National Forest management in the Pacific Northwest is shifting from a focus on commodity production to ecosystem management, in which the health of the entire forest ecosystem is considered, rather than that of a few key species. Ecosystem management includes retention of some live trees following timber harvest (green-tree retention) to preserve biodiversity, imitating the natural fire regime of large, but patchy fires that leave many live trees. How ecosystem management will affect growth and species composition of future forests is an important question. This study takes a retrospective approach to this question by using past disturbance as an analogue to green-tree retention following timber harvest. Using USDA Forest Service timber inventory plot data from the Cascade Mountains of Oregon and SW Washington, 132 unmanaged stands were identified with a tree cohort of 70-110 years old (regeneration) or a tree cohort of 70-110 years old with an overstory of large trees 200+ years old (remnants). All stands were in the Tsuga heterophylla (Raf.) Sarg. zone (Franklin and Dyrness 1973). Single-aged stands represented clearcuts, while two-aged stands served as analogues to stands harvested with green-tree retention. Regeneration basal area/hectare (ba/ha) declined when remnant-tree densities exceeded about 15 remnant trees/ha (R��=0.51) in a relationship roughly described by a sigmoidal curve. Conceptually removing remnant-tree space occupancy effects decreased remnant-tree density's value as a predictor of regeneration ba/ha by about 50% at management-level remnant densities (���45 remnant trees/ha). Thus, it appears that remnant "effects" were a result of both remnant-tree space occupancy and remnant resource use. Douglas-fir ba/ha in the regeneration also declined when remnant-tree densities exceeded about 15 remnant trees/ha (R��=0.60). Western hemlock ba/ha in the regeneration increased slightly with increasing remnant-tree densities (R��=0.19). Western redcedar ba/ha in the regeneration was apparently not related to remnant-tree density (R��=0.02). The degree of aggregation in remnant trees did not appear to affect regeneration ba/ha, but few stands contained the isolated clumps of remnant trees likely under a management scenario. Neither measured site characteristics nor regeneration density was related to regeneration ba/ha across species. Remnant-tree density was apparently unrelated to tree-species diversity in the regeneration. Total-stand ba/ha remained relatively constant across remnant densities. / Graduation date: 1994

Montane meadows comprise less than 5% of the landscape of the western Cascades of Oregon, but they provide habitat for diverse species of plants and pollinators. Little is known about plant-pollinator network structure at these sites. This study quantified plant-pollinator interactions over the summer of 2011, based on six observations of 10 permanent subplots in 15 meadows, stratified by size and isolation. The study examined (1) relationships between richness and abundance of flowers, pollinators, and interactions; (2) distribution of abundance and richness of flowers, pollinators, and interactions with regards to surrounding meadow habitat; (3) change in flower and pollinator abundance over the season; (4) factors associated with the presence of various guilds of pollinators; and (5) the structure of plant-pollinator networks. The study showed that (1) richness of pollinators increased 2x faster than richness of flowers with increased abundance; (2) density of flowers and interactions was positively correlated with meadow size and diversity of pollinators and interactions were both correlated with surrounding habitat at two spatial scales; (3) peak flower abundance coincided with or preceded peaks in pollinator populations; (4) abundance of three guilds of bees exhibited different patterns of association to surrounding habitat and meadow soil moisture corresponding to various dispersal potential and phenology of guild species; and (5) the number of network pairings for plants and pollinators increased with increasing species richness of potential interaction partners and all networks were found to be significantly nested. Results of this study indicate that plant-pollinator networks are complex assemblages of species, in which spatial and temporal patterns of habitat affect species composition and network structure. In particular, flower and pollinator abundance and richness are depressed in small and isolated meadows. Significant nestedness emerged as a pattern of network level organization across the study meadows. / Graduation date: 2013