Note:

Please read the abstract in the section 00front of this document / Thesis (DPhil (Entomology))--University of Pretoria, 2005. / Zoology and Entomology / unrestricted

Scarabaeid pests in South Africa and especially KwaZulu-Natal are characterised by a very long larval life cycle and short pupal and adult periods. However, it has nearly always been the adults of the species that have been identified, with very little attention paid to the larval identification of the species. This is unfortunate as it is nearly always the larval stage that is found to be associated with crop damage. Accurate identification of the species of these larvae is important for the management of scarabaeid pest species, as it unlocks the necessary information on the biology and ecology of many species, which allows the adaptation of control methods for different species. Inadequate keys for the taxonomy of larvae of these groups, as well as the lack of morphological taxonomists working on these groups have been identified as constraints. When a species is difficult to identify using traditional taxonomic methods, DNA diagnostic tools can be useful. Chapter 2 investigated the feasibility of identifying scarabaeid larvae using mitochondrial DNA data. Variation in the base pair sequence of the mitochondrial cytochrome c oxidase sub unit I (cox 1) gene was used. DNA sequences of cox 1 from scarabaeid larvae collected from sugarcane fields were compared with sequences from scarabaeid adults of known species in order to identify the species attacking sugarcane. Neighbour-joining and maximum parsimony analyses of 658 bp cox 1 sequences identified groups of larvae that linked to adult specimens. The major groupings delimited specimens belonging to the subfamilies Dynastinae, Melolonthinae and Rutelinae. Within-group sequence divergence ranged from 0 - 3.4 % and divergence between sister groups ranged from 2.6 - 25.1 %. The recorded divergence range within and between tribes was 0 - 21.3 % and 17.3 - 28.5% respectively. Similarly, the divergence range observed within and between genera was 0 - 19.2 % and 17.1 - 25.4% respectively. The maximum sequence divergence observed within subfamilies was 23.7 % and divergence between subfamilies ranged from 16.8 - 26.7 %. Examination of pairwise sequence divergence levels as well as node support allowed 68% of the unidentified larval specimens to be associated with identified adult specimens. Phylogenetic analysis matched identified adult mtDNA with unidentified larval mtDNA. This allowed the identification of those larvae through morphological characteristics unique to certain species. To create a field key to the subfamilies of Dynastinae, Melolonthinae and Rutelinae the most useful character distinguishing larvae of different species was the raster but additional morphological characteristics were included. These relationships between larval and adult scarabaeid specimens from sugarcane were examined using various phylogenetic tools. The data set included a total of 19 morphological characters as well as 166 partial cox 1 gene sequences. Maximum parsimony analyses were performed on morphological, molecular and combined data. The same morphological and molecular data sets were run both separately and as a combined analysis with MrBayes. In both types of analyses the morphological data performed poorly and crude groupings resulted, dividing taxa to tribe level only. Molecular data showed greater resolution than the morphological data and taxa were separated into groups equivalent to species and morphospecies designated in Chapter 2. A partition homogeneity test indicated that both data types could be combined. It is recommended that both morphological and molecular data be utilised in identification of scarabaeid sugarcane pests and that a character-based approach be implemented. Further molecular data from other genes should be included to test the accuracy of these results. The keys produced during this study will allow workers to focus on a single species biology, and subsequently allow an analysis of between species interactions, and within species control. These advances are a start to the improvement of knowledge of the species composition of scarabaeid larvae in sugarcane fields, thus making management and biological control of these pests a greater possibility. Further recommendations for future work are discussed in Chapter 5. / Thesis (M.Sc.)-University of KwaZulu-Natal, Pietermaritzburg, 2008.

