Graduation date: 2002

No description available.

The South African grassland biome is disappearing rapidly through advancing development and change in agricultural land use. One of the most threatened grassland types, Midlands Mistbelt, in the KwaZulu-Natal Midlands is an extremely diverse and home to many endemic species across an array of taxa. Three taxa, namely, grasses, grasshoppers and butterflies represent various trophic levels, which are important to the functioning of the grasslands. Ten grasslands were sampled by walking ten fifty metre transects for a twelve-month period. The grasslands were selected as they represented a range of management practices and varying environmental conditions. Using Indicator Species Analysis (ISA) twenty-two species of grasshopper were identified as indicators of environmental variables and management practices. The abundances of the various species indicated the intensity of the management regimes or disturbances. Using the twenty-two grasshopper species abundances and a three hundred point sampling assessment of the grasses creates an assessment tool that can rapidly appraise the management of the grassland, but due to lack of data for other taxa, cannot assess whether management practices for the focal taxa create congruent results for non-focal taxa. Two of the three taxa proved to be good indicators of grassland health, whilst the third, butterflies were ineffectual, due to low abundance and richness. From the results it was concluded that burning was taking place to frequently, and required a reduction to every four years, as this would improve butterfly richness and abundance, and increase abundance of endemic and flightless grasshopper speCies. A rotational grazing system needs to be implemented at sites where continual grazing takes place, wildlife or livestock, impacts on the grassland condition and species diversity. Increasing habitat heterogeneity increases species diversity, and allows later successional species to be included in the grasshopper assemblage. Management of the grasslands in the KwaZulu-Natal Midlands needs to be more responsive and adaptive. In addition, small fragment management needs to be intensified to provide a range of habitats and refugia that will suit all species. This study advocates the use of grasshoppers and grasses as suitable biotic indicators of grasslands in the KwaZuluNatal Midlands. / Thesis (Ph.D.) - University of KwaZulu-Natal, Pietermaritzburg, 2005

Plants are the major source of food or energy required to sustain life on the planet, but humans are grappling with the deteriorating conditions of natural ecosystems such as compositional change, desertification, invasive plants and soil erosion. In the face of global climate change and growing demands for agricultural productivity, future pressures on grassland ecosystems will intensify, therefore sustainable utilization of all plant resources is of vital importance to enhance food security within the limits of good conservation. The semi-arid grasslands of southern Africa represent major grassland resources for grazing. Herbage production in these areas is determined not only by water and nutrient availability, but also by controlled and uncontrolled fires. Since fire is regarded as a natural factor in savannas, it is essential to develop a deeper understanding of the role of fire in community structure and function for the development of appropriate burning regimes. A study was conducted in the Eastern Cape of South Africa where the rural communities are faced with the challenges of rangeland degradation in the form of encroachment by unacceptable bush, karroid, macchia and less desirable grass species, as well as soil erosion. The main objective of this thesis was to investigate the roles of competition and disturbance regimes (fire and simulated non-selective grazing) on species composition, habitat productivity and the performances of selected species from this semi-arid savanna. Long-term effects of burning frequency on herbaceous species composition, Leaf Area Index (LAI), Photosynthetically Active Radiation (PAR) within the herbaceous canopy, biomass production and soil chemical properties were investigated. These studies were conducted on a fire trial set up in 1980 at the University of Fort Hare research farm in the Eastern Cape, South Africa. The treatments comprise an annual, biennial, triennial, quadrennial, sexennial and no burn control, all replicated twice in a Complete Randomized Design. The data from the trial collected between 1980 and 2008 were used to determine compositional variation for herbaceous species using the Non-metric Multidimensional Scaling and Bray-Curtis Dissimilarity tests. The PAR ceptometer was used to determine LAI and intercepted PAR, while random samples were harvested from 1m² quadrats from each plot. Soil samples were taken at four depths (0-2 cm; 2-4 cm; 4-6 cm and 6-8 cm) from each plot and analyzed for pH, Ca, K, P, total C and total N. The Resin-Bag technique was used to determine nitrogen mineralization. Burning frequency caused significant variation in herbaceous species composition over time. The species were distributed along gradients of increasing burning frequency, and these responses were in three categories: Those that increased with burning frequency such as Themeda triandra; those that decreased with burning frequency such as Melica decumbens, and those that showed little response such as Panicum maximum. The three-year burn resulted in the highest compositional variation, light interception, Leaf Area Index, aboveground biomass production, while the annual, biennial and no burn treatments resulted in the lowest. The fact that infrequent burning resulted in higher species variation, improved habitat productivity due to increased leaf area for light interception shows that appropriate use of fire can maintain a more diverse and productive savanna system. Burning frequency had significant effects on the soil properties, while soil depth did not show any significance. Frequent burning