the Karoo landscape. It falls into four sections, the first of which was a three-year study which was undertaken on and around a prominent South African mesa to determine its role as an elevational conservation refugium for grasshoppers in a sea of grazed flatlands (Chapter 2). The number of grasshopper species and individuals on the summit, slopes and flatlands varied significantly, in relation to measured environmental variables. The summit, through inaccessibility to livestock grazing, was effectively a conservation refugium for one grasshopper species, Orthochtha dasycnemis. There was no significant difference in species richness between years of sampling, although there were significant variations in grasshopper abundance between years. The difference in rainfall between years was significant and appeared to be the key factor influencing grasshopper population dynamics. This clearly shows that a mesa can act as a conservation island and refugium supporting an insect assemblage that would be otherwise altered by heavy livestock grazing on the surrounding flatlands. This summit assemblage is strongly linked with that on the slopes and below, and is determined not so much by an island effect per se, but by low grazing intensity and associated soil and vegetation structure. The second part of the study focussed on the interaction between grasshopper assemblage response and three hill sizes at a regional scale (Chapter 3). Small hills contained a significantly higher grasshopper species richness and abundance than medium and large hills. There were significantly higher number of small-sized grasshopper species and individuals than medium and large-sized ones. Flatlands surrounding small hills had significantly higher grasshopper species richness and abundance than those surrounding medium and large hills. The slopes of the three hill sizes did not show significant difference in species richness and abundance. There was no significant variation between the summits of the three hill sizes in species richness but they varied in grasshopper abundance. The summits of small hills had significantly higher grasshopper abundance than the summits of medium and large hills. Detrended Correspondence Analysis showed two clear grouping of sampling site and grasshopper species. While the flatlands of small hills formed a separate assemblage of several grasshopper species, slopes and summits of all hills formed another clump of few grasshopper species. Canonical Correspondence Analysis revealed that flaltands surrounding small hills occurred along increasing gradients of shrub cover whereas those surrounding medium and large hills occurred along increasing gradients of grass cover, vegetation density and greenness of grasses. Slopes and summits of all hill sizes occurred along increasing gradients of rock cover, cragginess, grass height and soil temperature. Patterns of grasshopper dominance were markedly variable among sites. There were low dominance patterns on flatlands of small hills where most species were rare. The distributional patterns varied of higher taxonomic groups varied among the three hill sizes. Small hills contained species from four families and nine subfamilies, but medium and large hills had only members of Acrididae in five subfamilies. About 50% of the total grasshopper abundance were associated with small hills. The study revealed the patterns of grasshopper assemblages at regional scale, and showed that variability in hill sizes across the Karoo has marked role in grasshopper conservation, and that grasshoppers interact differentially with variable hill sizes across the Karoo. The third part of the study was undertaken at twelve grassland sites in the Mountain Zebra National Park (MZNP) and the surrounding farms to assess changes in grasshopper assemblages to grazing by indigenous mammals inside the park in comparison with grazing by domestic cattle outside (Chapter 4). The MZNP has been restored from cattle-grazed farmland to indigenous mammal parkland for 62 years. The number of grasshopper species and families inside the park was not significantly different from outside the park, but the number of individuals inside the park was significantly higher. Multivariate statistics did not reveal any strong site groupings based on simple inside/outside comparisons, but there were clear groupings of sites based on vegetation characteristics and other environmental variables. The park boundary, therefore, does not significantly determine grasshopper assemblages, although intensity of grazing does. The indigenous mammals inside the park had the same effect as the domestic cattle outside, and it was the level of defoliation and trampling that was important rather than type of mammal. Very intensive livestock grazing and trampling leads to bush encroachment and reduction in cover and/or disappearance of several grass species. In response to this pressure, grasshopper populations dropped, with localized extirpation of some species. Vegetation composition and structure showed a significant influence on grasshopper assemblages, particularly grass height and percentage cover. The MZNP is thus a localized area of elevated grasshopper abundance in comparison with the surrounding farm landscape, and presumably represents a situation prior to the current, intensive farming activities. Such elevated grasshopper The significantly lower population of grasshoppers on the surrounding farms, with local extirpation of some species also suggests that the MZNP could be viewed as a local centre to which species with higher capacity for mobility may seek refugia from anthropogenic pressures. Hence the MZNP serves as a reference