Goniopholididae is a distinctive and common clade of crocodyliforms known from the Jurasic and Cretaceous of Europe, Asia, and North America. Their position within Neosuchia makes them makes them very important to the study of crocodyliform evolution, but unfortunately their phylogenetic status is ambiguous. This study describes the osteology of two North American fossil taxa: Amphicotylus stovalli, a previously under-represented goniopholidid from the Jurassic Morrison Formation of Oklahoma, and new material of Denazinosuchus kirtlandicus from the Cretaceous Kirtland Formation of New Mexico. A phylogenetic dataset is then constructed including a full sampling of Jurassic North American forms and incorporating new new goniopholidid taxa not previously treated in a phylogenetic context. The affinity of North American and European members of Goniopholis is assessed as are the taxonomic status of putitive goniopholidids Vetisuchus and Denazinosuchus. Goniopholididae is monophyletic, and North American goniopholidids form a nested clade united by a distinctive palate morphology. Furthermore, all Morrison Formation goniopholidids are further nested in the North American clade. North American "Goniopholis" are generically distinct from European forms, and should be ascribed to Amphicotylus. Amphicotylus gilmorei is a junior synonym of A. lucasii. Denazinosuchus and Vectisuchus are not constituents of Goniopholididae, and instead are more closely related to pholidosaurs and thalattosuchians.

No description available.

Macrofossils from a Mesozoic fossil flora from Ellesmere Island, North West Territories are described and their affinities discussed, Several rarely occurring genera and species ore discussed in detail. The deposit in which they occur is dated using the known ranges of identified macro fossils and miospores, rand the results compared with those of other workers. The described flora is compared with floras of similar age in other parts of the world. / Thesis / Master of Science (MS)

A well preserved, fossiliferous Middle Triassic to Early Cretaceous section lies on the west side of Harrison Lake in the southern Coast Mountains. The study of this area involves a re-evaluation of the stratigraphic nomenclature first described by Crickmay (1925, 1930a) together with a lithologic description of the units and age determinations based on collected, identified and described fossils by the writer. Discussions on the biostratigraphy, paleogeography, regional correlations and structure of the thesis area and an overview of the regional tectonics of southwestern British Columbia and northwestern Washington, help to better understand the relation of this Mesozoic section to other rock assemblages in this geologically complex region. The oldest unit, the Middle Triassic Camp Cove Formation, comprises conglomeratic sandstone, siltstone and minor volcanic rock. Unconformable7 overlying this unit is the Toarcian to Early(?) Bajocian Harrison Lake Formation, divided into four distinct members by the writer, Celia Cove Member (basal conglomerate), West Road Member (siltstone, shale), Weaver Lake Member (flows, pyroclastic rocks, minor sediments) and Echo Island Member (interbedded tuff, siltstone, sandstone). Thickness of this formation is estimated at 3000 m. A hiatus probably is present between this unit and overlying shale, siltstone and sandstone of the Early Callovian Mysterious Creek Formation which is 700 m thick. Conformably above this are 230 m of sandstone and volcaniclastic rock of the Early Oxfordian Billhook Creek Formation. Late Jurassic fluvial conglomerate, sandstone and siltstone of the Kent Formation, perhaps 1000 m thick south of Harrison River, unconformably(?) overlies the last two units mentioned. Berriasian to Valanginian conglomerate and sandstone, 218 m thick, of the Peninsula Formation overlies the Billhook Creek Formation with slight angular unconformity. The Peninsula Formation is conformably overlain by tuffaceous sandstone, volcanic conglomerate, crystal tuff and flows of the Valanginian to Middle Albian Brokenback Hill Formation which is several km thick. Nine Jurassic ammonite genera are identified and described in this report. Triassic radiolaria and conpdonts and Cretaceous ammonites and bivalves are also present in the section. The most significant structure in the thesis area is the post-Albian to pre-Late Eocene Harrison Fault which strikes north-northwest through Harrison Lake, separating the Mesozoic section along the west side from the northern extension of the Cascade Metamorphic Core on the east side of the lake. A strong sub-horizontal stretching lineation within the fault zone may indicate right-lateral strike-slip movement. / Science, Faculty of / Earth, Ocean and Atmospheric Sciences, Department of / Graduate

published_or_final_version / abstract / Earth Sciences / Master / Master of Philosophy

published_or_final_version / abstract / Earth Sciences / Master / Master of Philosophy

published_or_final_version / Earth Sciences / Doctoral / Doctor of Philosophy

The first part of this dissertation deals with the timing of Mesozoic igneous activity in southern Cordillera of North America and its tectonic implications. A representative section in Santa Rita Mountains is dated using the zircon U-Th-Pb isotopic method. The oldest unit, the lower member of the Mt. Wrightson Formation, is concordantly dated at 210 ± 3 Ma. Initial basaltic andesite to andesite volcanism was followed by deposition of redbeds and associated volcanic rocks that are dated at 200 Ma. Felsic volcanism and eolian sand deposition may have spanned from 190 to 170 Ma. The Piper Gulch Granodiorite, representing the earliest Mesozoic intrusive equivalent, gives concordant dates of 188 ± 2 Ma. A second cycle of andesite and rhyolitic volcanism and sedimentation is dated at 151 ± 5 Ma using the whole-rock Rb-Sr isotopic method. The Hovatter Volcanics in the Little Harquahala Mountains, southwestern Arizona is dated at 165 Ma. Whole-rock Rb-Sr isotopic method on the same rocks gives a coherent reset isochron of 70 ± 3 Ma (Appendix III). A new stratigraphic correlation is proposed based on the dating data. Tectonic models proposed by previous workers to account for what seemed to be the lack of Triassic volcanic rocks are not necessary. This part of the Cordillera was an uplifted arc terrane during the Early Mesozoic (Appendix II) and may have provided volcanic detritus to the Late Triassic Chinle Formation in the Colorado Plateau. The second part of the dissertation deals with magma evolution and crust modification during arc magmatism. Rocks in southeastern Arizona have ƭ(N)(d) values of -3.4 to -6.4, while rocks to the west have ƭ(N)(d) values ranging from -8.5 to -9.2. An ƭ(N)(d) value of +2 for a Jurassic basalt indicates the presence of depleted mantle under the arc. Using lower crust and mantle end-members, 20 to 40% mantle input is estimated. This seems to argue for continuous growth model of the continental crust. Combined REE and isotopic data indicate that assimilation of lower crust by mantle melts followed by fractional crystallization took place. Detailed study indicates that the lower crust along sites of arc magmatism gets progressively hybridized by the mantle, becoming more mantle-like with time.