increased soil pH, K, Ca, and Na, but reduced C, N, P and N mineralization. There was a negative correlation between burning frequency and N mineralization, but no correlation existed between N mineralization and total N, total C or the C:N ratio. These results imply that frequent burning can cause nutrient losses and a greater nutrient limitation to plants in the long-term, especially soil C and N loss from combustion of organic material in the soil top layer. The ability of shade-tolerant plants to persist under shade and regular defoliation such as in burnt and grazed systems may be of greater importance for long-term productivity and sustainability of forage crops. It is therefore imperative to explore the mechanisms by which some species were favoured by frequent burning which created low shade conditions, while others were favoured by high shade conditions where burning is infrequent or absent. A pot experiment was conducted to investigate the shade tolerances of seven grass species that were abundant in the long-term fire trial. The test species were Cymbopogon plurinodis, Digitaria eriantha, Eragrostis curvula, Melica decumbens, Panicum maximum, Sporobolus fimbriatus and Themeda triandra. Individual grass tillers of each species were collected from the natural vegetation, propagated in separate seedling trays and transplanted into individual pots, and were grown under five shading treatments: full sun (0 % shading), 55 %; 70 %; 85 % and 93 % shading respectively. Shading significantly reduced the dry matter production of all the species. Biomass production of all the species decreased linearly to varying degrees with an increase in shade intensity. Digitaria eriantha and Eragrostis curvula were most adversely affected by shading, hence are classified as shade intolerant, while Melica decumbens was the least affected by shading, and is hence classified as shade tolerant. Cymbopogon plurinodis, Panicum maximum, Sporobolus fimbriatus and Themeda triandra are classified as moderately shade-tolerant. From the results it was apparent that some species could perform optimally in partial shade than in full sunlight, and these results lead to a conclusion that for satisfactory natural regeneration and seedling growth of this savanna vegetation would require a gap large enough to provide at least 30 % of ambient light. Investigating patterns in competitive effects and responses of species in these communities may not only explain the abundance of each species, but may also provide insight into the nature of forces that affect the structure and function of that community. Since fire, herbivory and soil nutrients are natural drivers of savanna community structure and function, their influence on competitive interactions of selected species were investigated. Two experiments were conducted to investigate the competitive effects and responses of eight selected common species in the area. The test species (phytometers) included one woody shrub, Acacia karroo and seven grass species namely: Cymbopogon plurinodis, Digitaria eriantha, Eragrostis curvula, Melica decumbens, Panicum maximum, Sporobolus fimbriatus and Themeda triandra. In an outdoor plot experiment the responses of the phytometers to competition from neighbours (0; 2 and eight neighbours respectively), fertility (fertilized, unfertilized) and clipping (clipping, no clipping) were investigated. The second comprised a pot experiment where the competitive effects of the species were investigated. Each species was grown under 3 levels of fertility (0 %; 50 % and 100 % Hoagland‘s solution) and clipping (clipping, no clipping) in pots filled with fine river sand and 4 neighbours. Competition intensity, soil fertility and clipping had significant effects on the biomass production of the phytometer species. Acacia karroo and Melica decumbens, exhibited the weakest competitive effects and responses, and incurred the highest mortalities after clipping and with 8 neighbours. Digitaria eriantha and Panicum maximum exhibited the strongest competitive effects and responses, especially in high fertility, and experienced the lowest mortalities. T.triandra exhibited stronger competitive effect after clipping in low fertility, while A. karroo and C. plurinodis exhibited stronger competitive effects in moderate (50 %) fertility. Cymbopogon plurinodis, Eragrostis curvula and Sporobolus fimbriatus ranked between these two extreme groups in terms of competitive effects and responses. Relative Competitive Interaction increased with soil fertility and number of neighbours in the absence of clipping. These results indicate that in general, taller or broad-leaved grass species outgrow the shorter ones, and this gives them a competitive advantage over light and soil resources. One of the range management objectives in the False Thornveld of the Eastern Cape is to promote the abundance of Themeda triandra, which is of high forage value and an indicator of rangeland that is in good condition. The general situation under livestock farming conditions in this area is that if the grass sward is optimally grazed and rested then there is a great potential for Themeda triandra to dominate.The results of the competition experiments indicated that the species exhibits strong competitive interaction, and also exhibited stronger competitive effect after clipping in low fertility. These results imply that it has a low response and a high effect, an attribute that would enhance its performance if it is moderately grazed or the area is burnt. The species is also moderately shade tolerant, and this may explain why it thrives in burnt and moderately grazed areas. These studies have demonstrated the important role that competition and disturbance in the form of fire and herbivory play in the maintenance of this savanna grassland. Through natural selection species are able to occupy different niches in the same area and coexist in a heterogeneous environment and minimize their chances of extinction. / Thesis (Ph.D.)-University of KwaZulu-Natal, Pietermaritzburg, 2012.

