Five species, Acacia karroo, A. robusta, A. nilotica, A. sieberana and A. tortilis, were inoculated with Rhizobium and grown in potted sand in a temperature controlled greenhouse. After six months, results showed a higher percentage plant nitrogen for all five species when inoculated plants were compared to uninoculated controls. Inoculated treatments of A. karroo and A. sieberana had the greatest growth in shoot length and biomass. Acacia robusta showed the highest nitrogenase activity when nodules were tested using acetylene reduction methods. Inoculants of A. tortilis showed the poorest growth for all parameters measured. A. karroo and A. nilotica were studied at a field site at Ashburton, 15km east of Pietermaritzburg. Acacia karroo and A. nilotica had similar mean percentage leaf nitrogen, but A. karroo had a significantly higher mean percentage stern nitrogen than A. nilotica. Rainfall, canopy throughfall and stemflow from A. karroo and A. nilotica were collected in late spring and examined for inorganic nitrogen content. Acacia nilotica yielded the highest nitrate levels in both throughfall and stemflow samples. Acacia karroo produced lower nitrate concentrations in samples of both throughfall and stemflow, than was found in rainfall. Both A. nilotica and A. karroo exhibited higher concentrations of ammonium in samples of throughfall and stemflow as compared to levels. Soil analyses yielded highest levels of organic nitrogen at the surface (0 - 5 cm) but this decreased significantly at 20 cm deep. Surface organic nitrogen was highest under A. karroo canopies and lowest in open grassland. At 20 cm, there was little difference in organic nitrogen content between soils sampled from open patches and those under canopies of A. nilotica or A. karroo. Nitrate showed little variation with species, but highest levels were found in the top five centimetres and levels were higher under grasslands than under canopies. Ammonium showed no significant differences between different depths but was higher in open grassland sites than under canopies. No pattern could be found to relate tree size to soil organic nitrogen content. / Thesis (M.Sc.)-University of Natal, Pietermaritzburg, 1995.

In the past, protected natural areas have excluded local communities from the land and denied them access to valuable natural resources. However, it is becoming accepted practice to ensure that neighbouring communities benefit from the conservation of these areas. In accordance with their neighbour relations programme, the Natal Parks Board initiated a study to establish the need for indigenous wood in the region of the Thukela Biosphere Reserve (TBR), particularly for the woodcarving industry, and to determine sustainable methods and levels of harvesting. Part of this study was to determine the socio-economic issues surrounding the woodcarving industry and other users of indigenous trees, and these are addressed in this thesis. A multidisciplinary approach was adopted to address as many aspects of natural resource use as possible. The principle of sustainable development was employed to explore the nature of the often complex relationships between local communities and protected areas, and local communities and natural resource use. This principle calls for the integration of social, economic and ecological issues, with special attention to the notions of futurity, equity and the environment. The biosphere reserve is considered to be an appropriate vehicle for achieving sustainable development and the sustainable utilisation of resources, both internationally and in the South African context. However, in practice there are many obstacles to overcome as was observed in the case of the TBR, where security of land tenure and the associated control of and access to natural resources are a source of major conflict in the area. In view of this conflict, a flexible and sensitive methodology that promoted rapport-building was selected, namely Rapid Rural Appraisal (RRA). Mainly verbal RRA techniques were used to gather information on the use of and demand for indigenous trees by the local communities residing within and surrounding the TBR. This information included species names, species uses, estimations of quantities harvested, perceptions of the resource base, conservation practices and harvesting techniques, economic relations, constraints, and relationships between the resource manager and the resource user. Indigenous trees were found to be an important resource for fuel, construction, medicine, carving, and to a limited degree, food, to local people living within and surrounding the TBR. The predominant uses of wood were for fuel and construction materials. Access to these resources varied, depending on the area or farm where people resided. People living in degraded areas outside of the TBR experienced great difficulty in harvesting wood for fuel or building, and either harvested it illegally off privately-owned land or purchased it at great cost. Generally, it was found that on farms where there were very few families present, residents were allowed greater access to wood compared to those living on farms where many families resided. There were also specialist users living in the area, namely traditional healers and woodcarvers. Limited information was collected on the medicinal use of trees. However, the preliminary data suggests that there is a great need for this resource. It was found that there are very few woodcarvers present in the study area. As the carving industry was the original focus of the study, detailed information was collected from these men. It was found that carved products are largely produced for local markets and included traditional weapons and traditional household implements such as meat trays and spoons. Carvers were finding it increasingly difficult to access wood, and the income they derived from this trade was supplementary. Although it is not perceived possible that the indigenous wood requirements of all local people in the area can be met by the resources within the TBR on an ongoing basis, management of bush encroachment may increase the supply of firewood and construction materials, especially to those farm residents who were experiencing difficulty in this regard at the time of the study. Through partnerships with more specialist users of indigenous trees such as woodcarvers and traditional healers, access to these resources too may be improved. Although more detailed and participative research is needed before substantiated management plans can be formulated, it is hoped that through this study a foundation will be laid to direct future research efforts, dispel misunderstandings, and be part of the effort required to ensure sustainable development of natural resources. / Thesis (M.Sc.)-University of Natal, 1996.