showing the difference between restored through- natural-succession and anthropogenically-disturbed habitats, and compares desirable with undesirable ecosystem changes for herbivorous invertebrates such as grasshoppers. The fourth part of the study was on grasshopper assemblage response to seasonal grazing (including summer, winter, spring and autumn grazing), rotational grazing, continuous resting and continuous grazing at a long-term experimental site (Chapter 5). Rotationally grazed sites supported the highest number of grasshopper species and abundance, while continuously-grazed sites had the lowest. Cluster analysis revealed that spring-grazed and winter-grazed sites were the most similar, with continuously-rested sites being the next most similar to these. Rotationally-grazed sites showed the lowest similarity to the rest of the sites. DCA showed clear groupings of sites and grasshopper species, with most species associated with rotationally grazed sites. Continuously-grazed sites had a different grasshopper assemblage. CCA showed that the assemblages followed definite gradients of measured environmental variables. Rotationally-grazed sites occurred along gradients of increasing bare ground, while continuously-grazed and summer-grazed sites occurred along increasing gradients of shrub cover and soil temperature. Spring-grazed, autumn-grazed, winter-grazed and rotational1y-grazed sites were characterized by high vegetation density. Grasshopper dominance patterns were very different at different sites. Summer-grazed sites had the highest percent dominance (40%) by Picnodictyaflavipes, while winter-grazed sites showed higher percent dominance (32%) by Pseudogmothela sp. The significance of variable grazing management systems for maintaining floral and grasshopper diversity is discussed. It is recommended that rotational grazing in this arid system is most suited to maintaining plant and insect diversity. These four parts in this study all clearly showed that grasshoppers interact with the landscape in a way that their assemblage patterns are dictated by patch as well as by regional dynamics. Topography in particular contributes significantly to biodiversity patterns at the spatial scale of landscape. But these patterns are also strongly determined by differential grazing pressures from domestic livestock which in turn interact with the various topographical features. These findings enable recommendations to be made on optimal grazing regimes relative to the hilly features of the landscape. The results also show that restoration which incorporates low-pressure grazing regimes and which takes cognizance of topographical features can maintain grasshopper abundance and diversity in the long term. / Thesis (Ph.D.)-University of Natal, Pietermaritzburg, 2001.

An hierarchical syntaxonomic classification of the vegetation of the Camdebo and Sneeuberg regions of the karoo biome is presented as a second approximation after the earlier work by Acocks (1953). Details on the geomorphology, geology, climate, and early vegetation history of the area are given. The vegetation of the study area was stratified with the aid of Landsat imagery and the community classification was generated using two-way indicator species analysis (Twins pan) which produced ordered phytosociological tables. Tabular comparisons and final sorting of tables are according to the methods and techniques of the ZiirichMontpellier school of phytosociology. Syntaxonomic ranks are defined as five classes, nine orders and seventeen communities. The classes are Grasslands, Karoo Shrublands, Karoo Dwarf Shrublands, Sub-tropical Transitional Thicket, and Riparian Thicket. The distribution of syntaxa corresponds with the steep precipitation gradient experienced in the study area. These vegetation concepts are applied to the description of the flora of the Karoo Nature Reserve and an analysis of the total flora of the reserve is provided. The communities of the pediments, which contain the highest number of endemics, are poorly conserved. I test the validity of the vegetation classification by interpreting the results of an analysis of soils within the hypothesized vegetation units. There is a gradient of increasing Na, silt and pH levels from the Shrublands and Grasslands to the Succulent and Grassy Dwarf Shrublands of the pediments. A qualitative model of the vegetation history during the glacial-interglacial sequence in the Graaff-Reinet region of the eastern Cape is presented. Using a descriptive approach, the distribution patterns of 68 taxa, which are differential species for Karoo Shrublands, Succulent Thicket and Karoo Dwarf Shrublands, are investigated relative to major southern African biomes. The results indicate that a large proportion of the differential species in the phytosociological classification show affinity with Grassland and Savanna Biomes. Three species groups encountered in the Dwarf Shrublands show affinity with the Nama-Karoo biome. The differential species of the Succulent Thicket have a predominantly subtropical distribution. Using an historical approach, the palaeoenvironment of the region during the past 20 000 years is discussed briefly. On the basis of the descriptive and historical perspectives, a qualitative model of vegetation history is presented. The Succulent Thicket may have become established on edaphically favourable sites in the ameliorating conditions of the warmer, wetter Holocene subsequent to the Last Glacial Maximum. The Dwarf Shrubland and Succulent Dwarf Shrubland are depauperate in relation to ccmmunities in other southern African biomes, but the relatively large number of endemics suggests a long history in the