The purpose of this study was to develop and test a predictive understanding of the vegetation dynamics of the Lowveld of South Africa (30°35'E to 30°40'E and 24°00'S to 25°00'S). The study covered about 5000 km2 in Adjacent Private Protected Areas (APPA) adjoining the Kruger National Park (KNP). Data gathering (800 sites; 23 properties) commenced in 1989 and those recorded up to 2004 are reported here. The value, both ecological and economic, of the wildlife and tourism industry dependent on this savanna region is discussed in both historical and current perspectives. A range of land-use objectives and anthropogenic interventions were exposed. The properties ranged in size from 30 to 800 km2 and formed an effective and extensive manipulative experiment for investigating interaction of bush density, animal stocking, use of fire and landscape-scale processes. The first descriptive classification (at 1:250 000) of the area was developed using Inverse Distance Weighted interpolations. This confirms similar landscape/vegetation patterns in the KNP and Mocambique. The current mode of determining stocking density or carrying capacity was interrogated and indices suitable for complex multi-species systems developed. This was done in the context of equilibrial/disequilibrial paradigms. Application of the original indices resulted in drought-related declines in animal biomass of 4000 kg km2 over 20 years due to overestimation of carrying capacity. The model proposed here uses rainfall, animal type, biomass and vegetation parameters to determine stocking density for both coarse (regional) and ranch-specific scales. Principal driving determinants (rainfall, geology, soil type, tree density canopy cover, animal numbers, feeding classes and fire) of vegetation structure and their influence on the herbaceous layer were investigated. Groupings on ecological potential showed 'high' potential areas are less sensitive to animal impact than those classified as 'low' potential. Sustainability, embedded in a forward-looking component viz. Strategic Adaptive Management (SAM) with well-articulated endpoints viz. Thresholds of Potential Concern (TPCs) was used to study fluctuations in animal populations with Connochaetes taurinus (Blue wildebeest) as the case study. The TPC approach provides strong pointers for proactive management aimed at maintaining the system within bands defined by TPCs supporting operationally practical and periodically reviewed objectives. / Thesis (Ph.D.)-University of KwaZulu-Natal, Pietermaritzburg, 2005.

Aristida junciformis subsp. junciformis Trin. et Rupr. is an indigenous, densely tufted, evergreen perennial grass associated with the degeneration of grasslands over large areas of South Africa. More than two hundred thousand square kilometres of veld, c. 17% of the total land area of South Africa, contains A. junciformis. The aim of this study was to improve our understanding of the mechanisms by which this species invades and dominates grassland, specifically in this study, Southern Tall Grassveld. Aristida junciformis also has a low nutritional value resulting in a very low grazing value. The unpalatability of the plant is due to the high tensile strength of the leaves, whose hard, fibrous laminas are very difficult for a grazing animal to crop once their length attains 30 cm or more. This species lack of response to conventional grazing practices has often resulted in a grassland in which the carrying capacity has diminished to such an extent as to be virtually useless for grazing in a relatively short time. The extent of encroachment of veld by A. junciformis appears dependant on the frequency of disturbance and rest afforded to the veld. Annual burning and mowing maintained the species composition of A. junciformis at levels <10% whilst protection from fire, burning or grazing allows this species to dominate the herbaceous layer at levels approaching 90%. Aristida junciformis does not become moribund and is unlikely to die if left undefoliated. As the abundance of A. junciformis increases, veld deterioration gradually accelerates through increased selective grazing on remaining palatable species. Competition from adult A. junciformis plants increases the mortality of seedlings and constrains growth of surviving seedlings. Tiller production of Aristida junciformis seedlings declined from an average of 5.2 tillers per seedling in a no competition situation to 2.2 tillers per seedling when subjected to full competition. Tiller production of T. triandra seedlings decreased from 9.6 tillers per seedling free from competition to 3.3 tillers per seedling subject to full competition for resources. Once these seedlings have become adult plants they are avoided by grazing animals and grazing pressure on the remaining palatable species consequently increases. This allows the A. junciformis plants to increase their size and density in the sward. Once this density is sufficiently high, grass seedlings of either A. junciformis or more desirable grass species such as T. triandra are unable to establish, eventually resulting in a monospecific stand of A. junciformis if left undisturbed. Large amounts of caryopses are produced by A. junciformis - up to a 19 000 caryopses from a large mature plant (c. 38 000 caryopses/m²). Of these c. 40% is likely to be infertile but the remaining c. 60% viable caryopses are dispersed in a typically leptokurtic distribution, the number of caryopses dispersed rapidly declining within a 10 