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Filetički odnosi unutar sekcije SynochetaLegrand, 1946 (Crustacea, Isopoda, Oniscidea)Balkanskog poluostrva / Phyletic relations within Synocheta Legrand, 1946 section (Crustacea, Isopoda, Oniscidea) of the Balkan PeninsulaHorvatović Mladen 23 September 2014 (has links)
<p>Aktuelna sistematika sekcije Synocheta na gotovo svim nivoima ne odražava realne filetičke veze. Neobična distribucija, nejasni diferencijalni karakteri familije Styloniscidae i njene veze sa familijom Trichoniscidae su problematični. Kriterijumi podele Trichoniscidae na podfamilije su nejasni, nedosledni i često neprimenljivi. Otuda ne čudi krajnje uprošćena podela koja ne oslikava realne filetičke odnose unutar grupe, koji su znatno kompleksniji. </p><p>Od 593 vrste Synocheta, trećina (većinom endemita), naseljava Balkan, što nameće <br />ovo područje kao jedan od centara diverzifikacije i diverziteta grupe, i čini ga idealnim za sagledavanje realnih filetičkih odnosa. Ovo je definisalo glavne ciljeve naše studije: što potpunije sagledavanje faune balkanskih Synocheta kroz taksonomsku obradu; utvrđivanje filetičkih veza na osnovu kompleksa relevantnih karaktera uporednim analizama, sa težištem na konzervativnijim karakterima, što do sada nije učinjeno. </p><p>Konstatovali smo u balkanskoj fauni iz familije Styloniscidae 4 roda sa 15 vrsta, od toga 3 roda i 4 vrste su nove za nauku; iz familije Trichoniscidae 33 roda sa 176 vrsta, od toga 1 novi rod i 18 novih vrsta. Od tog broja 27 rodova i 161 vrsta Synocheta je endemično za Balkan. </p><p>Kod Styloniscidae smo utvrdili znatno veći diverzitet i heterogenost od onoga što je bilo do sada poznato, što ukazuje na moguće poreklo ove grupe sa prostora severne Gondvane (delom inkorporirane u prostore Balkana), zajedničko sa familijom Trichoniscidae.</p><p>Rekonstruisali smo najznačajnije momente u filogeniji Trichoniscidae: vrlo rano razviće konglobacije, uz masivan integument i razvijenu ornamentiku kao adaptacije; prelazak na zaštitu stereotaksacijom; evolucija ka aktivnijoj zaštiti i gubitku „oklopa” što dovodi do velike adaptivne radijacije. Pri tome smo pokazali da su karakteri koji su smatrani izvedenim osobinama zapravo pleziomorfni.</p><p>Predstavili smo osnovne evolutivne tokove u familiji Trichoniscidae, kao i filetičke <br />veze koje proizilaze iz ove studije: Haplophthalminae su stara grupa koja poseduje niz pleziomorfnih karaktera; Buddelndiellinae su stara grupa sa znatno bližim filetičkim vezama sa Haplophthalminae; Thaumatoniscellinae imaju vrlo davno zajedničko poreklo sa Haplophthalminae; Trichoniscinae su mlađa, parafiletička grupa, a mnoge linije su nezavisno evoluirale od podfamilije Haplophthalminae. </p><p>Ovom studijom smo u nekim segmentima dokazali dijametralno suprotne, znatno kompleksnije filetičke veze unutar Synocheta u odnosu na do sada prezentovane.</p> / <p>Current systematic Synocheta section at almost all levels fails to reflect the real phyletic relations. Unusual distribution and vague differential characters of Styloniscidae family and its relationships with the Trichoniscidae family are problematic. The division criteria for forming Trichoniscidae subfamilies are unclear, inconsistent and often unenforceable. Hence, the extremely simplified division that does not reflect the real phyletic relations within the group, which are much more complex, is not surprising.</p><p>Of 593 Synocheta species, one third (mostly endemic) inhabit the Balkans, which imposes this area as one of the centers of diversification and diversity of the group, and makes it ideal for the analysis of real phyletic relations. This defined the main goals of our study: as complete assessment of the fauna of the Balkan Synocheta as achievable through taxonomic treatment; determining phyletic relations through comparative analyses based on the complex of relevant characters, with the emphasis on the more conservative ones, which has not been accomplished thus far.</p><p>In the Balkan fauna of the Styloniscidae family, 4 genera with 15 species were identified, of which 3 genera and 4 species are new to science; similarly, the 33 genera with 176 species from the family Trichoniscidae found in Balkan fauna included one new genus and 18 new species. Of that number, 27 genera and 161 Synocheta species are endemic to the Balkans.</p><p>In Styloniscidae, we found much greater diversity and heterogeneity than was previously established, which indicates that these groups possibly originate from the northern Gondwana (partly incorporated into the Balkans area), in common with Trichoniscidae family.</p><p>We reconstructed the most important moments in the Trichoniscidae philogeny: early development of conglobation, with massive integument and developed rnamentation as adaptations; transition into protection via stereotaxation; evolution toward more active protection and loss of "armor", which leads to extensive adaptive radiation. In addition, we have shown that the characters that were previously considered derived properties are actually plesiomorphic.</p><p>We have presented the basic evolutionary trends in Trichoniscidae family, as well as phyletic relations arising from this study: Haplophthalminae are the ancient group ossessing a wide range of plesiomorphic characters; Buddelndiellinae are the ancient group with a much VI closer phyletic relations to Haplophthalminae; Thaumatoniscellinae share a common, and very distant, origin with Haplophthalminae; Trichoniscinae are a younger paraphyletic group and many lines have evolved independently of the Haplophthalminae subfamily.</p><p>In this study, in some segments, we have demonstrated substantially more complex phyletic relations within Synocheta, diametrically opposed to the previously presented findings.</p>
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Taksonomija i zoogeografija paukova (Arachnida,Araneae) Republike Makedonije / Taxonomy and zoogeography of spiders (Arachnida, Araneae) of Republic of MacedoniaMarjan Komnenov 30 September 2019 (has links)
<p>Doktorska disertacija predstavlja studiju faune paukova (Arachnida, Araneae) Makedonije baziranu na kritičkoj analizi publikovanih podataka koji obuhvataju period od 1907. do 2018. godine, taksonomskoj reviziji relevantnih arahnoloških zbirki iz osam evropskih prirodnjačkih muzeja i rezultatima sopstvenih tridesetogodišnjih istraživanja. Terenski rad je baziran na sakupljanju materijala<br />korišćenjem standardnih metoda kao što su ručno sakupljanje i sakupljanje putem klopki i noviju metodu upotrebom motornog usisivača. Materijal je<br />laboratorijski obrađivan standardnim postupcima uz pomoć binokularne lupe i mikroskopa. Ukupno su registrovane 742 vrste paukova iz 40 familija. Pet taksona pretstavljaju nove vrste za nauku. Dve vrste predstavljaju nove nalaze za faunu paukova Balkanskog poluostrva. Šezdeset i dva taksona su po prvi put zabeležene za faunu Makedonije. Drevna familija Anapidae je sa jednom vrstom po prvi put registrovana u fauni Makedonije. Revizijom publikovanih podataka, ustanovljeno je da je179 vrsta pogrešno identifikovano i navedeno za<br />faunu Makedonije. Vrste su kategorisane prema trenutnom poznavanju njihove rasprostranjenosti, na osnovu čega je izvršena zoogeografska analiza faune. Ustanovljena su 32 horotipa u fauni paukova Makedonije, svrstana u četiri horološka kompleksa: šire rasprostranjene, evropske, mediteranske i endemske vrste. Najveći broj vrsta registrovano je u kompleksu široko rasprostranjenih vrsta (44,3%) koji daju glavni karakter fauni. Visok stepen zastupljenosti evropskih (19,1%) i eurosibirskih (11,3%) horoloških elemenata, kao dva pojedinačno najzastupljenija u fauni paukova Makedonije, je odraz dominantnog planinskog karaktera reljefa Makedonije. Horološki kompleks mediteranskih vrsta zastupljen je sa 99 vrsta (13,3%). Visok udeo vrsta (11,3%) koje pripadaju horološkim elementima koji su zastupljeni na prostorima Južne Evrope, Mediterana, Zapadne i<br />Srednje Azije govori o bogatoj fauni paukova kserofilnih, termofilnih i aridnih terena. Lokalni karakter faune može se sagledati u relativno velikom broja vrsta iz kompleksa endemskih vrsta (87). Iz ovog kompleksa najdominantniji su vrste iz horotipa "endemiti užeg područja" sa 45 vrsta.Doktorska disertacija predstavlja studiju faune paukova (Arachnida, Araneae) Makedonije baziranu na kritičkoj analizi publikovanih podataka koji obuhvataju period od 1907. do 2018. godine,<br />taksonomskoj reviziji relevantnih arahnoloških zbirki iz osam evropskih prirodnjačkih muzeja i rezultatima sopstvenih tridesetogodišnjih istraživanja.Terenski rad je baziran na sakupljanju materijala korišćenjem standardnih metoda kao što su ručno sakupljanje i sakupljanje putem klopki i noviju metodu upotrebom motornog usisivača. Materijal je laboratorijski obrađivan