Submitted by Dominick Jesus (dominickdejesus@hotmail.com) on 2016-02-02T19:44:58Z No. of bitstreams: 2 Dissertação_Héctor Jaime Gasca Alvarez.pdf: 13762745 bytes, checksum: e958548660b8b2eef43f22abf72486f6 (MD5) license_rdf: 23148 bytes, checksum: 9da0b6dfac957114c6a7714714b86306 (MD5) / Made available in DSpace on 2016-02-02T19:44:58Z (GMT). No. of bitstreams: 2 Dissertação_Héctor Jaime Gasca Alvarez.pdf: 13762745 bytes, checksum: e958548660b8b2eef43f22abf72486f6 (MD5) license_rdf: 23148 bytes, checksum: 9da0b6dfac957114c6a7714714b86306 (MD5) Previous issue date: 2008-04-22 / Conselho Nacional de Pesquisa e Desenvolvimento Científico e Tecnológico - CNPq / The Oryctini (Coleoptera: Scarabaeidae: Dynastinae) it a large tribe of worldwide distribution with approximately 26 genera and 230 species. Fourteen genera and 113 species are distributed in the Neotropical region. This group of scarab beetles is characterized by a generally large and robust body, black or brown coloration, exposed mandibles, protíbia tri- or quadridentate with large teeth, and pronounced sexual dimorphism with the males possessing prominent tubercles or horns on the head and/or pronotum. Adults are nocturnally active and generally feed on rotting fruits or decaying vegetation. Knowledge of the tribe in the Neotropics is fragmentary, necessitating further studies that address taxonomy, biology and geographical distribution patterns. This study is a faunistic survey of the Oryctini of the Brazilian Amazon. The composition of the group in the study area is 7 genera and 19 species found in 7 states, 91 municipal districts, and approximately 167 specific localities. States with larger number of species are Amazonas and Pará, with 17 and 13 respectively. Heterogomphus eteocles Burmeister, 1847, Heterogomphus aidoneus (Perty, 1830), Heterogomphus telamon Burmeister, 1847, Megaceras crassum Prell, 1914, and Megaceras laevipenne Prell, 1914, are reported for the first time from the study area. Megaceras laevipenne is reported for the first time from Brazil. The taxomony, descriptions, distribution maps, and biological and ecological data are provided for all species. A characters analysis is provided as well as an identification key for all Oryctini species that occur in Brazilian Amazon. / A tribo Oryctini (Coleoptera: Scarabaeidae: Dynastinae) é um grupo de distribuição mundial formada por 26 gêneros e aproximadamente 230 espécies, dos quais 14 gêneros e 113 espécies estão distribuídas na região Neotropical. Este grupo de coleópteros pode ser reconhecido pela presença de um corpo geralmente grande e robusto, coloração escura, preta ou castanha, mandíbulas largas e expostas, protíbias com três a quatro dentículos grandes e um acentuado dimorfismo sexual, onde os machos possuem tubérculos ou cornos proeminentes no pronoto e/ou a cabeça. Os adultos têm hábitos noturnos e se alimentam usualmente de matéria vegetal em decomposição ou de frutos doces apodrecidos O conhecimento da tribo no Neotrópico ainda é escasso sendo necessários estudos que ajudem a esclarecer aspectos relacionados com a taxonomia, biologia e padrões de distribuição geográfica das espécies. Realizou-se um estudo faunístico para a tribo Oryctini da Amazônia brasileira. A diversidade no grupo para a região estudada está representada por 7 gêneros e 19 espécies, registradas em 7 estados, 91 municípios e aproximadamente 167 localidades. Os Estados com maior número de espécies foram o Amazonas e o Pará, com 17 e 13 respectivamente. São registradas pela primeira vez para a Amazônia brasileira as espécies Heterogomphus eteocles Burmeister, 1847, Heterogomphus aidoneus (Perty, 1830), Heterogomphus telamon Burmeister, 1847, Megaceras crassum Prell, 1914 e Megaceras laevipenne Prell, 1914, sendo esta última novo registro para o Brasil. A taxonomia, descrições morfológicas, dados biológicos e ecológicos, e mapas de distribução geográfica são apresentados para cada espécie. A partir de um análise de caracteres taxonômicos, foi elaborada uma chave dicotômica para a identificação das espécies de Oryctini que ocorrem na Amazônia brasileira.

No description available.

Thesis (M.A.)--The University of North Carolina at Greensboro, 2010. / Directed by Jan Rychtar; submitted to the Dept. of Mathematics and Statistics. Title from PDF t.p. (viewed Jul. 5, 2010). Includes bibliographical references (p. 54-57).

1. Twenty four species of the genus Sisyphus known in southern Africa are listed with their synonyms and their taxonomy is discussed. 2. A key to the adults of 23 of these species is given. 3. The known distribution of 23 species and seasonal occurrence of 19 species are illustrated and briefly discussed. 4. The general biology of the genus Sisyphus is discussed and the major behavioural and biological differences between the two subgenera Sisyphus and Neosisyphus are given. 5. The laboratory and field investigation into variations in behaviour and biology in eight species occurring in the Mkuzi Game Reserve, with regards to interspecific competition, is presented. 6. The known possible natural enemies of Sisyphus are mentioned and the extent of competition from other genera, based on field and laboratory observations, is outlined. 7. Summary tables of all data with a final tabulated evaluation of the suitability of species for introduction into Australia is given. 8. Nine species have been highly recommended for Australia. Six more species could be considered but most of these still need further investigation. The remaining nine species were found to be unsuitable for transfer to Australia.

No description available.