metre radius. A large proportion of the caryopses was trapped in surrounding foliage but in open swards caryopses had greater opportunity to be blown further distances than in a closed sward. The density of A. junciformis caryopses on the soil surface was positively correlated with the density of flowering adult plants in the area and varied from 400 caryopses/m² (density of parent plants c. 0.6 plants/m²) in less effected areas to 11000 caryopses/m² (density of adult plants c. 6 plants/m²) in severely encroached areas. The primary function of the three awns appears to be orientating the caryopsis correctly for in its descent from the parent plant to expedite germination. Caryopses orientated vertically with the awns uppermost exhibited the highest and most rapid germination (67%) compared to caryopses lying horizontally (35%) whilst only 1% of inverted caryopses germinated. Caryopses trapped in litter and effectively held off the soil surface failed to germinate. Removal of the glumes from A junciformis seed greatly enhanced the rate and overall germination of the seed except for inverted seed of which <1% germinated. The highest numbers of A. junciformis seedlings (32 seedlings/m²) were found in those areas with the highest density of caryopses on the soil surface. Despite the large amounts of caryopses produced, dispersed and landing in apparently suitable micro-sites for germination, comparatively few A. junciformis seedlings (n=992) were found and overall germination ranged between 1 % and 4% of the initial caryopses density on the soil surface. Seedling survival through winter was low with only 13% surviving to the following spring. The basal areas of A. junciformis increased overall by 66% whilst that of other grass species increased overall by only 3% in the time monitored. These results suggest that the primary method of encroachment of A. junciformis in the grass sward appears to be through vegetative expansion and not seedling recruitment. Frequent defoliation of the sward and avoidance of overgrazing to enhance the competitive abilities of palatable species and provide as high a fuel load as possible appear to be the most economically and logistically feasible ways to remove or at least inhibit A. junciformis veld encroachment at present. / Thesis (M.Sc.)-University of Natal, Pietermaritzburg, 1998.

The mountains of Lesotho form the catchments for the Lesotho Highlands Water Project (LHWP), which is presently under construction, and their condition will determine the longevity of the LHWP. The mountain rangelands also support an extensive livestock system. However, there is concern that grazing is negatively affecting the mountain vegetation to the detriment of both livestock production and catchment function. Therefore, the impact of environment and grazing on the vegetation was investigated to aid the development of management policy for the conservation of the grazing, floristic and water resources of the mountains. Vegetation surveys were conducted in the mountains in the east (Study Area 1: 2 625 - 3 350 m a.s.l.) and in the west (Study Area 2: 2 240 - 3 125 m a.s.l.). Indirect gradient analysis (IGA) and classification were used to investigate the influence of environment on vegetation pattern. Results of the IGA indicated that variation in species composition in the mountains is related primarily to topographic variation, in particular elevation and aspect. Five vegetation communities were identified in Study Area 1 and seven in Study Area 2. These communities occurred consistently in specific topographic positions in the landscape and were arranged along a temperate/subtropical grass species continuum which was associated with a gradient in elevation and aspect. In Study Area 1, the elevation boundary between the high-lying temperate grasslands and the lower subtropical grasslands corresponded with the generally recognised boundary between the Alpine and Subalpine vegetation belts (viz. c. 2 950 m a.s.l. on northerly aspects and c. 2 750 m a.s.l. on southerly aspects). This boundary was lower in Study Area 2 (viz. c. 2 800 m a.s.l. on northerly aspects and c. 2 300 m a.s.l. on southerly aspects). Vegetation-insolation relationships were investigated in Study Area 1 using a model for simulating solar radiation, temperature and potential evaporation patterns on sloping terrain (RADSLOPE). The spatial distribution of the identified vegetation communities and the ratio of temperate (C₃) and subtropical (C₄) grasses in the sward were related to solar irradiance patterns, as influenced by topography. Results suggest that exposure, which increases with altitude, is probably also an important determinant of vegetation pattern in the mountains. The influence of grazing on the vegetation was studied by examining changes in species composition and cover that were associated with gradients in grazing intensity that exist around cattleposts in the mountains. There was little evidence of a shift in species composition and cover under grazing in the Alpine Belt but there was an identifiable grazing gradient in the Subalpine belt. There, short dense grasslands, dominated by palatable species, degrade to a dwarf karroid shrubland with sparse cover under prolonged, intense grazing. The optimum position along the grazing gradient of the more abundant species was identified. It was proposed that the relative positions, or scores, of these species along the grazing gradient can be used in a weighted scoring procedure to provide an index for monitoring the response of the mountain vegetation to grazing. However, the species’ scores still require verification. The need for monitoring temporal changes in vegetation composition and cover in order to assess the possible effects of the LHWP and other development initiatives was noted. Such monitoring should be undertaken in conjunction with an overall programme to assess the dynamics of the socio-economy in the mountains. Therefore, interdisciplinary monitoring programmes are required to achieve this. These programmes should be focused in a few key study locations rather than spread over a wide area. / Thesis (M.Sc.)-University of KwaZulu-Natal, Pietermaritzburg.

