When making decisions on which areas to harvest in a sustained yield, even-flow manner in mountainous areas such those in coastal British Columbia, it is important to know how timber productivity changes with elevation. This information allows foresters to decide at what elevation to start increasing the rotation age and to decide at what elevation sustainable harvesting becomes infeasible due to low productivity. Since Pacific silver fir (Abies amabilis Dougl. ex Forbes) has an elevation range that extends from sea level nearly to the tree line (0 m to approximately 1,650 m; from the Coastal Western Hemlock zone, through the Mountain Hemlock zone; to the lower limits of the Alpine Tundra zone), productivity-elevation relationships are especially important. To acquire quantitative measures of productivity decrease with increasing elevation a regression equation relating site index (the height of the dominant trees at a base age of breast height age of 50 years) to elevation in southern coastal BC was developed. In turn, we used this regression as an input into the height driven yield model named the Variable Density Yield Prediction model (VDYP). The use of the VDYP model allows the site index values to be translated into actual productivity measures (e.g., volume per hectare, mean annual increment at culmination age).

Site classification in the biogeoclimatic ecosystem classification system is based on three differentiating properties: climatic regimes (expressed by biogeoclimatic subzones or variants), soil moisture regimes (SMRs), and soil nutrient regimes (SNRs). A SNR represents a segment of a regional soil nutrient gradient, i.e., soils which provide similar levels of plant-available nutrients over a long period. SNRs are identified in the field using a number of easily observable soil morphological properties and indicator plant species. However, we need to know to what extent soil nutrient properties support these indirect field-estimates. There have been several studies that quantitatively characterize regional soil nutrient gradients in different climatic regions, but no study has yet been done in the subalpine coastal forest (Mountain Hemlock zone). Influenced by a maritime subalpine boreal climate, high-elevation coastal soils differ from low-elevation soils by having a thicker forest floor and a higher organic matter content. In the study summarized here, relationships between soil chemical properties and field-estimated SNRs are examined and soil chemical properties and field-identified SNRs are related to the site index of Pacific silver fir (Abies amabilis (Dougl. ex Loud.) Forbes) - one of the major timber crop species in the Coastal Western Hemlock and Mountain Hemlock zones.

Growth release of trees following canopy disturbances is of interest to ecological scientists and forest managers. Using dendroecological techniques, I examined growth release of canopy and subcanopy trees following the formation of natural, fine-scale canopy gaps in old-growth, western red cedar-western hemlock forests of coastal British Columbia. I aimed to quantify detailed information on release of the three shade-tolerant tree species that constitute these stands: western red cedar (Thuja plicata), western hemlock (Tsuga heterophylla), and Pacific silver fir (Abies amabilis). As a first step, I calibrated the radial-growth averaging method to account for regional-scale variability and capture a more complete range of growth releases that may occur following the formation of fine-scale gaps in the study stands. A 25% threshold, 5-year moving average, and 10-year window emerged as appropriate parameters for detecting releases using radial-growth averaging. Basal area increment was also the most appropriate growth index for detecting releases. Establishing these empirically-based criteria was important for quantifying the magnitude and duration of releases. Tree diameter and growth rate prior to release were the most important predictors of the magnitude and duration of releases, but identity of the tree species and distance from the gap center were also important predictors. Western hemlock and Pacific silver fir were often growing slowly both in the canopy and subcanopy, giving them tremendous potential to release. For these species, releases were generally intensive and persistent. In contrast, western red cedar were often growing quickly both in the canopy and subcanopy, giving them less potential to release. Compared to western hemlock and Pacific silver fir, western red cedar releases were less intensive and persistent. Patterns related to distance from the gap center emerged for trees growing along the north-south axis of gaps. Regardless of species, increasing distance from the gap center resulted in decreasing magnitude and duration of releases. However, patterns for duration were complex, as the distance effect was greater for trees north of the gap center. Information on growth release of trees is useful for reconstructing the history of past canopy disturbances, elucidating mechanisms of tree species coexistence, and assessing and predicting stand changes due to forest management in coastal British Columbia.