Browse selection, intake, utilisation, palatability, quality and production are tightly linked and need to be considered together in trying to improve our understanding of browsing dynamics and the interactions between browsers and vegetation. Such an understanding is necessary in order to re-evaluate determinations of browser carrying capacities and evaluating actual and potential impacts of browsing animals on vegetation composition and diversity. Browser carrying capacity is determined by both the quantity and the nutritive value of forage. The measurement of browse quantity and nutritive value and the matching of browse supply to browser demand are central to sustainable utilisation and the monitoring of vegetation health. South African savannas are poorly studied with respect to tree canopy growth and browse production making it difficult to quantify the available browse biomass on which browsing capacity estimations are based, and consequently difficult to estimate levels of browsing that are sustainable. This study addressed these issues by investigating browse dynamics, broadly aiming to (1) explore factors affecting browse production, biomass and nutritive value; (2) develop models to assess and monitor these parameters across seasons and properties; (3) use the resultant models in improving our understanding of how to determine browser carrying capacities. More specifically, our study sought to examine the effects of plant physiognomy, forage nutritive value, canopy stratum, defoliation, temperature, rainfall and soil nutrient status on the browse production of evergreen (Carissa bispinosa, Euclea divinorum, Gymnosporia senegalensis), semi-deciduous (Spirostachys africana, Ziziphus mucronata) and deciduous (Acacia nilotica, Dichrostachys cinerea) savanna tree species from June 2003 – June 2005 in three sites along the northern Zululand coastline of KwaZulu-Natal. Available browse biomass, during the dry season, of four key savanna tree species (A. nilotica, E. divinorum, G. senegalensis, and S. africana) was estimated through the development of allometric regression equations. Non-linear regression was used to investigate the relation between the leaf dry mass (LDM) and canopy volume (CVol) of each of the four tree species. Exponential regression (y = a + brlnx) of the natural logarithm of CVol data provided the most accurate and precise description of the tree CVol – LDM relation. A study was undertaken to determine which factors may influence browse production in a southern African savanna. Regression tree models for the browse production identified that the dominant factors influencing browse production were CVol (m3), season, species and height to the lowest leaves of the tree canopy (HL) (m). The length of the growing season had a marked effect on the production potential of savanna tree species, suggesting that improved conditions for growth, i.e. greater rainfall, soil moisture content and improved soil nutrient availability result in a longer period of rapid sustained growth. Species was identified as an important contributing factor to differences in browse production rates, suggesting the need for the development of species or species group models. Mean annual browse production of evergreen trees was greater than that of deciduous and semi-deciduous trees. Mean quarterly (three monthly) browse production was highest, for all trees, during the wet season, with the greatest difference between wet and dry season production being observed in deciduous forms. Evergreen forms showed continuous growth over the whole study, with enhanced growth over the wet season. Deciduous forms, on the other hand, concentrated growth in spurts, when environmental conditions became favourable, with most production occurring during a short growing season. Browse nutritive value was found to be greatest during the wet season, when growth and photosynthesis are at their greatest. Further, browse nutritive value was greatest in deciduous species. Evergreen trees were found to have greater acid detergent fibre (ADF) concentrations than both the deciduous and semi-deciduous trees. By contrast, crude protein (CP) concentrations were greater in semi-deciduous and deciduous species than in evergreen species. The daily CP requirements for maintenance for an adult impala (45 kg) were met by all species over all three study areas and all seasons. Daily CP requirements for growth and lactation, however, were only ever met by deciduous and semi-deciduous species, though this result was not consistent over study areas and seasons. Predictive models for the production of browse on deciduous, semi-deciduous and evergreen trees in northern Zululand were developed using multivariate adaptive regression spline functions. The best predictors of growing season browse production in all three tree guilds (defined here as a group of trees having a characteristic mode of living) were primarily measurable tree dimensions, while the prevailing environmental conditions had little impact. Differences in the production, nutritive value and available browse biomass between the different tree forms and seasons have a profound effect on the determination of browser carrying capacities and need to be incorporated into any game or conservation management plan. / Thesis (Ph.D.)-University of KwaZulu-Natal, Pietermaritzburg, 2012.