Grazing lands in Eritrea are degraded due to decades of overstocking and consequent overgrazing. Since the rangelands are accessible to entire village communities, organisation and coordinated decision making regarding the management of these resources is not often achieved. Farmers are not motivated enough to make investments to improve a communally owned resource due to the prevailing common access grazing systems. A field survey was undertaken interviewing 12 farmers in the private access commercial and 80 farmers in the common access subsistence grazing using face to face interviews in the Barka and Arado cattle farming communities in four out of the six regions in Eritrea. Debub, Gash-Barka, S. Keih Bahri and Maekel regions were selected using stratified and simple random sampling methods. The regions were chosen based on various agroecological zones where the representatives of different grass species and the two most common cattle breeds in Eritrea (Arado and Barka) are found. The survey included the collection of data on village and household characteristics focusing on rangeland grazing management systems and additional sources of supplementary forage. The study uses several stages of analysis like principal component analysis accompanied by regression analysis together with descriptive statistics and ordination diagram. The commercial farmers addressed grazing constraints by investing in improved grazing through planting 258 ha per farmer of drought resistant seeds and 1767 vs. 8 cactus slices per farmer and covered 75% vs. 40% of forage requirements from grazing resources compared to the subsistence farmers, respectively, during 2002. These results were achieved because 78% of the commercial farmers adopted controlled stocking rates. In common access grazing, the costs of collective action to control cattle stocking rates are high, making imple mentation of stocking rate controls difficult. As a consequence, 65% of the subsistence farmers were forced to migrate their cattle looking for grazing forage in the dry season during the year. The outcomes of migration were evidenced by the results of severe overgrazing and degradation on the rangelands proximity to villages in Debub and Maekel regions and the populated area of Gash_Barka region. The increased number of animals resulting in high grazing pressure was the consequence of migration. Ten vs. six percent of mortality rates was reported for the subsistence systems compared to the commercial systems respectively. The lower results of milk yield, calving rates and off- take rate productivity indicated in the different stages of analysis for the subsistence farmers were the consequences of the lack of the adoption of controlled stocking rates primarily constrained by the migration. The Barka and Arado cattle farming systems are kept under common access grazing systems. Compared to the Arado cattle farming, the Barka cattle farming region had relatively better access to grazing forage. The better quality of grazing in this region is attributed to a naturally low stock density in the region. During 2002, the Barka cattle farming had 1087 vs. 721 Lit of milk yield, 63% vs. 53% of calving productivity and 9.3% vs. 10.9% of mortality rates than the Arado cattle farming regions respectively, due to access to a wider area of grazing lands and more labour inputs. The Barka cattle area farmers are agro pastoralists and usually focus on grazing dairy cattle farming than crop farming. They increased calving rate productivity and decreased mortality rates by increasing the proportion of lactating cows and decreasing the proportion of oxen compared to the Arado cattle farming. The Arado cattle farming had higher offtake rates and income from cattle sales compared to the Barka cattle farming region. The higher off- take rate, which is an index of percentage of cattle sold, for the Arado cattle was probably linked to the shortage of grazing forage and increased herding costs. The Barka and Arado cattle farmers had a shortage of quality and quantity crop residue winter forage during 2002. Farmers were dependent only on rain fed cropping. The application of crop rotation, fallow and chemical fertilizers were low to enhance soil nutrients. Out of the total crop residues forage produced, only 22% and 15% of legumes residue DM forage was produced for the Barka and Arado cattle farmers respectively. Agro- industrial and crop farming by-products supplementary feeds were also limited due to the shortage of feeds in the country during the year. In general, government intervention is important to bring institutional changes to promote the adoption of controlled stocking rates to alleviate the shortage of grazing forage. / Thesis (M.Sc.)-University of KwaZulu-Natal, Pietermaritzburg, 2007.