Growth release of trees following canopy disturbances is of interest to ecological scientists and forest managers. Using dendroecological techniques, I examined growth release of canopy and subcanopy trees following the formation of natural, fine-scale canopy gaps in old-growth, western red cedar-western hemlock forests of coastal British Columbia. I aimed to quantify detailed information on release of the three shade-tolerant tree species that constitute these stands: western red cedar (Thuja plicata), western hemlock (Tsuga heterophylla), and Pacific silver fir (Abies amabilis). As a first step, I calibrated the radial-growth averaging method to account for regional-scale variability and capture a more complete range of growth releases that may occur following the formation of fine-scale gaps in the study stands. A 25% threshold, 5-year moving average, and 10-year window emerged as appropriate parameters for detecting releases using radial-growth averaging. Basal area increment was also the most appropriate growth index for detecting releases. Establishing these empirically-based criteria was important for quantifying the magnitude and duration of releases. Tree diameter and growth rate prior to release were the most important predictors of the magnitude and duration of releases, but identity of the tree species and distance from the gap center were also important predictors. Western hemlock and Pacific silver fir were often growing slowly both in the canopy and subcanopy, giving them tremendous potential to release. For these species, releases were generally intensive and persistent. In contrast, western red cedar were often growing quickly both in the canopy and subcanopy, giving them less potential to release. Compared to western hemlock and Pacific silver fir, western red cedar releases were less intensive and persistent. Patterns related to distance from the gap center emerged for trees growing along the north-south axis of gaps. Regardless of species, increasing distance from the gap center resulted in decreasing magnitude and duration of releases. However, patterns for duration were complex, as the distance effect was greater for trees north of the gap center. Information on growth release of trees is useful for reconstructing the history of past canopy disturbances, elucidating mechanisms of tree species coexistence, and assessing and predicting stand changes due to forest management in coastal British Columbia.

Growth release of trees following canopy disturbances is of interest to ecological scientists and forest managers. Using dendroecological techniques, I examined growth release of canopy and subcanopy trees following the formation of natural, fine-scale canopy gaps in old-growth, western red cedar-western hemlock forests of coastal British Columbia. I aimed to quantify detailed information on release of the three shade-tolerant tree species that constitute these stands: western red cedar (Thuja plicata), western hemlock (Tsuga heterophylla), and Pacific silver fir (Abies amabilis). As a first step, I calibrated the radial-growth averaging method to account for regional-scale variability and capture a more complete range of growth releases that may occur following the formation of fine-scale gaps in the study stands. A 25% threshold, 5-year moving average, and 10-year window emerged as appropriate parameters for detecting releases using radial-growth averaging. Basal area increment was also the most appropriate growth index for detecting releases. Establishing these empirically-based criteria was important for quantifying the magnitude and duration of releases. Tree diameter and growth rate prior to release were the most important predictors of the magnitude and duration of releases, but identity of the tree species and distance from the gap center were also important predictors. Western hemlock and Pacific silver fir were often growing slowly both in the canopy and subcanopy, giving them tremendous potential to release. For these species, releases were generally intensive and persistent. In contrast, western red cedar were often growing quickly both in the canopy and subcanopy, giving them less potential to release. Compared to western hemlock and Pacific silver fir, western red cedar releases were less intensive and persistent. Patterns related to distance from the gap center emerged for trees growing along the north-south axis of gaps. Regardless of species, increasing distance from the gap center resulted in decreasing magnitude and duration of releases. However, patterns for duration were complex, as the distance effect was greater for trees north of the gap center. Information on growth release of trees is useful for reconstructing the history of past canopy disturbances, elucidating mechanisms of tree species coexistence, and assessing and predicting stand changes due to forest management in coastal British Columbia. / Arts, Faculty of / Geography, Department of / Graduate

In the wetter climates associated with the coastal forests of northwestern North America, coarse woody debris (CWD) accumulations in the form of snags, downed boles, and large branches can be large in natural forest ecosystems. Seedlings often regenerate on stumps and downed logs in the understory of old-growth coastal forests. The question remains though, whether CWD is a necessary component for seedling survival and growth in forests managed for commodity production. This study addresses one concern of forest managers: is there an immediate nutritional or moisture supply advantage conferred by CWD for the survival and growth of seedlings in the coastal climate of British Columbia? We compared survival and growth of seedlings planted in decaying wood compared to non-woody humus forms and mineral soil under heavy shade and full light conditions. Low light environments are of particular interest since reports of the strong association between CWD and regeneration has primarily referred to understory seedlings and saplings in old-growth forests.