region. Their differential species groups occur under arid conditions, accompanied by soils with high base and fertility status. The Dwarf Shrublands may have been more extensive during the drier glacial times on those sites currently occupied by Shrubland. The Shrublands display the expected affInity with the Grassland and Savanna Biomes. The small number of endemics suggest that these communities may have occupied the region in the period since the Last Glacial Maximum. Species with Succulent Karoo Biome affInity are poorly represented. The reliability of using Landsat products to detect and map the vegetation of the region is assessed. The manual classification of Landsat standard products provides a poor reflection of the vegetation of the arid, sparsely-vegetated bottomlands and pediments. The products provide good representation of the boundaries of thicket vegetation, but this uni-temporal approach does not distinguish between floristically different thicket communities. After analyzing digital Landsat data, I suggest that the multi-spectral scanner detects the boundaries of broad soil pedons and geological formations in areas of low vegetative cover. I describe and map the vegetation categories of the region after manual interpretation of six Landsat scenes. This is an effIcient, cost-effective method of mapping vegetation in extensive regions. The mapping units do not reflect the syntaxonomic classification, representing rather an integration of physiographic, pedological, geological and floristic information. With the view to improving the classification of these units, I develop a qualitative model of the natural resources of the region using an expert system

The Beattie Magnetic Anomaly is the world’s longest terrestrial magnetic anomaly with a strike length of over 1000 km and a wavelength in excess of 100 km. Collinear with this is a large belt of elevated crustal conductivities called the Southern Cape Conductive Belt. Historical crustal interpretations proposed a common source of serpentinized ophiolite as an explanation for both the anomalous crustal magnetic susceptibility and electrical conductivities. Spreading between the Western and Eastern Cape of South Africa the mid- to lower crust that hosts these anomalies is obscured by the overlying Cape and Karoo Supergroups. Between 2003 and 2006, three high resolution geophysical experiments were completed across the surface maximum of the Beattie Magnetic Anomaly (BMA) and the Southern Cape Conductive Belt (SCCB). These included a magnetotelluric (MT) survey and near vertical reflection and wide angle refraction seismic profiles. Within the MT inversion model the SCCB appeared as a composite anomaly, which included a mid-crustal conductor which is spatially associated with the BMA and a laterally continuous upper crustal conductor which is located at depths equivalent to the lower Karoo Supergroup. Subsequently; the upper crustal conductor was identified in northern and eastern extensions of the magnetotelluric profile; a distance in excess of 400 km. Historical magnetometer and Schlumberger Sounding experiments have previously identified elevated conductivities in the Karoo sequences which were attributed to the Whitehill and Prince Albert formations. These carboniferous, transgressive sediments are known to be conductive from borehole conductivity surveys and direct measurements at surface. In order to constrain the conductive properties of these sediments, impedance spectroscopy (IS) experiments were completed on core samples collected from a historical borehole drilled near to the MT profile. Part One of this thesis presents the results of these experiments, which support the proposition that the Whitehill and Prince Albert Formations are responsible for the laterally continuous, sub-horizontal, upper crustal conductor visible in the MT inversion model. Vitrinite reflectance studies were performed on the same samples by the Montanuniversität, in Leoben, these results corroborate the proposition that elevated organic carbon, of meta-anthracite rank, is the primary conductive phase for the Whitehill and Prince Albert formations. Part two of this thesis completed forward modelling exercises using historical aeromagnetic data previously collected across the Beattie Magnetic Anomaly. Preliminary models were unable to fit the geometry of any single magnetic model with conductors present in the MT inversion model discounting the proposition that the SCCB and BMA arise from a single crustal unit. Two constrained models were arrived at through an iterative process that sought a best fit between the measured data and the NVR crustal interpretations. The first model, proposes a largely resistive unit which incorporates portions of elevated crustal conductivity; these conductors are spatially correlated to crustal portions also characterised by high seismic reflectivity. The size of this modelled body suggest the likely host of the BMA is an intermediate plutonic terrane, analogous with the Natal sector of the Namaqua Natal Mobile Belt as well as the Heimefrontfjella in Dronning Maud Land, Antarctica, with magnetite hosted within shear zones. This is in agreement with previous studies. The second model proposes a lower crustal sliver imaged in the NVR data at depths proximal to the Curie Isotherm for magnetite and hematite as the source of the BMA. At these depths geomagnetic properties such as burial magnetisation or thermo-viscous remanent magnetism (TVRM) can potentially be linked to regional scale tectonic processes and can theoretically elevate a body’s net magnetic susceptibility. TVRM has been proposed for long wavelength crustal anomalies elsewhere.