The Cabullona Basin of northeastern Sonora is a structural depocenter that was formed during Late Cretaceous time. The 2.5 km-thick sedimentary fill of this basin, the Cabullona Group, is composed in ascending order of the Corral de Enmedio formation, Camas Sandstone, Packard Shale, Lomas Coloradas formation, and, laterally equivalent to the last two units, El Cemento conglomerate. Abundant vertebrate and invertebrate as well as pollen identifications from these units indicate the Cabullona Group is of middle Santonian to Maastrichtian age. The Corral de Enmedio formation represents shallow lacustrine deposits. Lithofacies of the Camas and Lomas Coloradas formations indicate they were deposited by braided- and meandering-fluvial systems, whereas the eight members of the Packard Shale represent complex fluvio-deltaic-and-lacustrine systems. The El Cemento conglomerate is a thick clastic wedge of coarse-grained alluvial deposits that was deposited adjacent to the structural margin of the basin. Clast composition and paleocurrent directions of the El Cemento conglomerate indicate it was derived from strata of the nearby uplift of the Sierra Anibacachi-Cerro Cabullona. The low-angle, southwest directed Cabullona thrust fault that separates the uplift from the basin formed the structural margin of the basin. The tectono-sedimentary history of the Cabullona Group, its age and regional tectonic setting support the idea that this basin was formed because of typical Laramide-style deformation and indicates that the Rocky Mountain foreland province of Laramide deformation extended southward to northeastern Sonora.

Troodontidae is an important dinosaur taxon that closely resembles birds in both morphology and biology. The evolution of troodontids is crucial for understanding evolutionary transitions between non-avialan theropods and avialans. Despite the recent discovery of several troodontid taxa across the world and many new studies of coelurosaurian relationships, an overall survey of morphological variation in troodontids and a comprehensive analysis of ingroup troodontid relationships have yet to be accomplished. In the first four chapters of this dissertation, the osteology of two new troodontid taxa and two closely related paravians are described in detail. These descriptions are based on new specimens recovered from the Mesozoic of China and Mongolia. These new taxa include the basal dromaeosaurid Microraptor zhaoianus, the basal avialan Anchiornis huxleyi, a new troodontid taxon represented by IGM 100/1323, and a second new troodontid taxon represented by IGM 100/1126 and IGM 100/3500. These paravian taxa are all small-sized, with a basal paravian body plan resembling Archaeopteryx, yet they represent members of all three major paravian lineages (Troodontidae, Dromaeosauridae and Avialae), and support the traditionally recognized paravian interrelationships. Osteological description of Microraptor zhaoianus is based on an excellently preserved new specimen BMNHC PH881. This specimen preserves significant morphological details that are not present, or are poorly preserved, in the other Microraptor specimens, including aspects of the skull, rib cage, and humerus. These new characters corroborate Microraptor as a member of the Dromaeosauridae and support the close relationship of troodontids with dromaeosaurids. Four new specimens (PKUVP 1068; BMNHC PH804, BMNHC PH822 and BMNHC PH823) of Anchiornis huxleyi reveal new osteological details of this important paravian taxon. Anchiornis huxleyi shares derived features with avialans, but it lacks derived deinonychosaurian characteristics such as a laterally exposed splenial and a specialized raptorial pedal digit II. IGM 100/1323 represents a new troodontid taxon from the Late Cretaceous Djadokhta Formation of Mongolia, diagnosed from other troodontids by the absence of the lateral groove on the dentary, a posteriorly curved pterygoid flange, a distinct spike-like process on the ischium, and elongate chevrons. Despite generally having a basal paravian body plan, IGM 100/1323 displays many derived troodontid features. IGM 100/1126 and IGM 100/3500 represent another new Late Cretaceous troodontid taxon from the Djadokhta-Formation-like rocks at Ukhaa Tolgod, Mongolia. It is unique and distinct from other troodontids in having closely packed peg-like teeth, a twisted suborbital process of the jugal, a quadratojugal with a crescentic ascending process that braces the quadrate posteriorly, reduction of the basal tubera, and presence of a posterior fossa on the proximal fibula. This new taxon is morphologically more derived than Early Cretaceous troodontids but is more primitive than other Late Cretaceous troodontids. A new and comprehensive phylogenetic analysis of coelurosaurian theropods, focusing on troodontids is presented in Chapter 5. This is an updated version of the Theropod Working Group (TWiG) analysis (2015.1). This new analysis incorporates new paravian taxa and new characters, most of which are relevant to paravians, especially the troodontids that are the focus of this dissertation. The new phylogenetic analysis agrees with previous studies on the general relationships of coelurosaurians, yet some important differences from previous TWiG analyses are present in paravians, including: 1), the Jianchang paravians are recovered as basal avialans; 2), Late Cretaceous troodontids form a monophyletic group; and 3), Jinfengopteryginae is not monophyletic.