standardnim postupcima uz pomoć binokularne lupe i mikroskopa. Ukupno su registrovane 742 vrste paukova iz 40<br />familija. Pet taksona pretstavljaju nove vrste za nauku. Dve vrste predstavljaju nove nalaze za faunu paukova Balkanskog poluostrva. Šezdeset i dva taksona su po prvi put zabeležene za faunu Makedonije. Drevna familija Anapidae je sa jednom vrstom po prvi put registrovana u fauni Makedonije. Revizijom publikovanih podataka, ustanovljeno je da je179 vrsta pogrešno identifikovano i navedeno za<br />faunu Makedonije. Vrste su kategorisane prema trenutnom poznavanju njihove rasprostranjenosti, na osnovu čega je izvršena zoogeografska analiza faune. Ustanovljena su 32 horotipa u fauni paukova Makedonije, svrstana u četiri horološka kompleksa: šire rasprostranjene, evropske, mediteranske i endemske vrste. Najveći broj vrsta registrovano je u kompleksu široko rasprostranjenih vrsta (44,3%) koji daju glavni karakter fauni. Visok stepen zastupljenosti evropskih (19,1%) i eurosibirskih (11,3%) horoloških elemenata, kao dva pojedinačno najzastupljenija u fauni paukova Makedonije, je odraz dominantnog planinskog karaktera reljefa Makedonije. Horološki kompleks mediteranskih vrsta zastupljen je sa 99 vrsta (13,3%). Visok udeo vrsta (11,3%) koje pripadaju horološkim elementima koji su zastupljeni na prostorima Južne Evrope, Mediterana, Zapadne i<br />Srednje Azije govori o bogatoj fauni paukova kserofilnih, termofilnih i aridnih terena. Lokalni karakter faune može se sagledati u relativno velikom broja vrsta iz kompleksa endemskih vrsta (87). Iz ovog kompleksa najdominantniji su vrste iz horotipa "endemiti užeg područja" sa 45 vrsta.</p> / <p>The PhD research project is a study of fauna of spiders (Arachnida, Araneae) of Macedonia based on a critical analysis of published data covering the period from 1907 to 2018, taxonomic review of relevant arachnological collections from eight European Natural history museums and the results of author’s thirty years of research. Field work is based on the collection of materials using standard methods such as manual collection and trapping and a newer method using a motor vacuum cleaner. The material was processed with standard procedures using a dissecting binocular and a microscope.A total of 742 species of spiders from 40 families have been registered. Five taxa are new species for science. Two species present new findings for the fauna of the spiders of the Balkan Peninsula. Sixty two taxa were recorded for the first time for the fauna of Macedonia.An ancient family Anapidae with one species has been registered for the first time in the fauna of Macedonia. By reviewing the published data, it was established that 179 species were incorrectly identified and listed for the fauna of Macedonia. Species are categorized according to the current knowledge of their distribution, on the basis of which a zoogeographic analysis of the fauna was performed.32 chorotypes were established in the fauna of the spiders of Macedonia, classified into four chorological complexes: widespread, European, Mediterranean and endemic species. The largest number of species is registered in the complex of widespread species (44.3%), which give the main character of the fauna. The high degree of representation of European (19.1%) and Eurosiberian (11.3%) chorological elements, as the two individually most represented in the fauna of the spiders of Macedonia, is a reflection of the dominant mountain character of the relief of Macedonia. The chorological complex of Mediterranean species is represented by 99 species (13.3%). The high proportion of species (11.3%) belonging to the chorological elements that are present on the territory of Southern Europe, the Mediterranean, West and Central Asia speaks of the rich fauna of spiders of xerophilic, thermophilic and arid terrain. The local character of the fauna can be seen in a relatively large number of species from the complex of endemic species (87). From this complex are the most dominant species from the chorotype "narrower endemic" with 45 species.Western Macedonia is the richest with fauna of spiders, with 23 species from the group of endemites of the narrower area, which speaks of the great importance of this region as a refugium and space in which the processes of speciation were intensively carried out.</p>
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Strateginės analizės instrumentarijaus modeliavimas mažose ir vidutinėse įmonėse / Modelling of Strategic Analysis Instrumentarium at Small and Medium - sized EnterprisesVaitkevičius, Sigitas 19 April 2006 (has links)
The doctoral thesis is concerned with the issues that are faced when selecting strategic analysis tools to be used in small and medium-sized enterprises (SME) in the process of strategic management. The paper comprises three parts.
The first part presents a discussion on the methodology of strategic analysis and the application of its tools in SME. As a result, the taxonomy of 40 strategic analysis tools applied in business organizations (including SME) was designed. In addition, the „model of strategic tune“, defining the correlation of „a dominating strategy type“, „the size of an enterprise“, and „the nature of strategic analysis“, was formed in this part.
The second part describes the methodologies to study the application of strategic analysis tools. In the course of research project, two individual empirical studies were conducted: 1) expert evaluation study of strategic analysis tools, during which the criteria for expert evaluation of 40 strategic analysis tools were prepared; 2) SME owners, executives and specialists‘ testing and survey.
The third part of the thesis indicates the results of the study into the application of strategic analysis tools. While combining theoretical and empirical methods and using various classification criteria, four typologies of strategic analysis tools were designed. Factual material reflecting SME executives‘ scope of knowledge and their perceptions and attitudes towards strategic analysis was collected by means of testing and... [to full text]
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Taxonomic and phylogenetic review of the genus Trichocera Meigen, 1803 (Diptera, Trichoceridae) / Taksonominė ir filogenetinė genties Trichocera Meigen, 1803 (Diptera, Trichoceridae) analizėPetrašiūnas, Andrius 02 December 2009 (has links)
Winter gnats of the genus Trichocera are one of the main insect groups found in mild climates during the autumn, winter and spring months and so are unique in their way of adapting to living in colder seasons. Phylogenetic links of the genus Trichocera are examined, taxonomy of the genus renewed, species composition of several regions is ascertained and characters for dividing into subgenuses are specified in this thesis. Based on the material collected in Lithuania by standard entomological techniques and by analysis of literature data, a list of 22 species of winter gnats is given with 9 species new for our country. One new for science species of winter gnats is described from Sardinia, Italy. Based on the selected morphologic characters, phylogenetic analysis was performed and 15 species were transferred from one subgenus to another as a result. Characters of male aedeagal complex were found to be a basis for attributing the species into particular subgenus. For the first time important structures of the females of four species, male of one species and type specimens of 15 more species were illustrated. World catalogue of the 110 recent species of Trichocera is compiled and different structures of 106 species are illustrated. / Genties Trichocera uodai yra viena pagrindinių vabzdžių grupių, aptinkamų vidutinėse platumose rudens, žiemos ir pavasario mėnesiais, ypatinga savo prisitaikymu gyventi šaltuoju laikotarpiu. Disertacijoje nagrinėjami genties Trichocera rūšių filogenetiniai ryšiai, atnaujinta genties taksonomija, išaiškinta atskirų regionų rūšinė įvairovė, patikslinti skirstymui į pogentes naudotini požymiai. Standartiniais entomologiniais metodais Lietuvoje surinktų žieminių uodų mėginių pagrindu bei pagal literatūros šaltinius buvo sudarytas 22 rūšių katalogas, papildant jį 9 naujomis mūsų šalies faunai rūšimis. Aprašyta nauja mokslui žieminių uodų rūšis iš Sardinijos salos (Italija). Remiantis atrinktais požymiais atlikta filogenetinė analizė, kurios pagrindu penkiolika rūšių perkeltos iš vienų pogenčių į kitas. Nustatyta, kad patinų edeaguso komplekso ypatybės yra svarbiausios skirstant rūšis į pogentes. Pirmą kartą iliustruotos keturių rūšių patelių ir vienos rūšies patino svarbios identifikacijai struktūros bei dar penkiolikos rūšių tipiniai egzemplioriai. Sudarytas 110 recentinių Trichocera genties rūšių pasaulio faunos katalogas, pateiktos 106 rūšių įvairių identifikacinių struktūrų iliustracijos.