The overall aim of the study was to investigate re-vegetation of disturbed sites, using nursery grown plugs (from seedling trays) of Themeda triandra, Heteropogon contortus and Hyparrhenia dregeana in order to reach practical management guidelines for re-vegetation using indigenous grass plugs. A number of field trials were set up at Kamberg Nature Reserve (29°24'S, 29°40'E) on a site that was clear felled of A. mearnsii in October 1997. The trials were established in January 1998 and January 1999. A total of approximately 52 ,000 nursery raised plugs of T. triandra, H. contortus and H. dregeana were planted into an area of approximately 7,000 m2 . In the planting density trial , plugs of H. dregeana only and a combination of T. triandra/H. contortus were planted at 15 cm and 30 cm spacings. The T. triandra/H. contortus combination at 30 spacing showed the greatest survival and lateral plant growth (tiller number and basal area) and this combination is thus recommended. In the over-sowing trials, the H. dregeana and T. triandra/H. contortus combination at both 15 cm and 30 cm spacing were over-sown with E. curvula. The survival and lateral growth of the T. triandra/ H. contortus combination at 30 cm was again greater than the other treatments. Over-sowing with E. curvula suppressed the survival and lateral growth of the planted plugs across all treatments compared to not over-sowing. The over-sown conditions showed a significant decrease in the diversity of the plots, both in the number of species present and the Shannon diversity index. An area that had been cleared of A. mearnsii and sown to E. curvula 25 years previously was shown to have a lower number of species than the neighbouring veld. Nursery raised plugs of T. triandra were planted into the mature E. curvula in an attempt to improve the biodiversity of these areas. To re-introduce T. triandra into these E. curvula swards the plugs must be planted into the centre of a gap rather than around the base of an E. curvula plant. For improved survival of the plugs the E. curvula tufts must be clipped, while for best lateral growth the E. curvula tufts must be sprayed with a glyphosate herbicide three months prior to planting and clipping. However, the added expense of spraying and clipping is not warranted as the clipped treatments also showed good growth. Transplant shock is common when planting nursery raised plugs out into the field, as there is a relatively small root volume in the plug compared to the above ground leaf biomass. Alleviation of moisture stress at planting using a starch based polymer with high water holding capacity (Terrasorb®) and a white, needle punched geo-fabric (Agrilen®) to provide a seven day period of artificial shade after planting did not show significant improvements over the control with regards to survival or plant growth. Thus these methods of moisture amelioration are not recommended in revegetation through planting of plugs at this study site. A trial was established to investigate the biomass production of six different treatments to determine their potential to support a fire. The total biomass for the plots which were over-sown by E. tef and planted to only H. dregeana were on average sufficient for a fire, but there was a discontinuous fuel load across these plots, especially in the replications that had very low survival rates and thus these plots could not be burnt. The control and herbicide sprayed plots also showed sufficient fuel load for a fire, but this fuel load was made up of A. mearnsii saplings and bramble with very little grass cover and thus a fire would not have burnt through these plots either. The T. triandra/H. contortus combination did not produce sufficient fuel load, due to poor survival. Thus only the plots over-sown with E. curvula were able to burn in this trial and as a burning trial per se the trial was abandoned. Seed bearing hay (thatch) was collected in early summer (December 1997) and late summer (April 1998). Both times of year of harvesting proved to be successful in terms of grass cover, although the early harvested thatch had a greater number of species per plot. The Shannon diversity indexes of the two treatments were not significantly different. The multi-response permutation procedure technique confirmed that there was a compositional difference between the treatments. By the end of the trial Harpochloa falx and T. triandra and H. dregeana were indicators for the early and the late harvested thatch respectively. Comparing the thatching trial and the planting density trial indicated that the T. triandra/H. contortus combination at 30 cm spacing would be recommended