Despite belonging to the best described patterns in ecology, the mechanisms driving biodiversity along broad-scale climatic gradients, like the latitudinal gradient in diversity, remain poorly understood. Because of their high biodiversity, restricted spatial ranges, the continuous change in abiotic factors with altitude and their worldwide occurrence, mountains constitute ideal study systems to elucidate the predictors of global biodiversity patterns. However, mountain ecosystems are increasingly threatened by human land use and climate change. Since the consequences of such alterations on mountainous biodiversity and related ecosystem services are hardly known, research along elevational gradients is also of utmost importance from a conservation point of view. In addition to classical biodiversity research focusing on taxonomy, the significance of studying functional traits and their prominence in biodiversity ecosystem functioning (BEF) relationships is increasingly acknowledged. In this dissertation, I explore the patterns and drivers of mammal and dung beetle diversity along elevational and land use gradients on Mt. Kilimanjaro, Tanzania. Furthermore, I investigate the predictors of dung decomposition by dung beetles under different extinction scenarios. Mammals are not only charismatic, they also fulfil important roles in ecosystems. They provide important ecosystem services such as seed dispersal and nutrient cycling by turning over high amounts of biomass. In chapter II, I show that mammal diversity and community biomass both exhibited a unimodal distribution with elevation on Mt.Kilimanjaro and were mainly impacted by primary productivity, a measure of the total food abundance, and the protection status of study plots. Due to their large size and endothermy, mammals, in contrast to most arthopods, are theoretically predicted to be limited by food availability. My results are in concordance with this prediction. The significantly higher diversity and biomass in the Kilimanjaro National Park and in other conservation areas underscore the important role of habitat protection is vital for the conservation of large mammal biodiversity on tropical mountains. Dung beetles are dependent on mammals since they rely upon mammalian dung as a food and nesting resource. Dung beetles are also important ecosystem service providers: they play an important role in nutrient cycling, bioturbation, secondary seed dispersal and parasite suppression. In chapter III, I show that dung beetle diversity declined with elevation while dung beetle abundance followed a hump-shaped pattern along the elevational gradient. In contrast to mammals, dung beetle diversity was primarily predicted by temperature. Despite my attempt to accurately quantifiy mammalian dung resources by calculating mammalian defecation rates, I did not find an influence of dung resource availability on dung beetle richness. Instead, higher temperature translated into higher dung beetle diversity. Apart from being important ecosystem service providers, dung beetles are also model organisms for BEF studies since they rely on a resource which can be quantified easily. In chapter IV, I explore dung decomposition by dung beetles along the elevational gradient by means of an exclosure experiment in the presence of the whole dung beetle community, in the absence of large dung beetles and without any dung beetles. I show that dung decomposition was the highest when the dung could be decomposed by the whole dung beetle community, while dung decomposition was significantly reduced in the sole presence of small dung beetles and the lowest in the absence of dung beetles. Furthermore, I demonstrate that the drivers of dung decomposition were depend on the intactness of the dung beetle community. While body size was the most important driver in the presence of the whole dung beetle community, species richness gained in importance when large dung beetles were excluded. In the most perturbed state of the system with no dung beetles present, temperature was the sole driver of dung decomposition. In conclusion, abiotic drivers become more important predictors of ecosystem services the more the study system is disturbed. In this dissertation, I exemplify that the drivers of diversity along broad-scale climatic gradients on Mt. Kilimanjaro depend on the thermoregulatory strategy of organisms. While mammal diversity was mainly impacted by food/energy resources, dung beetle diversity was mainly limited by temperature. I also demonstrate the importance of protected areas for the preservation of large mammal biodiversity. Furthermore, I show that large dung beetles were disproportionately important for dung decomposition as dung decomposition significantly decreased when large dung beetles were excluded. As regards land use, I did not detect an overall effect on dung beetle and mammal diversity nor on dung beetle-mediated dung decomposition. However, for the most specialised mammal trophic guilds and dung beetle functional groups, negative land use effects were already visible. Even though the current moderate levels of land use on Mt. Kilimanjaro can sustain high levels of biodiversity, the pressure of the human population on Mt. Kilimanjaro is increasing and further land use intensification poses a great threat to biodiversity. In synergy wih land use, climate change is jeopardizing current patterns and levels of biodiversity with the potential to displace communities, which may have unpredictable consequences for ecosystem service provisioning in the future. / Gradienten der Biodiversität, wie der Breitengradient der Artenvielfalt, gehören zu den bestbeschriebenen Mustern in der Ökologie. Dennoch bleiben die Mechanismen, die diese Gradienten steuern, unzureichend untersucht. Bergmassive eignen sich aufgrund ihrer hohen Artenvielfalt, ihrer räumlichen Begrenzung, der gleichmäßigen Veränderung abiotischer Faktoren mit der Höhe und ihres