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Fauna osolikih muva (Diptera: Syrphidae) Grčke / Fauna of hoverflies (Diptera: Syrphidae) of GreeceLikov Laura 28 August 2018 (has links)
<p>Osolike muve predstavljaju veoma bitnu i korisnu insekatsku grupu. Pružaju nezamenljive ekološke i ekonomske usluge, kao i usluge za životnu sredinu. Navedene usluge su omogućene njihovom sposobnošću oprašivanja biljaka u prirodnim i agroekosistemima, kao i zahvaljujući larvama– predatorima lisnih vaši i drugih insekata, smanjujući ekonomske gubitke u prinosu kultivisanih biljaka. Takođe mogu poslužiti kao bioindikatori klimatskih promena i stanja staništa. Neke vrste imaju i konzervacioni značaj, obzirom da su zbog antropogenih uticaja postale ugrožene, što zahteva preduzimanje mera za obezbeđivanje njihovog opstanka. Izuzetno bogatstvo flore i faune na Balkanskom<br />poluostrvu predstavlja jedinstveni ekološki i biogeografski fenomen u Evropi. Grčka, kao jedna od najvažnijih centara biodiverzita na Balkanu i u Mediteranu se odlikuje širokim spektrom važnih tipova ekosistema, od kojih su najbitniji šumski, budući da najveći broj osolikih muva preferira upravo taj tip staništa.<br />Značaj faunističkih istraživanja se ogleda u otkrivanju sastava i brojnosti vrsta na određenim staništima ili područjima. Faunistički sastav pruža uvid u biodiverzitet istraživane oblasti, koja se neprekidno menja, naročito u poslednje vreme pod uticajem negativnog antropogenog uticaja. Podaci o vrstama mogu biti od velikog značaja za eventualne programe zaštite i oporavka ugroženih vrsta, kao i u planiranju formiranja novih zaštićenih područja ili u uvođenju novih mera zaštite u već postojećim prirodnim dobrima.<br />Prepoznavanje ugroženih vrsta ili ugroženih staništa koje one naseljavaju, igra bitnu ulogu u planiranju konzervacionih strategija za njihovo očuvanje. Rezultati doktorske disertacije upotpunjujuznanje o faunističkom sastavu, taksonomiji, distribuciji i diverzitetu osolikih muva u Grčkoj. Pomoću sakupljenih podataka formiran je identifikacioni ključ za razlikovanje kako rodova tako i vrsta registrovanih na ovom području, što u velikoj meri pomaže u daljim faunističkim istraživanjima. Kao rezultat ove doktorske disertacije dobijena je konačna lista od 418 vrsta iz 83 roda. Od ukupnog broja zabeleženih vrsta 64 vrste su prvi put registrovane na teritoriji Grčke, dok su 18 vrsta nove za nauku.<br />Osnovni cilj ovog rada je revizija faune osolikih muva Grčke na osnovu prikupljenog materijala iz publikovanih i nepublikovanih zbirki i dopunskih<br />terenskih istraživanja, sastavljanje liste o svim poznatim vrstama sa područja Grčke, i izrada identifikacionog ključa za razlikovanje zabeleženih rodova i vrsta<br />(praćeni fotografijama i crtežima dijagnostičkih morfoloških karaktera adultnih jedinki). Drugi deo doktorske disertacije se bavi prepoznavanjem retkih i<br />ugroženih vrsta Grčke sa naglaskom na njihovu zaštitu i konzervaciju. U okviru disertacije je izvršena prostorna analiza distribucije vrsta na području Grčke, koja su već ugrožena na evropskom i balkanskom nivou, a u cilju određivanja područja sa posebnim značajem za njihovo očuvanje.</p> / <p><!--[if gte mso 9]><xml> <w:WordDocument> <w:View>Normal</w:View> <w:Zoom>0</w:Zoom> <w:TrackMoves/> <w:TrackFormatting/> <w:PunctuationKerning/> <w:ValidateAgainstSchemas/> <w:SaveIfXMLInvalid>false</w:SaveIfXMLInvalid> <w:IgnoreMixedContent>false</w:IgnoreMixedContent> <w:AlwaysShowPlaceholderText>false</w:AlwaysShowPlaceholderText> <w:DoNotPromoteQF/> <w:LidThemeOther>EN-US</w:LidThemeOther> <w:LidThemeAsian>X-NONE</w:LidThemeAsian> <w:LidThemeComplexScript>X-NONE</w:LidThemeComplexScript> <w:Compatibility> <w:BreakWrappedTables/> <w:SnapToGridInCell/> <w:WrapTextWithPunct/> <w:UseAsianBreakRules/> <w:DontGrowAutofit/> <w:SplitPgBreakAndParaMark/> <w:DontVertAlignCellWithSp/> <w:DontBreakConstrainedForcedTables/> <w:DontVertAlignInTxbx/> <w:Word11KerningPairs/> 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Normal";mso-tstyle-rowband-size:0;mso-tstyle-colband-size:0;mso-style-noshow:yes;mso-style-priority:99;mso-style-qformat:yes;mso-style-parent:"";mso-padding-alt:0in 5.4pt 0in 5.4pt;mso-para-margin-top:0in;mso-para-margin-right:0in;mso-para-margin-bottom:10.0pt;mso-para-margin-left:0in;line-height:115%;mso-pagination:widow-orphan;font-size:11.0pt;font-family:"Calibri","sans-serif";mso-ascii-font-family:Calibri;mso-ascii-theme-font:minor-latin;mso-fareast-font-family:"Times New Roman";mso-fareast-theme-font:minor-fareast;mso-hansi-font-family:Calibri;mso-hansi-theme-font:minor-latin;mso-bidi-font-family:"Times New Roman";mso-bidi-theme-font:minor-bidi;}</style><![endif]--></p><p class="MsoNormal">Hoverflies are a very important and useful insect group. They provide irreplaceable environmental and economic services, as well as environmental services. These services are enabled by their ability to pollinate plants in natural and agroecosystems, as well as thanks to larvaepredators of aphids and other insects, reducing economic losses in the yield of cultivated plants. Hoverflies are extremely important for the environment, as they can serve as bioindicators of climate change and habitat conditions. Some species also have conservation importance, since they have become endangered due to anthropogenic effects, which requires taking measures to ensure their survival. The extraordinary richness of flora and fauna on the Balkan Peninsula represents a unique ecological and biogeographical phenomenon in Europe. Greece, as one of the most important center of biodiversity in the Balkans and in the Mediterranean, is characterized by a wide spectrum of important ecosystem types, the most important of which are forests, since the largest number of hoverflies prefer this type of habitat.The importance of faunal research is reflected in the discovery of the composition and number of species in specific habitats or areas. The faunal composition provides an insight into the biodiversity of the investigated area, which is constantly changing, especially lately under the influence of negative anthropogenic impact. Data about species can be of great importance for possible programs for the protection and recovery of endangered species, as well as in the planning of the formation of new protected areas or in the introduction of new measures of protection in already existing natural resources. Identifying endangered species or endangered habitats they inhabit plays an important role in planning conservation strategies for their conservation.The results of these thesis complement the knowledge of the fauna composition, taxonomy, distribution and diversity of hoverflies in Greece. Using the collected data, an identification key was formed to distinguish both the genera and species registered in this area, which greatly assisted in further faunistic research. As a result of these thesis a final list of 418 species rom 83 genus was obtained. Of the total number of recorded species, 64 species were first time registered in the territory of Greece, while a total of 18 species are new to science. The main goal of this paper is to revise the fauna of Greek howerflies based on the collected material from published and unpublished collections and additional field research, compile a list of all known species from Greece, and create an identification key to distinguish the recorded genera and species (accompanied by photographs and drawings of diagnostic morphological characters of adult individuals). The second part of the thesis deals with the identification of rare and endangered species of Greece with an emphasis on their protection and conservation. Within the framework of the dissertation, a spatial analysis of the distribution of species in Greece was carried out, already threatened at the European and Balkan levels, in order to determine areas of special<br />significance for their preservation.</p>