The grazing management project in the Okhombe ward of the Amazizi Tribal Authority formed part of the National Department of Agriculture's LandCare program to address communal natural resource management issues. Okhombe land is communal whereby every member of the community is the legal owner of the rangeland with individual ownership of stock with the chieftaincy playing a major role in land allocation. In order to avoid critics of the past and address the top-down approach of the past interventions, a participatory approach was conducted in the planning and implementation of the grazing system. The service providers held a series of visioning workshops with the community in an effort to better understand community resource use patterns, needs, constraints and opportunities as part of the participatory approach. Issues identified by the community were the need for fencing grazing camps, animal health improvement, subdivision of rangeland and crop fields and the development of a rotational grazing system. The main aims of this study were to develop a participatory grazing plan with the community, develop and support institutional structures governing range management, and build capacity of the community in range management. The effect of the current grazing system on species composition was determined. In addition to these, the project investigated the potential different fodder trees has on alleviating feed and nutritional deficit, particularly during the dry winter months of the Upper Thukela. Among the main achievements of this study was the development and strengthening of local institutional structures and effective liaison by all structures with the Inkosi and the tribal council. The community developed a rotational grazing plan, marked the camp boundaries, produced digital maps and successfully built fence boundaries (approximately 20 kms of fencing) to divide their land. The fence boundaries separated the crop fields and rangeland, closed ward boundaries in the upland to prevent access by cattle from neighbouring wards, and divided the land into three camps. Six crush pens were constructed in each subward of the Okhombe ward. A communal herders fund opened and fence construction improved crop yields due to a decrease in crop damage by cattle. Okhombe ward, located in the Highland Sourveld region of KwaZulu-Natal, experiences feed and nutrition deficits to ruminants during winter. The prevailing species composition in Okhombe was investigated as part of the grazing plan. The veld condition of the sites ranged from poor (40.7%) in the bottomland to an averaged of 47.0% in upland sites. The most distinctive feature of the rangeland in this area was the loss of highly palatable Decreaser grass species (P <0.05), such as Themeda triandra in the bottom slopes « 1300 m) when compared to the upland (> 1800 m). The proportional abundance of Decreaser species accounted for an averaged of 1.02% of the bottomland and an averaged of 11.5% of the upland compared to the values of 49% in the benchmark (grassland in optimal condition). The composition of the less palatable Increaser Il species was very high at all elevations (1200 m -80.7%, 1400 m - 75.8% and 1700 m - 55.7%) when compared to the low benchmark composition of 19%. The dominant grasses of the bottom slopes were Increaser Il species, such as Eragrostis curvula, Eragrostis plana and Sporobolus africanus and unpalatable Increaser III species such as Aristida junciformis. A significant difference (P < 0.05) in the composition of Decreaser, Increaser I and Increaser Il species was found between the bottom and slopes compared to the upland region. However, the grass cover formed by these tufted species was generally high, making it more resistant to physical degradation. The bottom slope ranged from reasonable to excellent cover (16.9%), the middle slope ranged from reasonable (15.9%) to a good cover of 18.1%, averaging 16.7% and a range of 16.1% to 17.9% for the upland plateau. In the agroforestry trial the potential of different fodder species for supplementing fodder was examined. Leucaena leucocephala had the potential of being a suitable fodder tree species for use in alley cropping (P < 0.05) compared to Morus alba and Acacia karroo. Results from the partially intercropped treatments showed that L. leucocephala yield (665 kg ha-I) varied significantly (P < 0.05) from the A. karroo (378 kg ha-I) and M alba yield (345 kg ha-I). Treatments that were fully intercropped varied, but no significant difference (P > 0.05) were recorded. Morus alba produced the least yield of 345 kg ha-I, A. karroo yielded 378 kg ha-1 and 