Thesis (MSc)--University of Stellenbosch, 2003. / ENGLISH ABSTRACT: Arid and semi-arid rangelands of the Nama-Karoo Biome are believed to have changed considerably since the arrival of domestic livestock in the veld. Severe grazing pressure is considered to be one of the prime factors responsible for the perceived degradation of vegetation and concurrent soil degradation. To understand the process of degradation and to make further recommendations for future veld restoration, a large-scale project was undertaken in the Eastern Cape. This project focused on the role that landscape heterogeneity plays in providing refuges for plant species. The key question asked in this project was: What role do these hypothetically less grazed mesas play in the conservation of rangelands in the Nama-Karoo of the Eastern Cape? This study, a component of the larger project, investigated grazing and soil landscape gradients on- and off- the three mesas (Tafelberg, Folminkskop and Buffelskop) in the Middelburg District of the Eastern Cape, South Africa and the possible influences that they might have on the veld. The flats surrounding the mesas were mostly used as grazing camps in contrast to the plateaux of the mesas, which had varied levels of accessibility. In the absence of direct observations, the primary objective of the study was to test the differential dung pellet abundance and impact of animals on different parts of the landscape. As such, dung pellet counts in this study were correlated with surrogates of soil physical properties including bare ground, trampling and litter cover. Variation in dung pellet density was found at the different habitats (flats, slopes and plateaux) of all mesas. The flats to the north-eastern and south-western of Tafelberg mesa were found to be more heavily utilized by livestock and herbivores, while the plateaux and southeastern slopes of Folminkskop and Buffelskop were also utilized by grazers. The Tafelberg mesa was the only study site that was consistent with the hypothesis which stated that grazers would be less concentrated on the plateau compared to the surrounding flats due to its inaccessibility, whilst the high mean dung pellet density on the plateaux of the smaller Folminkskop and Buffelskop mesas due to easier access contradicted the original hypothesis. It appeared that dung pellet density did not clearly turn out to be an indicator of habitat use in this study, but showed where slopes and plateaux were accessible to herbivores, as in the case for the Buffelskop mesa, a higher abundance of dung pellets were found suggesting that higher intensities of habitat use took place. During this study a strong pattern was observed of bare soil patches on the flats surrounding the mesas. There was a decrease in percentage of bare soil along the gradient of the three mesas with a high percentage of bare soil on the plateaux of the mesas. A significant correlation was found between bare soil and dung pellet density. However, the plateaux of Folminkskop and Buffelskop had a high percentage of bare soil compared to the plateau of Tafelberg mesa. Farmers mainly used these smaller mesas as grazing camps for their livestock and herbivores. A positive correlation between bare soil and litter cover of the different habitats was evident in this study. A lower percentage of litter cover at these sites was associated with a high percentage of bare soil. Litter is very important in a healthy vegetation community in terms of nutrient cycling and fertile patches. A detailed assessment of soil chemical and physical properties would reveal, firstly, if vegetation change is better explained by soil or grazing effects and, secondly, if changes in soil have resulted from land use. Differences in macro- and micro-site variations between open-canopy (between shrubs) and closed-canopy (under shrubs) sites for each habitat were determined to differentiate between local scales due to land use and landscape scales due to geomorphology. The results suggested that carbon, phosphorus, calcium, magnesium, copper and manganese levels in soils at landscape scale better explain vegetation changes between habitats. At a local scale (open- and closed-canopy sites) land use was responsible for little changes in soils. Changes in only soil potassium, zinc and boron elements were actually a consequence of local scales due to land use. The soil nutrient content on the slopes appeared to be intermediate between the flats and plateaux of all three mesas. It appeared that dolerite capped Tafelberg and Folminkskop mesas had high silt and clay content, while Buffelskop (sandstone) mesa had a lower silt and clay content. Consequently, the