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Morfološka varijabilnost vrsta roda Anacamptis Rich. (Orchidoideae, Orchidaceae) na području Balkanskog poluostrva i Panonske nizije / Morphological variability of species of the genusAnacamptis Rich. (Orchidoideae, Orchidaceae) in the Balkan Peninsula and the Pannonian PlainRadak Boris 04 November 2019 (has links)
<p>Pored rodova evropskih orhideja koji se odlikuju velikim stepenom diverzifikacije, kao i značajnim brojem novoopisanih vrsta, kao što su <em>Ophrys </em>ili<em> Epipactis,</em> rod<em> Anacamptis </em>je ostao poprilično zapostavljen u orhidološkim istraživanjima. Na predstavnicima ovog roda uglavnom su vršena molekularna istraživanja, u kojima je bio uključen i veći broj drugih rodova.Takođe, urađene su i opsežne studije, ali samo na pojedinim vrstama. Morfološka istraživanja su bila veoma retka i uz par uzuzetaka, ograničena na geografski mala područja i na jednu ili nekoliko morfološki sličnih vrsta. Analiza morfološke varijabilnosti ovolikog broja taksona roda<em> Anacamptis</em> na geografski relativno velikom području kakvo je Balkansko poluostrvo, kao i deo Panonske nizije koji se oslanja na njega, do sada nije bilo sprovedeno. Morfološkim analizama bili su podvrgnuti gotovo svi predstavnici ovog roda u rangu vrste i podvrste, a koji se javljaju na području kopnenog dela Balkanskog poluostrva, kao i južnog oboda Panonske nizije. Sprovedene su jednostrane i uporedne morfološke analize sa detaljnomobradom podataka metodama bazične, univarijantne i multivarijantne statistike. Do sada najvećim istraživanjem taksona roda<em> Anacamptis</em>, po broju obrađenih jedinki i području na kom je sprovedeno, bila je obuhvaćena 2001 jedinka iz 185 populacija. Analizirane su prirodne populacije, ali i materijal iz Herbarijuma BUNS. Ukupno je definisano 69 morfoloških karaktera, kvantitativnih i kvalitativnih, a među njima su traženi oni koji imaju potencijalno diferencijalni karakter između opisanih vrsta i infraspecijskih taksona. Analize su obuhvatile i interpopulacionu morfološku varijabilnost u okviru svakog taksona, a u cilju uočavanja postojanja određenih geografskih obrazaca njenih promena. Morfološki karakteri koji su se u sprovedenim analizama pokazali kao taksonomski informativni, iskorišćeni su za formiranje dihotomog ključa za determinaciju taksona roda <em>Anacamptis</em> prisutnih na istraživanom području. Analizama je utvrđen veliki broj karaktera koji su statistički značajno različiti između analiziranih populacija, podvrsta i vrsta. Većina karaktera je pokazala umerenu ili nisku varijabilnost, a u retkim slučajevima i povećanu. Karakteri cveta međusobno su statistički značajno korelisani,većina slabo do umereno, dok su oni na listićima kacige, kao i između labeluma i drugih delova cveta jako do izrazito jako povezani. Jedini takson koji nije pokazao nikakvu infraspecijsku diferencijaciju, a koji se odlikuje velikom unutari interpopulacionom varijabilnošću je<em> A. pyramidalis</em>. Iz istog razloga je pokazano kao neosnovano izdvajanje jedinki iz različitih delova njegovog areala u do sada opisane infraspecijske oblike. Potvrđena je niska morfološka varijabilnost uskorasprostranjene vrste <em>A. boryi.</em> Kod svih preostalih istraživanih taksona utvrđeno je postojanje određenih obrazaca geografske interpopulacione varijabilnosti i infraspecijske struktuiranosti.Populacije <em>A. laxiflora</em> subsp. <em>laxiflora </em>mogu se podeliti na jadranske i kontinentalne, pri čemu najzapadnije pokazuju i najmanje vrednosti analiziranih karaktera. <em>A. coriophora </em>subsp.<em> fragrans </em>se pokazala kao manje varijabilnom od tipske podvrste, ali je u okviru nje kao zaseban takson izdvojen var. <em>hermae </em>kao fenološki, ekološki i morfološki odvojen u odnosu na ostaleispitivane populacije. Tipska podvrsta pokazuje obrazac variranja ukupne morfološke varijabilnosti u pravcu zapad-istok. Kao karakteri koji su najznačajniji za diferencijaciju podvrsta vrste<em> A. palustris</em> izdvojeni su dužina i širina brakteje, stepen diferenciranosti labeluma na režnjeve, kao i dubina sinusa koja ih razdvaja. Populacije<em> A. palustris</em> subsp. <em>elegans</em> moguće je podeliti na tri geografski i morfološki definisane grupe – istočnobalkansku, zapadnobalkansku i periferne populacije sa severa Srbije i iz Slovenije. Kao diferencijalni karakteri između taksona sekcije <em>Laxiflorae</em> pokazali su se odnos između dužine plodnika i brakteje, kao i dužina srednjeg režnja labeluma. Vrsta<em> A. morio </em>je po prvi put jasno morfološki i arealno podeljena na dve podvrste na istraživanom području. Podvrste pokazuju velika preklapanja vrednosti morfoloških karaktera, ali se obe odlikuju specifičnim karakteristikama cveta, pre svega labeluma na osnovu kojih ih je moguće razlikovati. Definisani su areali podvrsta<em> A.morio </em>na istraživanom području – tipska podvrsta je prevashodno severnija i zapadnija, dok je subsp.<em> caucasica </em>južnija i istočnija. Prvu odlikuju krupniji cvetovi, bubrežastih labeluma sa srednjim režnjem koji ne prevazilazi značajno dužinu bočnih, dok je druga sa sitnijim cvetovima i snažno isturenim srednjim režnjem. Obe podvrste mogu se podeliti na grupe populacija sa jasno definisanim geografskim raspostranjenjem. Vrsta <em>A. papilionacea </em>je po prvi put morfološki istaživana na većem broju primeraka, a ne prostim poređenjem pojedinačnih primeraka iz različitih delova areala. Pokazana je morfološka diferenciranost na dve podvrste – tipsku i subsp. aegaea, koje se jasno morfološki razlikuju i zauzimaju različite delove areala vrste. U okviru subsp. <em>papilionacea</em> registrovan je klinalni raspored morfološke varijabilnosti u pravcu sever-jug. Registrovana su i tri hibrida, nova za područje pojedinih zemalja – A. <em>× gennarii</em> (Severna Makedonija),A. <em>× parvifolia</em> (Crna Gora) i A. × <em>timbali</em> (Srbija). Hibridi pokazuju generalnu morfološku intermedijarnost u odnosu na roditelje, ali se odlikuju i novim stanjima karaktera, nezabeleženim kod roditeljskih taksona. Slični obrasci morfološke varijabilnosti, kako na interpopulacionom nivou pojedinih vrsta, ali iizmeđu podvrsta svake istraživane vrste, pokazuju jasnu vezu između istraživanih taksona i još jednom opravdavaju njihovo zajedničko grupisanje u rod<em> Anacamptis.