The Mountain Hemlock (MH) zone includes all subalpine forests along British Columbia’s coast. It occurs at elevations where most precipitation falls as snow and the growing season is less than 4 months long. The zone includes the continuous forest of the forested subzones and the tree islands of the parkland subzones (Figure 1). Old-growth stands are populated by mountain hemlock, Pacific silver fir, and Alaska yellow-cedar, and are among the least-disturbed ecosystems in the world. Canopy trees grow slowly and are commonly older than 600 years, while some Alaska yellow-cedars may be up to 2000 years old. Early regeneration failures followed slashburning and the planting of unsuitable species. Currently, the most successful and feasible option for reforesting cutovers is natural regeneration with a mix of the three main tree species, but uncertainties remain about the temporal and spatial pattern of regeneration, changes in species composition, and the time required for stand establishment after cutting. Our study addressed these concerns by examining regeneration patterns on 6 sites that were clearcut 11-12 years prior to sampling and left to regenerate naturally. The sites were located at the lower limits of the zone in the Tetrahedron Range, near Sechelt, at elevations from 1060-1100m.

Ecosystem-specific forest management requires comprehension of tree species productivity in managed settings, and how this productivity varies with the ecological determinants of site quality, i.e., the environmental factors that directly affect the growth of plants: light, heat, soil moisture, soil nutrients, and soil aeration. A good understanding of this variation is necessary for making species- and site-specific silvicultural decisions to maximize productivity. Productivity of a given species is usually measured by site index (tree height at 50 years at breast height age). Quantitative relationships between site index and these measures of site quality provide predictive models for estimating site index. Pacific silver fir (Abies amabilis (Dougl. ex Loud.) Forbes) is an important timber crop species in the coastal forests of British Columbia. In relation to climate, its range in southwestern British Columbia extends from sea level to almost timberline, and from the hypermaritime region on western Vancouver Island to the subcontinental region on the leeward side of the Coast Mountains. In relation to soils, its range extends from slightly dry to wet sites and from very poor to very rich sites. In view of this relatively wide climatic amplitude, a large variability in productivity can be expected. It is particularly important to consider the growth performance of Pacific silver fir when decisions are made regarding whether or not to cut stands on high-elevation sites. In the study summarized here, relationships between Pacific silver fir site index and selected ecological measures of site quality were examined, and site index models using these measures as predictors were developed.

This study examines the influence of climate on tree-ring properties of several major tree species: Pacific silver fir (Abies amabilis (Dougl. ex Loud.) Forbes), subalpine fir (Abies lasiocarpa (Hook.) Nutt.), Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), and lodgepole pine (Pinus contorta var. latifolia Dougl. ex Loud.). Our three objectives were to determine how (1) tree-ring properties change along an elevation gradient, (2) short-term climatic influences are correlated with tree-ring properties, and (3) long-term climatic influence on tree-ring properties.

The Mountain Hemlock (MH) zone includes all subalpine forests along British Columbia’s coast. It occurs at elevations where most precipitation falls as snow and the growing season is less than 4 months long. The zone includes the continuous forest of the forested subzones and the tree islands of the parkland subzones (Figure 1). Old-growth stands are populated by mountain hemlock, Pacific silver fir, and Alaska yellow-cedar, and are among the least-disturbed ecosystems in the world. Canopy trees grow slowly and are commonly older than 600 years, while some Alaska yellow-cedars may be up to 2000 years old. Understanding regeneration patterns in the MH zone has become increasingly important as logging continues towards higher elevations of the zone where snowpacks are deeper.