texture and parent material of the soils contributed to the variations in soil nutrient composition between these mesas. High infiltration rate together with low nutrient content on the flats clearly showed that these flats, surrounding the mesas were degraded. The high infiltration rates were caused by high activity by livestock and other indigenous animals on the flats which breaks the surface crusting of bare soil and improve infiltration. It was concluded that high levels of grazing at these sites have also altered the textural and soil properties. Endozoochory dispersal and the deposition of dung pellets in areas of small patch disturbances play an important role in veld regeneration in degraded areas. Dung pellets collected from permanent study sites on the southeastern and northwestern flats and slopes, and all the study sites on the plateau of Tafelberg mesa, was sown in seedling trays, watered and monitored for seedling germination. Species list were then compared to below-ground soil seed bank data and above-ground vegetation data collected by other researchers at the same permanent study sites. Higher seedling percentages were recorded from dung pellets collected on the flats than on the plateau. A total of sixteen species were found to germinate in dung pellets collected on the flats compared to ten species germinating in dung pellets collected on the slopes and two species on the plateau of Tafelberg mesa. The seeds that germinated represent a variety of palatable grasses and shrub species. Aristida sp., Eragrostis bicolor, Eragrostis ch/orome/as and Eragrostis obtusa were palatable grasses recorded for dung pellets collected on the flats. Of the species recorded, Aristida sp., Chenopodium sp. and Pentzia sp. were found in dung pellets but were not recorded in parallel soil seedbank and vegetation studies. Successful restoration of veld conditions requires strict grazing management practices. Germination of seed in dung pellets might be considered to be a valuable means of indicating restoration potential and rangeland conditions for the identification of both degraded and conservation worthy areas. With appropriate land management skills and restoration measures, these challenges can be constructively and creatively faced. / AFRIKAANSE OPSOMMING: Daar word geglo dat die ariede en sernl-arlede weivelde van die Nama-Karoo bioom aansienlik verander het sedert die aankoms van mak lewende hawe in die veld. Swaar weidingsdruk word beskou as een van die vernaamste faktore verantwoordelik vir die waarneembare oorbeweiding van plantegroei en die gelyktydige grond agteruitgang (degradasie). Ten einde die proses van degradasie beter te verstaan, en om verdere aanbevelings te kan maak vir toekomstige veld herstel, is huidiglik 'n grootskaalse projek in die Oos-Kaap aan die gang oor die rol wat landskap ongelyksoortigheid speel in die voorsiening van skuiling aan plant spesies. Hierdie studie ondersoek weiding en grondlandskap gradiënte (op en af) van die drie mesas (Tafelberg, Folminkskop en Buffelskop) in die Middelburg streek van die Oos- Kaap, Suid-Afrika en die moontlike invloed wat hierdie gradiënte op die veld het. Die vlaktes was meestal gebruik vir weidingskampe in teenstelling met die kruin van die mesas. Hierdie studie het duidelike patrone vir habitat-gebruik op die vlaktes, hange en kruine van al die mesas aangetoon. Die vraag wat gestel was, was watter rol speel hierdie hipoteties minder beweide mesas in die bewaring van weivelde in die Nama- Karoo van die Oos-Kaap? Hierdie studie toets nie direk vir mis verspreiding as 'n plaasvervanger vir dier verspreiding in die landskap sedert werklike dier getalle nie oorweeg was nie. In die awesigheid van direkte waarneming, die primêre doelwit was om te toets die differensiaal misdigtheid en die impak van diere op verskillende dele van die landskap. Mis getalle in hierdie studie was gekorreleer met plaasvervangers van fisiese eienskappe insluitent onbedekte grond, vertrapping en droë plant material decking. Variasie in misdigtheid was gevind by die verskillende habitate (vlaktes, hange en kruine) van all die mesas. Die vlaktes aan die noord-oostelike en suid-westelike kante van die Tafelberg mesa was meer hewig benut deur lewende hawe en hêrbivore, terwyl die kruine en suid-oostelike hange van Folminkskop en Buffelskop ook gebruik was deur weidende diere. Die Tafelberg mesa was die enigste studie area wat konsekwent was met die hipotese, terwyl die hoë gemiddeld misdigtheid op die kruine van die kleiner Folminkskop en Buffelskop mesas, as gevolg van makliker toegang weerspreek die oorspronklike hipotese. Dit blyk dat misdigtheid duidelik opkom as 'n nie aanwyser habitat-gebruik in hierdie studie, maar wys well waar hange en kruine toeganklik was vir herbivore, soos in die geval van Buffelskop mesa, oorvloed van mis was gevind wat voorstel dat hoer intensiteit van habitat-gebruik voorgekom het. 