</em></p> / <p>Unlike the highly diversified and species-rich genera of European orchids, such as<em> Ophrys </em>or <em>Epipac</em>tis, the genus <em>Anacamptis </em>has remained quite neglected in orchidological research. Representatives of this genus, together with many others genera, were investigated mainly by molecular methods. Also, extensive studies have been done, but only on specific species. Morphological studies were very rare, and with a few exceptions, limited to geographically small areas and one or more morphologically similar species. Analysis of the morphological variability of so many taxa of the genus <em>Anacamptis</em>, in a is, in a geographically relatively large area such as the Balkan Peninsula, as geographically relatively large area such as the Balkan Peninsula, as well as the southern part of the Pannonian Plain, has not been conducted so far. Morphological analyses were<br />performed on almost all representatives of this genus,in the species and subspecies rank , occurring in the Balkan Peninsula mainland, as well as in the southern perimeter of the Pannonian Plain. Comparativemorphological analyses were conducted using the methods of basic, univariate and multivariate statistics.So far, the largest study of the genus nacamptis, by the number of processed individuals and the area in which it was conducted, included 2001 individuals from 185 populations. Natural populations and material from the BUNS Herbarium were analyzed. A total of 69 morphological characters, quantitative and qualitative, were defined, and those that have a potentially differential character among analyzed species and infraspecific taxa were selected. The analyzes also included interpopulation morphological variability within each taxon, to identify the existence of specific geographic patterns of its changes. The morphological characters that proved to be taxonomically informative, were used to form the dichotomous key for the determination of the genus Anacamptis members, presented in the study area. The analyzes identified a large number of characters that were statistically significantly different between the analyzed populations, subspecies, and species. Most of the characters showed moderate or low variability, and in rare cases increased variability. Flower’s characters were statistically significantly correlated with each other, most of them weakly to moderately, while those of the sepals and petals, as well as between the labellum and other parts of the flower, were very strongly correlated. The only taxon that has not shown any infraspecific differentiation (except <em>A. boryi</em>), and which is characterized by high intra- and interpopulation variability, is A. pyramidalis. For the same reason, it has been shown as unjustified to separate individuals from different parts of species range into infraspecific forms. The low morphological variability of the narrowly distributed A. boryi was confirmed. In all other taxa studied, the existence of certain patterns of geographical interpopulation variability and infraspecific structuring was detected. <em>A. laxiflora</em> subsp. <em>laxiflora</em> populations can be divided into two groups – Adriatic and Continental, with the lowest characters values measured among the westernmost populations. <em>A. coriophora s</em>ubsp. <em>fragrans</em> proved to be less variable than the type subspecies, but within it, a new variety (var. hermae) as phenological, ecological and morphologically distinct from other studied populations was described. The typesubspecies has a transition of morphological variability in a west-east direction. The characters that are the most important for differentiation of A. palustris subspecies are the bract length and width, the degree of labellum differentiation into lobes, and the depth of the sinuses that separate them. Populations of A. palustris subsp. elegan<em>s </em>can be divided into three geographically and morphologically defined groups – Eastern Balkan, Western Balkan and peripheral populations from the North Serbia and Slovenia. Differential characters among the Laxiflorae section taxa are ration between the ovary length and bract length, as well as the labellum middle lobe length. For the first time, species <em>A. morio </em>is morphologically and geographically divided into two subspecies, in the study area. Analyzed subspecies have great overlapping values for many morphological characters, but both are characterized by specific flower’s characteristics, above all labellum shape. On this basis, it is possible to distinguish them. Areals of <em>A</em>. <em>morio </em>subspecies were defined – the type subspecies is predominantly northern and western,while subsp. caucasica is southern and eastern. The first is characterized by larger flowers, renal shape labellum with a median lobe that does not exceed significantly the length of the lateral ones, while the second subspecies has smaller flowers and strongly projecting the median lobe. Both subspecies can be divided into groups of populations that have clearly defined geographical distribution. For the first time, A. papilionacea was morphologically investigated on a larger number of specimens, unlike previous researches in which individual specimens from different parts of the range were simply compared. Morphological differentiation into two subspecies was shown – type and subsp. aegaea. They are morphologically distinct and occupy different parts of the species range. Within the subsp. papilionacea a clinal distribution of morphological variability in the north-south direction was recorded. Three taxa of hybrid origin, new for the area of North Macedonia (A.<em> × gennari</em>i), Montenegro (A. <em>× parvifolia</em>) and Serbia (A. × t<em>imbali</em>) were registered. Hybrids exhibit a general morphological intermediacy between parents but are also distinguished by new character states that were not observed in<br />parental taxa. Similar patterns of morphological variability, both at the interpopulation level ofindividual species and between the subspecies of each species studied, show a clear relationship between the studied taxa and once again justify their grouping into the genus <em>Anacamptis.</em></p>
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Nuotolinių studijų kurso Programavimas grafinėje terpėje reinžinerija / Reengineering the Distance Study Course‚ Programming in GUI‘Demenis, Tomas 29 September 2008 (has links)
Šiame magistro darbe mes analizuojame nuotolinių studijų kurso bendrai ir mokymo medžiagos atskiromis kurso dalimis pertvarkymą. Reinžinerijos koncepcinis modelis yra įvairiai interpretuojamas ir yra taikomas programinės įrangos rengimo, ar vadybos moksluose. Tai yra daroma, kad sistemas būtų galima geriau panaudoti. Jos analizuojamos ir pertvarkomos tolimesniam naudojimui. Mes sujungiame minėtų sričių reižinerijos koncepto reikšmes ir naudojame tai kaip metodinį pagrindą nuotolinių studijų srityje. Mes analizuojame nuotolinių studijų kurso struktūrą. Tada pristatome kursus paruoštus ir naudojamus užsienio universitetuose, susijusius su ‘Programavimu grafinėje terpėje‘, trigubo pastovumo principu, Bloom taksonomija ir jos pritaikymu kompiuterijos mokslų studijavimui, informatikos mokymo programa 2001 (angl. computing curriculla 2001). Toliau, mes siūlome konceptualią nuotolinių studijų pertvarkos struktūrą iš dėstytojo perspektyvų ir pristatome atvejų analizę, kurioje pora temų yra pertvarkytos, atsižvelgiant į trigubo pastovumo principą ir reikalavimus kompiuterių mokslų studentams. / In this master thesis we analyse a problem of reengineering of a distance study system, in general, and the learning material of a separate course, in particular. Reengineering concept with its different interpretations is used in software engineering and management sciences. It deals with making systems better maintainable, examination and reconstitution of the system for further reimplementation. We combine the meaning of reengineering concept in both mentioned areas and employ it as methodological background in distance study area. We analyse the structure of the distance study course. Then we introduce the courses, prepared and delivered in foreign universities and related to ‘Programming in GUI’ course, triple consistency principle, Bloom taxonomy and its applicability to computer science studies, Computing Curricula 2001. Further, we propose a conceptual distance study course reengineering framework from the lecturer’s perspective and present a case study, in which two topics were reengineered, considering triple consistency principle and requirements for computer science students.
