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Old-growth stands are important for management, conservation, wildlife, recreation, and maintaining biological diversity in forested landscapes. However, we are lacking the information needed to adequately identify and characterize old-growth stands. This is especially true for high elevation, interior forests. The characterization of stand structure and regeneration pattern will help in the development of site-specific guidelines for identifying old growth stands and restoring some of the old-growth characteristics in managed stands. This pamphlet presents a synopsis of a study investigating stand structure and regeneration of old-growth stands in the Moist Cold Engelmann Spruce - Subalpine Fir (ESSFmc) Subzone near Smithers, B.C. The three stands selected for the study were located on zonal sites, each in different watersheds, and the stands were established after fire. The criteria used for selection were: i) absence of lodgepole pine, ii) presence of advanced regeneration, and iii) abundant snags and coarse woody debris. These stands were presumed to represent the old-growth stage of stand development or the final (climax) stage of secondary succession.

The species richness, abundance and composition of saproxylic Coleoptera was compared between 'old-growth' and mature-managed hemlock-hardwood forests in southeastern Ontario, Canada. Beetles were sampled weekly from 29 April until 3 October 2003 using large-area flight-intercept traps (FITs) and trunk-window traps (TTs). Analyses were conducted using the Fisher's alpha and Simpson's diversity indices, rarefaction, indicator species analysis and cluster analysis. A total of 11,888 fungivorous Coleoptera was collected from 11 families and 73 species (excluding Nitidulidae). Nitidulidae were analysed separately with traps yielding 2,129 sap beetles comprising 30 species. The species richness and abundance of fungivorous Coleoptera did not differ significantly between the two forest types. Conversely, the species abundance of nitidulid beetles was higher in managed forests and the species richness higher in 'old-growth' forests. Several species were strongly associated with either managed or 'old-growth' forest types. Certain species such as Anisotoma inops (Leiodidae) and Glischrochilus sanguinolentus (Nitidulidae) were much more frequently caught in TTs than in FITs.

I examined saproxylic beetle responses in two silvicultural systems of the eastern mixedwood boreal forest of Quebec. I first investigated habitat-use and aspen-host use of saproxylic and bark and wood-boring beetles in remnant forest patches (cut-bock separators and small patches) left after harvest, theorized to resemble natural post-fire residual trees and snags. A second study focused on effects of partial cutting, a method that may serve to imitate natural succession dynamics. / Remnant forest patches had the highest saproxylic and bark and woodboring beetle species richness and relative abundance. Although non-significant, higher larval densities were also collected from remnant forest patches. In the second study, partial cut patches had an intermediate saproxylic beetle assemblage compared to uncut forest and clearcuts. / This research has brought new information on the effects of alternative harvesting practices on saproxylic beetles, supporting the hypothesis that biodiversity is best preserved based on forest management that is diversified and based on natural disturbance dynamics.

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This research examines the relationship between understory plant diversity and logs in a Pacific Northwest (PNW) Sitka spruce (Picea sitchensis)-western hemlock (Tsuga heterophylla) old-growth, coastal forest. These forests are renowned for their high forest productivity, frequent wind storms, and slow log decomposition rates that produce unmatched accumulations of coarse woody debris (CWD) yet few studies have examined the relationship between CWD and understory vegetation ecology. My research addressed this topic by comparing understory plant census data between paired fallen log and forest floor sites (n=20 pairs). My objectives were to: 1) determine the influence of substrate type on community composition and diversity, and 2) examine successional pathways and species assemblage patterns on CWD in various stages of decomposition. To meet these objectives I employed non-metric multidimensional scaling (NMDS) ordinations and unsupervised cluster analyses to identify and compare community assemblages on both substrates. These methods revealed similar species diversity and evenness between log and forest floor sites with compositional differences within and between substrates corresponding to habitat availability for colonization and light and moisture gradients. My results also suggest understory successional pathways related to decay class and characterized by an initial abundance of bryophytes, forbs, and seedlings followed by woody shrubs. Understory communities developing on logs also experienced increasing diversity, evenness, and divergence from forest floor communities consistent with log decomposition. These results differ from findings for boreal forests that reveal increasing similarity between substrate communities with increasing decay class. Recommendations for future research include the employment of a more robust sample size and direct measurements of environmental variables. Additional comparator studies are also needed to confirm the effects of forest type and decomposition on the relationship between CWD and forest understory communities. This study demonstrates how fine-scale wind disturbance fosters biodiversity through the creation of CWD substrate. My results and future research are essential for the development of silvicultural models designed to promote biodiversity in PNW coastal forests.