'n Duidelike patroon is waargeneem tydens hierdie studie van onbedekte grond areas op die vlaktes rondom mesas. Daar was 'n afname in persentasie onbedekte grond langs die gradiënt van die drie mesas. Die kruine van Folminkskop en Buffelskop toon hoër persentasies onbedekte grond in vergelyking met die kruin van die Tafelberg mesa. Daar is 'n betekenisvolle korrelasie tussen onbedekte grond en misdigtheid gevind. Boere gebruik hierdie kleiner mesas hoofsaaklik as weidingskampe vir hulle lewende hawe en hêrbivore. In hierdie studie is daar 'n positiewe korrelasie gevind tussen onbedekte grond en droë plant material decking van die verskillende habitatte. Droë plantmateriaal bedekking speel 'n baie belangrik rol in 'n gesonde plant gemeenskap in terme van voedingsirkulering en vrugbare kolle. 'n Breedvoerige skatting van grond chemise en fisiese eienskappe wou openbaar eerstens of plantegroei verandering beter verklaar word deur grond of weiding effekte en tweedens, of verandering in grond veroorsaak was deur land-gebruik. Verskille in makro- en mikroterrein-wisseling tussen oop-blaredak (tussenplantruimtes) en toeblaredak (onderplantdekking) terreine vir elke habitat was bepaal om te onderskei tussen lokaal effek as gevolg van land-gebruik en landskap effek as gevolg van geomorfologie. Die resultate stel voor dat koolstof, fosfor, kalsium, magnesium, koper en mangaan vlakke in die grond by landskap effek beter plantegroei verandering tussen habitate verduidelik. By 'n lokaal effek (oop-en toe-blaredak terreine) land-gebruik was verantwoordelik vir min veranderinge in grond. Verandering in grond kalium, sink en baron elemente was werklik 'n gevolg van lokaal effek as gevolg van land-gebruik. Die nutrient inhoud van grond op die hange blyk intermediêr te wees tussen die van die vlaktes en kruine van al drie mesas. Hierdie studie toon ook dat die geologie en topografie van hierdie mesas die grond tekstuur en nutrient samestelling op en af van die mesas beinvloed. Dit wil voorkom asof die dolerite bedekte Tafelberg en Folminkskop mesas, hoë slik en klei materiaal bevat, terwyl Buffelskop (sandsteen) mesa 'n laer slik en klei inhoud het. Gevolglik, tekstueel en ouer materiaal grond dra by tot die variasie in grond nutrient samestelling tussen hierdie mesas. Hoë infiltrasie tempo tesame met lae voedingstof van die vlaktes dui daarop dat die vlaktes rondom die mesas oorbewei was. Hierdie hoë infiltrasie tempo word veroorsaak deur hoë dierlike aktiwieteite op die vlaktes deur lewende hawe en ander inheemse diere wat die oppervlakke van onbedekte grond versteur en infiltrasie verbeter. Die gevolgtrekking word gemaak dat die hoë weidingsvlakke van hierdie terreine ook die tekstuur en ander grond eienskappe verander het. Endozoochoreeuse verspreiding en die mis-afsetting in gebiede van klein areaversteurings, speel 'n belangrike rol in veld herlewing in oorbeweide gebiede. Mis versamel by permanente studie terreine op die SO en NW vlaktes en hange, as ook al die studie terreine op die kruin van die Tafelberg mesa, is gesaai in saailing bakke, besproei en gekontroleer vir saailing ontkieming. 'n Lys van spesies is vergelyk met onder-grondse saadbank gegewens en bo-grondse plantegroei gegewens wat deur ander navorsers by dieselfde studie terreine versamel was. Hoër saailing persentasies is gevind vir mis wat op die vlaktes versamel is as op die kruine. 'n Totaal van sestien geidentifiseerde spesies het uit die mis op die vlaktes ontkiem in vergelyking met die tien spesies wat ontkiem het uit die mis van die hange en twee spesies iut die mis van die kruin van Tafelberg mesa. Die ontkiemde saad verteenwoordig 'n verskeidenheid van vreetbare gras en struik spesies. Aristide sp., Eragrostis bicolor, Eragrostis chloromelas en Eragrostis obtusa is vreetbare grasse wat gevind was in die mis versamelop die vlaktes. Die spesies opname wys dat Aristida sp., Chenopodium sp. en Pentzia sp. Gevind was in die mis, maar nie in die parallelle grond saadbank en plantegroei studies nie. Die suksesvolle herstel van veldtoestande vereis "n streng weidingsbestuur toepassing. Die ontkieming van saad in mis kan beskou word as "n bekostigbare manier om veld herstel potentiaal te bepaal, asook weiveld toestande vir die indentifiseering van beide degradasie en bewaring van waardevolle areas. Met gepaste veldbestuur vaardighede en herstel maatreels, kan hierdie uitdagings konstruktief en kreatief aangespreek word.

