Mountain headwater catchments in the semi-arid Intermountain West are important sources of surface water because these high elevations receive more precipitation than neighboring lowlands. The hydrology of these mountain catchments is especially important as the region faces water shortages and conflicts. Conifer encroachment on aspen stands has been observed across the western US and can result in a decline in water yield. The overall objective of this study was to further our understanding of hillslope-stream connectivity in a headwater catchment of Northern Utah and any observable differences in this connection between aspen and conifer hillslopes. Hillslopes are the fundamental unit of a watershed. Therefore understanding processes at the hillslope scale is pertinent to managing valuable water resources. However, hillslope hydrology is understudied in the snow-driven, semi-arid west, leaving a gap in our knowledge of how watersheds function. This thesis focuses on how and when hillslope water contributes to stream water: hillslope-stream connectivity. Its specific objectives are (1) to compare peak snow accumulation under aspen and conifer stands, (2) to determine if shallow soil moisture shows organized patterns, indicating hillslope-connectivity and compare these patterns between vegetation types, (3) to examine hillslope-stream connectivity within deep layers of the soil profile and compare times of connectivity between vegetation types and (4) to find any thresholds past which hillslope-stream connectivity begins.

Improperly designed road-stream crossings can fragment stream networks by restricting or preventing aquatic organism passage. These crossings may also be more vulnerable to high flow events, putting critical human infrastructure at risk. Climate change, which will require access to suitable habitat for species persistence, and is also predicted to increase the frequency and magnitude of extreme floods, underscores the importance of maintaining stream connectivity and resilient infrastructure. Given the large number of road-stream crossings and the expense of replacement, it is increasingly important to prioritize removals and account for the multiple benefits of these management actions. I developed an aquatic barrier prioritization scheme that combines potential habitat gain, stream thermal resilience, aquatic organism passage, and culvert risk of failure. To assess relative thermal resilience, I deployed paired air-water thermographs in several New England watersheds and analyzed relative thermal sensitivity (relationship of water to air temperature) and exposure (duration, frequency, and magnitude of warm stream temperature episodes) among streams. These were combined into a single metric of thermal resilience corresponding with the distance of that stream’s sensitivity and exposure from the watershed median. To test the relationship between risk of failure, culvert dimensions, and stream connectivity, I developed a logistic regression to predict risk of failure using data from two watersheds that experienced extreme flooding from Hurricane Irene (2011). Finally, I applied the resultant prioritization scheme to 66 road-stream crossings in the Westfield River watershed (MA). Thermal habitat quality varied considerably within and among watersheds. Stream sensitivity was generally lower than the widely accepted 0.8 ̊C increase in stream temperature for every 1 ̊C increase in air temperature (Westfield median sensitivity = 0.44), with substantial differences among streams. Exposure also varied widely among streams, indicating that some headwater streams in New England are more thermally resilient than previously thought. Risk of infrastructure failure was predicted with a logistic regression using culvert constriction ratio and predicted aquatic organism passage as predictors (Likelihood ratio test, X2=59.1, df=3, p- value=9.2e-13), indicating that underdesigned culverts were more likely to be barriers to passage and more likely to fail in extreme flow events. To prioritize culverts, this study ultimately used a piecewise approach that identified culverts opening the longest reaches of thermally resilient habitat, and then ranked those culverts by infrastructure replacement need. In the Westfield River, the prioritization clearly identified crossing replacements most likely to yield multiple benefits. The scheme I developed can accommodate changes in the relative weights of the different criteria, which will reflect differences in management and conservation concerns in the confidence of inputs. In conclusion, increasing connectivity by removing barriers may be one of the most effective ways to mitigate the effects of climate change on aquatic systems, but it is important to remove the right barriers.