weltweiten Auftretens optimal zur Erforschung der Triebkräfte globaler Biodiversitätsmuster. Jedoch werden Gebirgs-Ökosysteme vermehrt durch menschliche Landnutzung und den Klimawandel bedroht. Da der Wissenstand über die Auswirkungen solcher Veränderungen auf die Biodiversität von Bergmassiven und zugehörigen Ökosystemdienstleistungen gering ist, nimmt die Erforschung von Höhengradienten auch aus der Perspektive des Artenschutzes eine besondere Bedeutung ein. In Ergänzung zur traditionellen, auf Taxonomie beruhenden Biodiversitätsforschung, wird die Wichtigkeit der Untersuchung funktioneller Merkmale und deren Bedeutung für Beziehungen zwischen Biodiversität und Ökosystemfunktionen (BEF) zunehmend anerkannt. In meiner Doktorarbeit untersuche ich entlang von Höhen- und Landnutzungsgradienten am Kilmandscharo (Tansania) die Muster und Triebkräfte der Artenvielfalt von Säugetieren und Dungkäfern als auch die Faktoren, die den Dungabbau durch Dungkäfer unter verschiedenen Aussterbe-Szenarien bestimmen. Säugetiere sind nicht nur charismatisch, sie nehmen auch wichtige Rollen in Ökosystemen ein. So erfüllen Säugetiere wichtige Ökosystemdienstleistungen wie die Verbreitung von Samen und sind maßgeblich am Nährstoffkreislauf durch den Umsatz großer Mengen von Biomasse beteiligt. Im zweiten Kapitel dieser Arbeit stelle ich dar, dass die Diversität und Biomasse der Säugetiergemeinschaft am Kilimandscharo eine unimodale Verteilung mit der Höhe aufweist. Dieses Muster wurde vor allem durch die Nettoprimärproduktion, ein Maß für die Nahrungsverfügbarkeit der Säugetiere, und den Schutzstatus der Untersuchungsgebiete bestimmt. Aufgrund ihrer Größe und Endothermie kann man schlussfolgern, dass für Säugetiere, im Unterschied zu den meisten Arthropoden, Nahrungsverfügbarkeit die Triebkraft der Diversität darstellt. Meine Resultate bestätigen diese Vorhersage. Die signifikant höhere Diversität und Biomasse der Säugetiere im Kilmandscharo Nationalpark und in anderen geschützten Gebieten unterstreicht die Wichtigkeit des Habitatschutzes für den Erhalt der Artenvielfalt großer Säugetiere in tropischen Bergmassiven. Dungkäfer stehen in enger Beziehung zu Säugetieren, da sie Säugetierdung als Nahrungs- und Nistmaterial benötigen. Dungkäfer übernehmen ebenfalls wichtige Ökosystemdienstleistungen: Sie spielen eine bedeutende Rolle im Nährstoffkreislauf und tragen entscheidend zur Bioturbation, der sekundären Verbreitung von Samen und der Unterdrückung von Schädlingen bei. Im dritten Kapitel weise ich nach, dass die Artenvielfalt der Dungkäfer mit der Höhe abnimmt, während die Abundanz der Käfer eine eingipfelige Verteilung zeigt. Im Unterschied zu den Säugetieren wurde die Diversität der Dungkäfer vor allem durch die Temperatur gesteuert. Obwohl ich versuchte, die vorhandenen Dungressourcen der Säugetiere möglichst genau durch die Berechung des Kotabsatzes zu quantifizieren, stellte ich keinen Einfluss von Ressourcenverfügbarkeit auf die Dungkäfer-Diversität fest. Stattdessen führte eine höhere Temperatur zu erhöhter Dungkäfer-Diversität. Abgesehen von ihrer Rolle als wichtige Ökosystemdienstleister stellen Dungkäfer auch Modellorganismen für BEF-Studien dar, da sie eine leicht zu quantifizierende Ressource benötigen. Im vierten Kapitel untersuche ich den Dungabbau von Dungkäfern entlang des Höhengradienten mithilfe eines Ausschlussexperiments: in der Gegenwart der gesamten Dungkäfergemeinschaft, unter dem Ausschluss großer Dungkäfer und in der Abwesenheit aller Dungkäfer. Der Dungabbau war am größten, wenn der Abbau durch die gesamte Dungkäfergemeinschaft erfolgen konnte. Waren nur kleine Dungkäfer anwesend, waren die Dungabbauraten deutlich geringer als in der Gegenwart großer Dungkäfer, während sie im Falle des Ausschlusses aller Dungkäfer minimal wurden. Außerdem konnte ich nachweisen, dass die Triebkräfte des Dungabbaus von dem Zustand der Dungkäfergemeinschaft abhingen. Während die mittlere Körpergröße von Dungkäfern der wichtigste Faktor darstellte, wenn die Lebensgemeinschaft vollständig war, erlangte die Artenvielfalt an Bedeutung, wenn große Dungkäfer abwesend waren. Im gestörtesten Zustand des Systems, wo der Dungabbau ohne Dungkäfer erfolgte, war Temperatur der einzige Faktor, der den Dungabbau bestimmte. Abiotische Faktoren nehmen an Wichtigkeit als Triebkräfte von Ökosystemdienstleistungen zu, je mehr das System gestört ist. Zusammenfassend wird in dieser Dissertation gezeigt, dass die Triebkräfte der Artenvielfalt entlang weitreichender klimatischer Gradienten am Kilimandscharo von der thermoregulatorischen Strategie der Organismen abhängen. Während die Diversität von Säugetieren vor allem durch die Nahrungsverfügbarkeit beeinflusst wurde, wurde die Dungkäfer-Diversität vor allem durch die Temperatur gesteuert. Außerdem sind geschützte Flächen für den Erhalt der Artenvielfalt großer Säugetiere unerlässlich. Weiterhin veranschauliche ich die herausragende Bedeutung großer Dungkäfer für den Dungabbau, da letzterer deutlich abnahm, wenn große Dungkäfer ausgeschlossen wurden. Betreffend der Landnutzung war insgesamt kein Einfluss auf die Dungkäfer- oder Säugetier-Diversität oder den Dungabbau durch Dungkäfer feststellbar. Anders sah es auf Ebene der am meisten spezialisierten trophischen Gilden der Säugetiere und funktionellen Gruppen der Dungkäfer aus: Hier waren bereits negative Auswirkungen sichtbar. Obwohl unter dem derzeitigen gemäßigten Ausmaß der Landnutzung am Kilimandscharo eine hohe Artenvielfalt aufrechterhalten werden kann, steigt der Druck durch das Bevölkerungswachstum, und eine zunehmende Intensivierung der Landwirtschaft stellt eine große Bedrohung für die Biodiversität dar. Im Zusammenspiel mit der Landnutzung gefährdet der Klimawandel das Niveau und die Verteilung der Biodiversität, mit dem Potential, Gemeinschaften von Organismen zu verdrängen, was unvorhersagbare Auswirkungen auf die Bereitstellung von Ökosystemdienstleistungen in der Zukunft haben könnte.