Ph.D. (Botany) / Please refer to full text to view abstract

The Eastern Cape Karoo region is semi-arid with highly variable rainfall. This variability in rainfall sustains a mosaic of surface freshwater bodies that range from permanently to temporarily inundated. These waterbodies provide habitats for diverse invertebrate assemblages. The imminent hydraulic fracturing for shale gas has a potential to modify the water regime, with particular risk of salinisation. Accumulation of salts in freshwater wetlands results in loss of biodiversity, as invertebrate species shift from salt intolerant to salt tolerant species. This study therefore aims to expand on existing knowledge and provide new information on the distribution, diversity and structure of macroinvertebrate assemblages associated with various freshwater bodies in the region prior to shale gas exploration. Limnological and ecological aspects of thirty-three waterbodies (rivers, dams and depression wetlands) were investigated between November 2014 and March 2016. An experimental study on the effects of salinity on hatching success of branchiopod resting eggs was also included in the research. Rivers were characterised by high conductivity and depression wetlands by high turbidity, while dams had relatively higher pH than the other two waterbody types. In terms of global phosphorus interpretation guidelines, the results indicate that freshwater systems in the study region are predominantly eutrophic, indicating that agricultural run-off, particularly from livestock dung (goats, cattle and sheep), is an important source of phosphorus in the freshwater systems studied. Our results revealed new distribution records for branchiopod crustaceans in the Eastern Cape region, including the first record of Laevicaudata. Results showed that the sampled variables were unable to explain the variation in physicochemistry and invertebrate assemblage of several sites. Waterbody type, whether a depression wetland or a river, was the only factor that consistently showed an effect on the composition of both physicochemical data and invertebrate data. Depression wetlands ranged from completely bare to being extensively covered by macrophytes. Therefore, the effect of macrophyte cover in structuring macroinvertebrate assemblages was the focus of further investigation. The results indicated that the macrophyte cover gradient had little influence on the structure of the invertebrate assemblages in the depression wetlands, while only the presence/absence of vegetation significantly influenced the structure of the invertebrate assemblages in these systems. Surface area, dissolved inorganic nitrogen and pelagic chlorophyll-a were the environmental variables that best explained the variation in the macroinvertebrate assemblages among the sites. However, the differences in macroinvertebrate richness, abundance and distribution patterns among sites were only weakly influenced by local and regional environmental factors. These findings suggest that invertebrate in temporary wetland systems are adapted to the highly variable nature of temporary habitats, thus the influence of local variables is negligible. Results of the experimental study, on the effect of salinity on hatching success of branchiopod resting eggs, revealed that hatchling abundance and diversity of large branchiopods was significantly reduced at salinities of 2.5 g L−1 and above. Salt-tolerant taxa such as Copepoda and Ostracoda were the only ones to emerge in the highest salinity of 10 g L−1. Thus, should the region continue to experience increasing aridity and possible shale gas development, which all aggravate the salinisation problem, severe loss of branchiopod diversity (Anostraca, Laevicaudata, Spinicaudata and Notostraca) is likely to occur. This may lead to considerable decline in invertebrate diversity in the region, with cascading effects on food webs and ecosystem functions. The findings of this study can potentially be used in comparative studies on wetland invertebrate ecology in other semi-arid regions and in the formulation of policy and strategies for biodiversity conservation.

The formation of the Ellisras basin was influenced by the repeated tectonic instability that is also be noted through the changes in the energy levels of the depositing media of the basin-fill, from the early Carboniferous period, as already indicated by Siepker (1986). This theory is also supported by MacRae (1988) who suggests that the sediment accumulation in certain sections of the Ellisras Basin was indicative of periods of palaeoslope rejuvenation resulting in sandstones, grits and minor conglomerates, alternating with periods of stasis when extensive coal seam deposition occurred. Studies completed in this thesis indicate the influence of basin movement and depositional thickness of the basin’s formations. These depositional patterns are closely related to mobile geological structures such as lineaments or faults of continuously active geological structures. Bumby and van der Merwe (2004) and Bordy (2000) indicate the possible influence of the Limpopo Mobile Belt as well as the failed East African Rift System on the formation of neighbouring basins. Geological structures also present in the Ellisras Basin can also be observed in these neighbouring bodies such as the Tuli, Tshipise and the Soutpansberg basins. Observations within the region of the Ellisras Basin, made in neighbouring Botswana by Arnott and Williams (2007) describe the Soutpansberg trough as the main influence in the formation of the Ellisras, Mmamabula and Mopane coalfields. These observations are in line with the findings of this thesis which indicates the consistent presence of continuously active geological structures within this region such as the greater intracratonic Soutpansberg trough which indicated re-activation during late Permian to early Triassic times. Copyright / Dissertation (MSc)--University of Pretoria, 2010. / Geology / unrestricted