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Uporedni pregled morfo-anatomskih karakteristika biljnih organa i analiza etarskih ulja sa njihovom primenom u taksonomiji odabranih rodova tribusa Inuleae Cass. (Compositae) / Comparative review of morpho-anatomical characteristics of plant organs and analyses of essential oils with their application in taxonomy of selected genera of the tribe Inuleae Cass. (Compositae)Karanović Dunja 27 September 2017 (has links)
<p>Komparativnom analizom morfo-anatomskih i mikromorfoloških karakteristika lista, stabla, rizoma, korena, ploda i receptakuluma, kao i biohemijskom analizom etarskih ulja podzemnih organa, obuhvaćeno je ukupno 16 vrsta iz 4<br />roda, Inula (I. aschersoniana, I. bifrons, I. britannica, <em> I. conyza, I. ensifolia, I. germanica, I. helenium, I. oculus-christi, I. saslicina ssp. salicina, I. salicina ssp. aspera, I. spiraeifolia), Dittrichia (D. graveolens, D. viscosa), Limbarda (L.<br />crithmoides) i Pulicaria (P. dysenterica, P. vulgaris) iz</em> tribusa Inuleae, familije Compositae. Vrste iz različitih rodova analizirane u ovoj disertaciji ranijom klasifikacijom svrstavane su u okviru različitih sekcija roda Inula, a takođe i tipska vrsta ovog roda,<em> I. helenium L</em>., svrstavana je u okviru zasebne sekcije<em> Corvisartia.</em> Na osnovu najnovijih molekularnih filogenetskih analiza, baziranih na sekvencama plastidne i jedarne DNK, kao i na osnovu morfoloških i karioloških podataka, predloženo je da sekcija <em>Corvisartia </em> bude izdvojena u zaseban rod. Ciljevi ove<br />disertacije definisani su u skladu sa činjenicom da su podaci o morfo-anatomskoj, mikromorfološkoj i biohemijskoj građi vegetativnih i reproduktivnih organa vrsta ovog tribusa malobrojni i nepotpuni, dok su analize vršene parcijalno i<br />nezavisno kod pojedinih predstavnika, sa slabim akcentom na njihovu komparativnu analizu. Istraživanje je obuhvatilo veliki broj kvalitativnih i kvantitativnih karaktera koji su analizirani i mereni pomoću stereomikroskopa, svetlosnog, fluorescentnog i skening elektronskog mikroskopa. Dobijeni podaci statistički su obrađeni različitim metodama numeričke analize koje se inače<br />koriste pri taksonomskim istraživanjima. Ovakav tip analize za vrste rodova <em> Inula,</em> <em>Dittrichia, Limbarda i Pulicaria</em> nije zabeležen u dosadašnjoj literaturi.<br />Sprovedena analiza pokazala je da ni jedan vegetativni biljni organ pojedinačno ne pruža dovoljno informacija na osnovukojih bi se sa statističkom sigurnošću mogle izdvojiti određene grupe vrsta, niti rodovi prema postojećoj podeli. U okviru analiziranih anatomskih i mikromorfoloških karakteristika ploda i receptakuluma, organizacija sklerenhimskog tkiva u zrelom plodu i karakteristike receptakuluma predstavljaju karaktere koji su dijagnostički za rodove. Karakteri koji razdvajaju analizirane taksone jesu: karakteristike papusa kod vrsta roda <em> Pulicaria; </em>prstenasato suženje između baze papusa i cipsele, organizacija<br />sklerenhimskog tkiva ploda i tip trihoma na površini receptakuluma kod vrsta roda<em> Dittrichia</em>; sekretorni kanali u perikarpu ploda, odsustvo trihoma i slabo izražena rebra na površini receptakuluma kod vrste<em> L. crithmoides</em>. Plod vrste <em> I.</em><br /><em>helenium</em> razlikuje se od plodova svih preostalih analiziranih vrsta na osnovu njegovih dimenzija, većeg broja sklerenhimskih vrpci koje nisu istaknute u rebra, kao i na osnovu slabo izraženih rebara na receptakulumu, između karpopodijalnih otisaka, na kojima se nalaze žlezdane i nežlezdane trihome. Rezultati su<br />ukazali na to da se vrsta I. helenium izdvaja kao zaseban takson na osnovu anatomskih i mikromorfoloških karakteristika ploda i receptakuluma sa jednako malim stepenom statističke podrške koji takođe podržava izdvajanje rodova <em>Dittrichia, Limbarda i Pulicaria</em> u zasebne rodove. Stoga, naši rezultati podržavaju tvrdnju da vrsta I. helenium treba biti izdvojena od preostalih<br />vrsta roda Inula u zaseban rod, pri čemu je njena autentičnost evidentna na nivou anatomije i mikromorfologije vegetativnih i reproduktivnih organa.<br />Rezultati hemijske analize etarskog ulja pružili su do sada nepoznate podatke o kvalitativnom i kvantitativnom sastavu etarskog ulja podzemnih organa analiziranih taksona i ukazali na mogućnost njihove primene u taksonomiji ispitivanog tribusa na nivou identifikacije pojedinačnih taksona. Takođe, dobijeni<br />rezultati ukazali su na nove potencijalno lekovite vrste koje predstavljaju potencijalni resurs za farmaceutsku industriju. Rezultati analize osnovnih komponenti kao i rezultati korespodentne analize ukazali su na odsustvo tendencije grupisanja taksona koji pripadaju istom rodu prema trenutno<br />važećoj klasifikaciji. Zbog velike varijabilnosti uslovljene faktorima spoljašnje sredine sastav etarskih ulja ne može se koristiti samostalno kao pouzdan hemotaksonomski marker. Ipak, rezultati o sastavu etarskih ulja značajni su u smislu njihove primene u medicinske i kozmetičke svrhe, u industriji<br />gde se etarska ulja koriste kao sirovina, te su dobijeni rezultati značajni i u kontekstu evaluacije kvaliteta etarskih ulja ispitivanih vrsta.<br />Sprovedeno istraživanje daje značajan doprinos u pogledu tačnije klasifikacije unutar tribusa koja se danas bazira uglavnom na morfološkim karakteristikama. Detaljno opisani morfo-anatomski i mikromorfološki karakteri analiziranih<br />vegetativnih i reproduktivnih organa doprinose boljem poznavanju karakteristika vrsta analiziranih rodova i predstavljaju dopunu postojećim podacima o analiziranimtaksonima. Takođe, na bazi morfo-anatomskih karakteristika ploda i receptakuluma izrađen je dihotomi ključ za determinaciju analiziranih rodova i vrsta. Korišćenjem metoda diskriminantne i korespodentne analize izdvojeni su i karakteri na osnovu kojih je moguće izvršiti diskriminaciju određenih grupa u okviru<br />analiziranih taksona.</p> / <p>A comparative analysis of morpho-anatomical and micromorphological characteristics of leaf, stem, rhizome, root, fruit and receptacle, as well as biochemical analysis of essential oils of underground organs, was conducted over 16 species of 4 genera, <em> Inula (I. aschersoniana, I. bifrons, I. britannica, I.conyza, I. ensifolia, I. germanica, I. helenium, I. oculus-christi, I. saslicina ssp. salicina, I. salicina ssp. aspera, I. spiraeifolia), Dittrichia (D. graveolens, D. viscosa), Limbarda (L. crithmoides) and Pulicaria (P. dysenterica, P. vulgaris)</em> tribe Inuleae, Compositae. In addition to the fact that the species from different genera analysed in this paper were previously placed in different sections within <em>Inula, I. helenium L</em>., as the type species of the genus <em> Inula,</em> has also been placed in a separate section Corvisartia. According to the latest molecular phylogenetic analyses, based on plastid and nuclear DNA sequence data, as well as morphological and karyological data, it is suggested that section Corvisartia should be segregated from the remaining Inula species as a separate genus, thus achieving a greater morphological homogeneity of the genus <em>Inula. </em> The objectives of dissertation are defined in accordance with the fact that the data on the morpho-anatomical, micromorphological and biochemical structure of the vegetative and reproductive organs of species of examined tribe are sparse and incomplete, while the analyses were carried out on few species of <em>Inula, Pulicaria, Dittrichia</em> and<em> Limbarda</em>, but less emphasis was placed on comparative purpose. The research involved a large number of qualitative and quantitative characters that were analysed and measured using a stereomicroscope, light, fluorescent and scanning electron microscope. The obtained data were statistically processed by various numerical analysis methods usually used in taxonomic research. These types of analyses for the species of the generaInula<em>, Dittrichia, Limbard </em> and <em> Pulicaria </em> have not been recorded in the literature so far. The conducted analysis showed that vegetative organs do not provide enough information on the basis of which certain groups of species could be singled out with statistically significant suport, neither the genera according to the actual classification. The yielded results indicated that the species from different genera were clearly delimited on the basis of their carpological and receptacular micromorphological and anatomical characteristics. Receptacle features and organisation of sclerenchyma tissue in a fruit are traits that tend to be diagnostic for genera. The features that separate analysed taxa are: two rows of pappus hairs in <em> Pulicaria </em>species; the ring constriction of the fruit over which it passes into pappus, the sclerenchymatous tissue in fruit in form of continuous ring, the type of trichomes on the receptacular surface in <em> Dittrichia </em>species; the fruit with secretory ducts, and the receptacular surface with weakly pronounced ridges without glands in L. crithmoides. I. helenium fruits differ from those of all the other species by their size, greater number of inconspicuous sclerenchymatic bundles, and poorly expressed ridges among carpopodial prints with glandular and non-glandular trichomes distributed along them. Results revealed that I. helenium stands out as a separate taxon, based on the anatomical and micromorphological characteristics of