In nature, both fundamental and sophisticated ecological functionality alike are driven by the display or, conversely, the concealment of colour and visual contrast. Through scientific investigation, the evolutionary genius manifested in numerous light manipulating biological systems has provided many blueprints for innovative photonic devices and other smart materials. The development of innovative bioinspired materials that harness the optical properties exhibited by chiral structures, such as those found in certain Scarabaeidae, offer widespread potential. In this thesis, the species-specific circularly polarised optical behaviour corresponding to a variety of Scarabaeidae beetles is investigated. The helicoidal morphology giving rise to circularly polarised photonic properties is first described for several beetles displaying a narrowband colour appearance. A structural analysis of the native surface structures encountered in these species is presented alongside a discussion of their role in defining the species' optical characteristics. Experimental measurements of the circularly polarised scattering behaviour are presented and discussed with respect to the electron microscopy and surface morphology measurements comprising the structural analysis. Following this, the origins of the subtle differences in the metallic colour appearance of several broadband coleopteran structures are deciphered. The broadband response of these species is attributed to chirped configurations of the helicoidal lamellar structure, elucidated using electron microscopy. Results of this structural analysis are applied to theoretical simulations of the structures' electromagnetic response and consolidate experimental measurements of the circularly polarised reflectance. Rigorous experimental measurement demonstrated that the circularly polarised spectral response is relatively robust to small structural defects or `biological noise'. Building from this, the intraspecific colour variation displayed in the species Chrysina aurigans is a useful lens through which the nuances of the chirped multilayer structure are further examined. As a result, this work provides detailed insights into the fine-tuning of chirped biological photonic systems. Likewise, the topic of biological wave retarders is afforded significant attention, primarily with reference to the biological half-wave plate encountered in the cuticle structure of Chrysina resplendens. A rigorous analysis of the photonic system in C. resplendens is presented, leading to novel conclusions describing this system's opto-structural relationship. Additionally, this thesis reports, for the first time, the presence of a biological wave retarder in the cuticle structure of Chrysina cupreomarginata.