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Mat-forming ectomycorrhizal (EcM) fungi represent a prevalent constituent of many temperate forest ecosystems and create dramatic changes in soil structure and chemistry. EcM mat soil have been shown to have increased microbial respiration rates and have been hypothesized to harbor unique assemblages of fungi and bacteria. The objectives of this dissertation were to characterize and examine the fungal and bacterial communities associated with EcM mats in old-growth forests of the H.J. Andrews Experimental Forest located in the Oregon Cascades. Additionally, this work assessed the application of traditional, emerging, and novel molecular sampling techniques for determining microbial communities of environmental samples. This research investigated the microbial communities associated with two common EcM mat genera found in old-growth Douglas fir stands in the Pacific Northwest; Piloderma (Atheliales, Basidiomycota) and Ramaria (Gomphales, Basidiomycota). Soil samples were collected from Piloderma and Ramaria mats and surrounding non- mat soil for molecular analysis of nucleic acids. First, a comparative study was conducted to determine the most appropriate rDNA molecular sampling technique for microbial community characterization. Two next-generation sequencing methods, Roche 454 pyrosequencing and Illumina-based environmental sequencing, the latter developed by the author, were compared to a more traditional sequencing approach, i.e., Sanger sequencing of clone libraries. These findings informed the subsequent sampling of the fungal ITS and bacterial 16S rDNA fragment with 454 pyrosequencing to determine the microbial communities within mat and non-mat soils. Second, this work utilized a pyrosequencing approach to explore fungal community structure in EcM mat and non-mat soils. This work concluded that differences in microbial communities do exist between Piloderma mat, Ramaria mat, and non-mat soils, but the differences are largely quantitative with relatively few distinct taxonomic shifts in microbial constituents. Piloderma, Ramaria and Russula, in addition to being the dominant taxa found on mycorrhizal root tips, were found to be the most abundant taxa in bulk soils within their respective mat types or non-mat sample. The background fungal communities within the EcM mats in this study exhibited considerable taxonomic overlap with the exception of Piloderma vs. non-mat comparisons; Russula species dominated nonmat soils but tended to be excluded or significantly underrepresented in Piloderma mats. Lastly, this study explored the bacterial communities associated with Piloderma and Ramaria mats using lower- coverage 454-Jr pyrosequencing. Bacterial communities exhibited significant structure as a function of mat-type, soil horizon and pH, but this finding should be interpreted with respect to the nonrandom distribution of Piloderma-mats in the O- horizon and the Ramaria-mats in the A-horizon, and the tendancy for EcM mats to be more acidic than surrounding soils. Nonetheless, the total microbial (bacterial and fungal) community was typically dominated by the mat-forming taxa, or Russula, in the case of non-mat soils. While the presence of Piloderma mats did enrich or restrict some bacterial groups, soil pH was also found to be a significant driver of bacterial richness and taxonomic diversity. Fungal and bacterial richness were also found to be positively related to one another, regardless of soil horizon or EcM mat type. This work, taken together, contributes to the understanding of hyperdiversity and heterogeneity of microbial communities of temperate forest soils and highlights the potential for fungal and bacterial communities to be influenced by the presence of EcM mats. / Graduation date: 2012

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