This study details results of small mammal surveys at 24 sites in four biomes (Nama- and Succulent Karoo, Albany Thicket, Grassland) as part of the Karoo BioGaps project to augment baseline biodiversity information needed to guide proposed fracking activities in the Shale Gas Development Area (SGDA) (Holness et al. 2016). A strategic Environmental Assessment (SEA), commissioned by the South African Government, evaluated the potential to exploit the supposedly substantial reserves of shale gas using hydraulic fracturing (“fracking”) in the Great Karoo Basin of South Africa. Terrestrial micromammals (<500g) were captured using Sherman livetraps (September 2016 to March 2017). Sampling over 6580 trap-nights produced 339 captures of 271 individuals representing 14 taxa. Trapping success was low 5.14% (mean per site 0.37± 0.61%). Most captures (87%) and individuals (83%) were recorded in the Nama-Karoo (294 captures, 226 individuals, 15 sites), whereas only 5 captures were recorded in Albany Thicket (3 sites). Four xerophilous/generalist species (Micaelamys spp. (Rock rats), Gerbilluscus paeba (Hairyfooted Gerbil), Macroscelides proboscideus (Round-eared Sengi), and Elephantulus spp. were numerically dominant at most sites, and within most biomes/bioregions; while five rare species were only ever recorded once. Mean α diversity (observed species richness Sobs) per site (2.88 ±1.99) and Shannon-Wiener diversity (1.70 overall, 1.04 ±0.33 per site) were low, with only 5.47 effective species (mean = 3.04 ±1.08 per site) and low equitability (0.64 overall). Sobs was highest in Nama-Karoo (13 species), and lowest in the Grassland and Albany Thicket biomes (2-5 species). Species accumulation/rarefaction curves did not reach asymptotes, and Sobs values for most sites/biomes/bioregions were significantly lower than Chao1 predicted species richness, suggesting that sampling effort did not accurately estimate species richness. However, trapping efficiency was generally high (56-100%; mean 86.7%) which compares favourably with that of two recent published studies in South Africa. Multiplicative beta diversity (βMt) across the SGDA was 4.56 indicating high species turnover between sites/biomes/bioregions. Species turnover was high across biome boundaries, notably Albany Thicket-Grassland (15), Nama-Karoo-Albany Thicket (14), and Succulent Karoo-Grassland (12). Biomes and bioregions tended to plot apart in ordination analyses with relatively low (40-60%) Sorenson similarity, indicating that most regional small mammal communities were well-differentiated. Despite data limitations, 66 new distribution records for 21 sites are reported for the SGDA. Total species richness (including historical records) was highest in the Nama-Karoo (19), particularly the Upper Karoo bioregion (19, mean 6.45 ±2.16, 11 sites), followed by the Grassland (16), and Albany Thicket (5) biomes. Total species richness records for most sites/regions fell within the iChao2 CI bounds, thus integrating trapping and historical records provided a relatively robust data set for subsequent spatial diversity analyses. However, even the total species richness dataset is likely to underestimate true diversity owing to not sampling arboreal species or detecting some cryptic species. Generalized linear analyses indicated that small mammal diversity indices were significantly associated with certain environmental/climatic parameters (livestock, drought). Despite the west to east increase in precipitation, highest diversity was concentrated in the arid north-west Nama-Karoo where dwarf shrubs and succulents predominate. This suggests that environmental and niche filtering are significant proximate factors shaping small mammal assemblages. No significant effects of biotic interactions (particularly competition following Diamond's (1975) first two rules) or resource-mediated niche limitations were evident for SGDA species assemblages. However, results for the Nama-Karoo (i.e. for a natural phytogeographical rather than geoeconomically-defined area) were significant suggesting that biotic interactions may also be proximate factors shaping local assemblages. Site assemblages were significantly nested, indicating that species at species-poor sites were subsets of those at richer sites; and thus, that site communities may have been structured by either long-term (ultimate) regional biogeographic processes (e.g. immigration and extinction related to distances between sites) or habitat filtering operating at local scales. Data deficiencies notwithstanding, my results present the most comprehensive landscape-level analysis for small mammals, and the only baseline dataset (based on randomized sampling) for the Greater Karoo and SGDA. While my results must be treated with caution, I am confident that the recommendations I make on species, sites and regions potentially vulnerable will be a useful guide to possible impacts of fracking in the study area.

The rocks of the Carboniferous to Jurassic aged main Karoo Supergroup of South Africa preserve an internationally significant and stratigraphically continuous retro-arc foreland depositional sequence. This succession documents environmental change from glacial-marine, through fluvio-deltaic to continental fluvial and aeolian, culminating in rift associated continental flood basalt extrusions. The fluvio-deltaic transition from marine to continental deposition has been the subject of much recent research and corresponds with the position of the Ecca-Beaufort contact. Over the entire basin this transition comprises three separate lithofacies associations deposited in the prodelta, deltafront and delta plain environments. Anomolously the southeastern contact is currently mapped as reflecting fluvial deposits of the Koonap Formation lying unconformably on prodelta deposits of the Fort Brown Formation. Detailed study across the Ecca-Beaufort contact in this part of the basin now reveals the same lithological transition as is present in the rest of the basin. Contrary to previuos work, this has shown that the Waterford Formation is indeed present in this part of the basin. This in effect means that the Grahamstown map sheet (1:250 000, Map 3326) requires modification to include this Formation and the new contact placement of the Ecca-Beaufort contact occurs some 70 to 120m above the presently mapped contact. Fossils collected during this and previous studies show that biostratigraphically Tapinocephalus Assemblage Zone fauna occurs in the rocks of the Koonap Formation in this part of the basin, indicating that the Ecca-Beaufort contact in the southern part of the basin youngs towards the east. This supports the prograding shoreline deposition model that has been previously proposed for the Ecca-Beaufort contact.