its fruit and receptacle, with equally small percentage of statistical support, which also supports the recognition of<em> Pulicaria, </em> <em>Dittrichia</em> and<em> Limabarda</em> as separate genera. Obtained results support the claim that I. helenium should be separated from the Inula genus, whereby its authenticity is evident at the level of anatomy and micromorphology of vegetative and reproductive organs. The chemical analysis of the essential oil has provided new data on the qualitative and quantitative composition of the essential oil of the underground organs of the analysed taxa and pointed out the possibility of their application in taxonomy. Also, the obtained results indicated potentially new medicinal plants that represent a potential resource for the pharmaceutical industry. Principal component analysis as well as the results of the Correspondent analysis indicated the absence of the grouping tendency of taxa belonging to the same genus, according to the actual classification. Due to the high variability conditioned by the environmental factors the composition of essential oils cannot be used independently as a reliable chemotaxonomic marker. However, the results of the composition of essential oils are significant in terms of their application for medical and cosmetic purposes, in the industry where oils are used as raw materials, and the results obtained are also significant in the context of evaluation of the quality of investigated essential oils.The conducted research makes a significant contribution to more precise classification within the tribe, which is now basedmainly on morphological characteristics. The detailed description of morpho-anatomical and micromorphological characters of the analysed vegetative and reproductive organs contributes to better knowledge of the characteristics of the analysed species and complements the existing data on them. Also, on the basis of morpho-anatomical characteristics of the fruit and the receptacle a dichotomous key for the determination of the analysed genera and species was developed. According to the Discriminant and Correspodent analysis the characteristics on which is possible to discriminate certain groups within the analysed genera are also distinguished.</p>
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Таксономија тестатних амеба које насељавају маховине на подручју Источне Херцеговине / Taksonomija testatnih ameba koje naseljavaju mahovine na području Istočne Hercegovine / Taxonomy of moss-dwelling testate amoebae from East HerzegovinaLuketa Stefan 08 December 2020 (has links)
<p>Докторска дисертација представља таксономску студију тестатних амеба Источне Херцеговине базирану искључиво на резултатима сопствених истраживања с обзиром да на овом подручју тестатне амебе до сада нису проучаване, те не постоје историјске музејске колекције. На подручју Источне Херцеговине регистровано је 40 врста тестатних амеба које су сврстане у 10 фамилија и један род без јасног места у класификационом систему. Сви регистровани таксони тестатних амеба су нови за фауну Босне и Херцеговине. Укупно је анализирано 24.549 јединки, од чега су 23.242 јединке припадале групи тестатних амеба са лобоподијама (супергрупа Amoebozoa), а 1307 јединки је припадало групи тестатних амеба са филоподијама (супергрупа Cercozoa). Најзначајнији резултат ове дисертације је опис пет нових врста за науку које припадају родовима<br />Centropyxis, Heleopera и Nebela. Морфотип означен као C. cf. aerophila се од врсте C. aerophila разликује по томе што се на крају љуштурице не налази пар крупних честица кварца,а и љуштурица је нешто дужа (46‒81 μm код врсте C. aerophila према 67‒ 88 μm код врсте C. cf. aerophila). Морфотип означен као C. cf. platystoma значајно се пре свега морфолошки разликује од врсте C. platystoma, те је закључено да се ради о неописаној врсти.У оквиру рода Heleopera описан је нови морфотип сличан врсти H. rosea који представља нову врсту за науку. Морфометријске разлике су релативне, тј. нису строго дискриминаторне, те се морају комбиновати са морфолошким разликама које су такође тешко yочљиве. Наиме, поред разлике у боји љуштурице, најбољи дискриминаторни морфолошки карактер је општи облик љуштурице. Љуштурице врсте H. rosea су робусног облика, док су љуштурице врсте Heleopera cf. rosea знатно елегантније ‒ уже су и имају облије ивице. Највеће морфометријске разлике у индексним карактерима су забележене за однос ширине и дужине љуштурице и однос ширине апертуре и ширине љуштурице.Морфотип Nebela cf. collaris се од врсте N. collaris јасно разликује пре свега морфолошки и еколошки, а морфометријски веома мало. Наиме, најважнија морфолошка одлика која морфотип N. cf. collaris раздваја од врсте N. collaris су таласасте ивице љуштурице, а еколошка разлика се јавља у смислу да врста N. collaris насељава зелене маховине док морфотип N. cf. collaris насељава сфагнумске маховине. Морфотип N. cf. tincta var. major се од морфотипа N. cf. collaris разликује пре свега по јасно израженом сужењу у делу близу апертуре, тј. израженом врату. Такође, морфотип N. cf. tincta var. major никада нема таласасте ивице љуштурице,док се код јединки морфотипа N. cf. collaris ова карактеристика често јасно уочава.</p> / <p>Doktorska disertacija predstavlja taksonomsku studiju testatnih ameba Istočne Hercegovine baziranu isključivo na rezultatima sopstvenih istraživanja s obzirom da na ovom području testatne amebe do sada nisu proučavane, te ne postoje istorijske muzejske kolekcije. Na području Istočne Hercegovine registrovano je 40 vrsta testatnih ameba koje su svrstane u 10 familija i jedan rod bez jasnog mesta u klasifikacionom sistemu. Svi registrovani taksoni testatnih ameba su novi za faunu Bosne i Hercegovine. Ukupno je analizirano 24.549 jedinki, od čega su 23.242 jedinke pripadale grupi testatnih ameba sa lobopodijama (supergrupa Amoebozoa), a 1307 jedinki je pripadalo grupi testatnih ameba sa filopodijama (supergrupa Cercozoa). Najznačajniji rezultat ove disertacije je opis pet novih vrsta za nauku koje pripadaju rodovima<br />Centropyxis, Heleopera i Nebela. Morfotip označen kao C. cf. aerophila se od vrste C. aerophila razlikuje po tome što se na kraju ljušturice ne nalazi par krupnih čestica kvarca,a i ljušturica je nešto duža (46‒81 μm kod vrste C. aerophila prema 67‒ 88 μm kod vrste C. cf. aerophila). Morfotip označen kao C. cf. platystoma značajno se pre svega morfološki razlikuje od vrste C. platystoma, te je zaključeno da se radi o neopisanoj vrsti.U okviru roda Heleopera opisan je novi morfotip sličan vrsti H. rosea koji predstavlja novu vrstu za nauku. Morfometrijske razlike su relativne, tj. nisu strogo diskriminatorne, te se moraju kombinovati sa morfološkim razlikama koje su takođe teško yočljive. Naime, pored razlike u boji ljušturice, najbolji diskriminatorni morfološki karakter je opšti oblik ljušturice. LJušturice vrste H. rosea su robusnog oblika, dok su ljušturice vrste Heleopera cf. rosea znatno elegantnije ‒ uže su i imaju oblije ivice. Najveće morfometrijske razlike u indeksnim karakterima su zabeležene za odnos širine i dužine ljušturice i odnos širine aperture i širine ljušturice.Morfotip Nebela cf. collaris se od vrste N. collaris jasno razlikuje pre svega morfološki i ekološki, a morfometrijski veoma malo. Naime, najvažnija morfološka odlika koja morfotip N. cf. collaris razdvaja od vrste N. collaris su talasaste ivice ljušturice, a ekološka razlika se javlja u smislu da vrsta N. collaris naseljava zelene mahovine dok morfotip N. cf. collaris naseljava sfagnumske mahovine. Morfotip N. cf. tincta var. major se od morfotipa N. cf. collaris razlikuje pre svega po jasno izraženom suženju u delu blizu aperture, tj. izraženom vratu. Takođe, morfotip N. cf. tincta var. major nikada nema talasaste ivice ljušturice,dok se kod jedinki morfotipa N. cf. collaris ova karakteristika često jasno uočava.</p> / <p>The PhD thesis is a taxonomic study of testate amoebae from East Herzegovina based exclusively on the results of our own research, given that testate amoebae have not been studied in this region so far, and there are no historical museum collections. In the region of East Herzegovina,40 testate amoeba species have been registered, which are classified into 10 families and one genus without a clear place in the classification system. All registered testate amoeba taxa are new to the fauna of Bosnia and Herzegovina. A total of 24,549 individuals belonged to the group of testate amoebae with lobopodia (supergroup Amoebozoa), and 1307 individuals belonged to the group of testate amoebae with filopodia (supergroup Cercozoa). The most significant results of this PhD thesis are the descriptions of five new species for science belonging to the genera Centropyxis, Heleopera, and Nebela. The morphotype Centropyxis cf. aerophila differs from C. aerophila in that there is no large quartz particles at the shell end, and the shell is slightly longer (46‒81 μm in C. aerophila versus 67‒88 μm in C. cf. aerophila). The morphotype C. cf. platystoma differs significantly morphologically from C. platystoma, so it was concluded that it is an undescribed species. Within the genus Heleopera a new morphotype similar to H. rosea has been described, representing a new species for science. Morphometric differences are relative, i.e. they are not strictly discriminatory, and must be combined with morphological differences that are difficult to detect. Namely, in addition to the difference in the color of the shell, the best discriminatory morphological character is the general shell shape. Shells of H. rosea are red and robust in shape, while shells of H. cf. rosea are volet and much more elegant ‒ they are narrower and have rounded edges. The largest morphometric differences in the index characters were observed for shell width/shell length ratio and aperture width/shell width ratio. The morphotype Nebela cf. collaris clearly differs from N. collaris primarily morphologically and ecologically, but morphometrically very little. Namely, the most important morphological character that N. cf. collaris separates from N. collaris are the wavy edges of the shell, and the ecological difference occurs in the sense that N. collaris inhabits green mosses while N. cf. collaris inhabits Sphagnum mosses. The morphotype N. cf. tincta var. major from the morphotype N. cf. collaris differs primarily by a clearly pronounced narrowing in the part near the aperture, i.e. pronounced neck. Also, the morphotype N. cf. tincta var. major never has a wavy edge of the shell, while in N. cf. collaris this feature is often clearly observed.</p>