to maximize biodiversity. The summer months have been shown to be the best time to plant the plugs, although the actual success will be dependant on the conditions within a particular year. The plugs should not be kept in the nursery for longer than three months and larger plugs (96 seedlings per tray) should be used. Nursery raised plugs of T. triandra and H. contortus were planted in an equal mix in an area that was cleared of A. mearnsii in 1996. By June 1998 661 H. contortus seedlings and 14 T. triandra seedlings had germinated naturally. The November 1998 population consisted of 418 H. contortus seedlings and 18 T. triandra seedlings. By May 2000 the June 1998 population showed a survival of 78.4% and the November 1998 population showed a survival of 91 .1 %. In the various trials, the ability of the nursery raised plugs used for re-vegetation to suppress the regrowth of A. mearnsii was investigated by determining the number of A. mearnsii seedlings per metre squared. The plant spacing and species of plugs used did not have a significant effect on the number of A. mearnsii seedlings per metre squared. Over-sowing with E. curvula did, however, significantly suppress the wattle re-growth. In the thatching trial the early harvested plots showed lower numbers of A. mearnsii per metre squared than the late harvest plots, as they were covered with a thick layer of thatch soon after the A. mearnsii was cleared which suppressed the A. mearnsii re-growth. Although E. curvula is able to produce a high biomass and suppress the A. meansii seedlings, it has a detrimental effect on the biodiversity of the area. Therefore, in conservation areas, where biodiversity is of great importance the planted plugs (at 30 cm spacing) or seed bearing hay must be used in preference to sowing E. curvula , although it must be remembered that greater follow up control is likely to be needed with planted plugs or seed bearing hay. The area must be planted or thatched as soon as possible after clear felling to provide competition for the A. mearnsii seedlings. / Thesis (M.Sc.Agric.)-University of KwaZulu-Natal, Pietermaritzburg, 2005.

The dynamics and complexities of climate-soil-vegetation relations in the Kruger National Park are poorly known. Although primary production and composition of the grass layer are very important components of the Park's ecosystem, equally little is known about the determinants of these parameters. A better understanding of these processes and relations will be of value to the management of this Park, as well as providing a better insight into these complex dynamics. A study was consequently undertaken covering a 14-year period to identify the most important determinants of above-ground grass production and composition. At the core of the study is the soil water balance. The use of evapotranspiration data in a study of this nature is however not absolutely essential, provided a variety of rainfall parameters are used, though it has the important advantage of providing a much more detailed and more complete insight into the relations of the grass sward with its environment. Stepwise and tree regression procedures were used to identify the important factors. It is concluded that rainfall in its various forms is the primary determinant of grass production, standing crop, and composition, the latter either as perennials or Decreasers. Secondary determinants, in varying degrees of importance, are the thickness and base status of the A horizon, distance to permanent drinking water, and competition by woody plants. Herbivore utilization is insignificant or at most, plays a relatively minor role. Herbivores appear to exert a negative influence on Decreaser abundance only when soil moisture stress exceeds a threshold level. When this is exceeded, relatively low herbivore densities are apparently sufficient to reduce Decreaser abundance. The definitions of Decreasers and Increasers consequently require revision to take into account the overriding influence of environmental factors, particularly those of soil moisture stress. The calibration of the disc pasture meter was re-evaluated. The relation between mean disc height and standing crop is non-linear. Up to a mean disc pasture meter height of 260 mm, the correlation between this parameter and above-ground standing crop is very strong (r2 = 0.95; P<0.0005). Beyond this height, the correlation is very poor (r2 =0.09; P<0.0005), apparently being strongly influenced by the structure of the grass plant, with tall grasses, or grasses with highly lignified culms resulting in a weaker correlation. / Thesis (M.Sc.)-University of Natal, Pietermaritzburg, 2003.