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Visoke kognitivne funkcije u nastavi lingvistiĉkih predmeta na tercijarnom nivou obrazovanja / Higher cognitive functions in linguistic courses in tertiary educationVotls Isidora 22 June 2016 (has links)
<p>Iskustvo u radu sa studentima pokazalo je da studenti nedovoljno ĉesto ostvaruju zadovoljavajuće ishode uĉenja na lingvistiĉkim predmetima na studijama engleskog jezika, što je takoĊe opisano i u stranoj literaturi. Kao jedan od razloga navodi se nastava koja upućuje studente na pasivnost, te oni pribegavaju memorisanju i reprodukciji materijala kao najĉešćim ishodima uĉenja. Biggs (1999) razraĊuje koncepte dubinskog i površinskog pristupa uĉenju, koje relevantna literatura smatra kljuĉnim faktorima za kvalitet ishoda uĉenja. Dubinski pristup uĉenju korelira sa kvalitetnim ishodima uĉenja i funkcionalnim znanjem, a karakterišu ga motivacija, zadovoljstvo usled uĉenja, studentska aktivnost i to aktivnost visokih kognitivnih funkcija. Visoke kognitivne funkcije (Bloom i dr. 1956, Anderson i dr. 2001) i povezane kognitivne radnje (rešavanje problema, analitiĉko, kritiĉko i kreativno razmišljanje) jesu najvaţniji ciljevi visokog obrazovanja jer samo one, usled primene transfera uĉenja, govore o steĉenom i primenljivom, tj. funkcionalnom znanju. Obuka i razvoj visokih kognitivnih funkcija omogući će studentima da uĉenju pristupe dubinski što je još jedan razlog da budu osnovni nastavni cilj svih predmeta na ustanovama tercijarnog obrazovanja. U skladu sa ovim teorijskim postavkama postavljene su osnovna i pomoćna hipoteza: upotreba posebno konstruisanih veţbi za aktivaciju viših kognitivnih funkcija u nastavi lingvistiĉkih predmeta dovešće do sticanja funkcionalnog znanja na teorijskom i praktiĉnom nivou; steĉeno znanje kroz ovakvu eksperimentalnu nastavu i upotreba tog znanja odraţavaće kognitivne funkcije ne samo niţeg nego i višeg reda: primeniti, analizirati, proceniti, stvoriti, kao i kritiĉko i kreativno razmišljanje i rešavanje problema. Kako bi se proverile hipoteze, sproveden je eksperiment sa studentima prve godine engleskog jezika (N=34) na Fakultetu za pravne i poslovne studije dr Lazar Vrkatić u Novom Sadu. U istraţivanju sa paralelnim grupama, eksperimentalna grupa je imala veţbe sa aktivnostima koje razvijaju više kognitivne funkcije na predmetu uvod u opštu lingvistiku tokom zimskog semestra školske 2012/2013. godine. UporeĊeni su kvantitativni rezultati kolokvijuma eksperimentalne i kontrolne grupe na kraju semestra, a potom je sproveden intervju sa po pet studenata iz svake grupe radi utvrĊivanja kvalitativnih razlika u kognitivnim procesima kod ove dve grupe. Obe grupe su ostvarile podjednak uspeh na kolokvijumu, te je osnovna hipoteza odbaĉena. Kodirani podaci iz intervjua pokazali su da obe grupe podjednako koriste kognitivne funkcije po broju i distribuciji, te je i pomoćna hipoteza odbaĉena. Kao objašnjenje za odsustvo većeg uspeha EG navedena su metodološka ograniĉenja istraţivanja: duţina eksperimentalne nastave, problem dokazivosti transfera i problem kodiranja intervjua. Drugi faktori koji mogu objasniti neuspeh su: prethodno steĉene navike u uĉenju, neshvatanje svrhe izuĉavanja predmeta i dr. UporeĊeni su rezultati boljih i slabijih studenata, te je utvrĊeno da bolji studenti pokazuju veći stepen samostalnosti, da upotrebljavaju više kognitivne funkcije kao i duţe nizove kognitivnih radnji. Posmatrajući kvalitativne podatke, bolji studenti eksperimentalne grupe pokazali su promenu gledanja na svet usled izuĉavanja lingvistike i izrazili su zadovoljstvo zbog uĉenja ovog predmeta. Oni pokazuju i upotrebu najduţih nizova vezanih kognitivnih radnji. Iz ovoga se moţe zakljuĉiti da su oni pristupili uĉenju dubinski i zbog toga ostvarili kvalitetnije ishode uĉenja. U cilju donošenja ĉvrstih zakljuĉaka neophodno je sprovesti dugotrajniji i obuhvatniji multidiciplinarni istraţivaĉki projekat, s obzirom da bi pozitivni rezultati bili od velikog znaĉaja za poboljšanje ishoda uĉenja na tercijarnom nivou obrazovanja. Ključne reči: uĉenje i nastava na tercijarnom nivou, taksonomija obrazovnih ciljeva, pristupi uĉenju, više kognitivne funkcije, transfer uĉenja, funckionalno znanje.</p> / <p>The experience of working with university students has shown that the learning outcomes of linguistic courses are infrequently satisfactory, which is also described in literature worldwide. Teaching philosophy in which students are forced into passives roles is one of the causes since such teaching results in low motivation with memorizing and reproduction of learned materials as the most frequent outcomes of learning. Biggs (1999) develops the concepts of deep and superficial learning approaches which have been declared in the relevant literature as key factors for the quality of learning outcomes. Deep approach to learning correlates with high quality learning outcomes, and is characterized by high motivation, satisfaction with learning and student activity of appropriately high cognitive levels. Higher cognitive functions (Bloom et. al. 1956, Anderson at al. 2001) and related cognitive activities (problem solving, analytical, critical and creative thinking) are the most important goals of higher education since these thinking skills are transferable and therefore represent applicable and functional knowledge. The training and development of the higher cognitive skills enables students to use deep approaches to learning, which is an additional reason to consider them as fundamental teaching goals in all courses in tertiary education. Based on this theoretical framework the main hypothesis and sub-hypothesis were formulated as follows: the use of specially designed practices which activate higher cognitive functions (HCF) will result in acquiring functional knowledge at both theoretical and practical levels; the knowledge gained through such teaching will reflect the use of higher cognitive functions: apply, analyze, evaluate, create, as well as show problem solving skills and critical and creative thinking. To test the hypotheses an experiment was conducted with the first year English language students (N=34) at the Faculty of Legal and Business Studies dr Lazar Vrkatić in Novi Sad. In the parallel groups design, the experimental group (EG) was involved with activities which develop HCFs in the course of Introduction to General Linguistics during the winter semester of the 2012/2013. Quantitative data were collected at the end of the semester (the final test) and compared between the two groups to determine whether the EG scored better results than the control group (CG). This was followed by interviews with five respondents from each group to qualitatively compare the cognitive processes. No statistically significant difference between test results in the two groups was found and so the main hypothesis was rejected. The coded data from the interviews showed an equal number of identified CFs with both groups with similar distribution patterns, thus the sub-hypothesis was also rejected. The absence of better scores of the EG can be explained by some methodological limitations of the experiment, such as the length of the experimental activities, the problem of proof of transfer and the coding of the interview data. Other factors include the existing learning habits of students, the inability to grasp the purpose of studying linguistics, etc. The results of better students were compared to those of the weaker ones, which showed that better students are more autonomous, use a greater number of HCFs and string more CFs into a complex response. Qualitative data also showed that better students of the experimental group expressed a change in how they see the world around them and express satisfaction because of studying linguistics. They also string the longest chains of cognitive activities. These findings lead to a conclusion that better students of the EG used deep approaches to learning which resulted in higher quality learning outcomes. In order to achieve conclusive results, a comprehensive long-term multidisciplinary research project should be carried out, since its results would have a significant impact on the quality of learning outcomes in tertiary education.</p>
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