1. leucocephala recorded the high of 664 kg ha-I. Results from the second season showed similar trend in that 1. leucocephala yielded a significant (P < 0.05) fodder production of 1715 kg ha-I in comparison to M. alba (1101 kg ha-I) and A karroo (1140 kg ha-I). M alba yielded the least dry matter production (P < 0.05) but had high potential (P < 0.05) for addressing lack of firewood in rural areas. Morus alba yielded high fuel wood production from both two seasons. There were no significant differences in fuel wood yield (P > 0.05) from the partially intercropped M alba (507.9 kg ha-I) and 1. leucocephala (455.0 kg ha-I) but the yield from both species varied significantly from the A. karroo yield (103.kg ha-I). With regard to fully intercropped plots, fuel wood yield from all tree species varied significantly, A karroo resulting in low yield (63 kg ha-I), 1. leucocephala recorded 243 kg ha-l and M alba the highest yield of 444 kg ha-I. In the second season, M. alba yielded an averaged fuel wood production of 728 kg ha-l and a low of 439 kg ha-I from 1. leucocephala. Acacia karroo, a slow growing indigenous tree, might be preferred by farmers due to its less branches resulting in minimal light competition with crops. Leucaena leucocephala tend to grow slowly in its initial establishment stage, but once roots become well established, it grows fast and produces high quantity of fodder. The effect all fodder trees had on crop yield was not negative during the trial period and further research on long term effects of alley cropping is recommended. The conclusions drawn here were based on tree growth and their likely impact in alley cropping. Leucaena leucocephala was also recommended as a preferred species for rural ruminants based on the forage quality study. The results showed high content of crude protein (19.27%), low NDF content (50.38%) and very low tannin content (1.19%) from 1. leucocephala compared to A. karroo with a high tannin content of 5.69%. Acacia karroo had a crude protein content of 13.60%, NDF percentages of 44.16 and 34.64% of ADF content. Morus alba also had a recommended chemical composition of 11.71% of CP, 42.86% of NDF, 36.96% of ADF and a low tannin content of 0.65%. L. leucocephala foliage proven is readily degradable under different diet ranging within 24 hrs of intake (P < 0.001) compared to other feeds. L. leucocephala had high dry matter loss degraded from the rumen under Eragrostis hay diet with poor nutrients to high protein concentrates diet. Under the Eragrostis hay diet for instance, L. leucocephala tend to degrade rapidly with values of dry matter loss ranging from 32.2% to 39% at 4 hrs to 16 hrs, when compared to low dry mater loss of 26% at 4 hrs to 31.33% at 16 hrs. Feeds such as M alba tend to degrade slowly within 24 hrs of intake and rapidly degrades after the stated period. The ep content of maize stover was very low ranging from 1.60% in maize stalks to 2.63% in maize leaves. The fibre content in maize stover was very high when compared to lower values in fodder samples. The NDF content ranged from 77.92% in maize leaves to 81.60% in maize stalks. Maize leaves when compared to a combination of maize leaves and maize stalks sole tend to degrade better within 24 hrs of intake. This was due to low (P < 0.05) degradability rate of maize stalks compared to a combination of maize stalks and leave and leaves sole and least NDF content in maize leaves might have attributed to these results. Due to poor chemical compositions of these roughage samples, the study recommended the establishment of fodder banks and agroforestry systems to curb the nutrients deficit during winter. In conclusion therefore, this study highlight that the sustainability of rural systems to manage communal grazing land should be further explored. Most of the challenging issues in communal range management are social in nature rather than technical concepts. These include ways of improving social contributions from cattle to the community while maintaining cultural values of the use of cattle. The interventions in communal range management by service providers should understand the institutional arrangements within a community and an attempt to strengthen such existing structures is recommended. Further interventions by service providers in Okhombe ward should bring in the planning discussions, experts from social sciences, to deal with understanding of community dynamics. Complexities in communal range management involve dealing with non-stock owners within project boundaries. Communities from neighbouring wards should not be ignored and ways of improving communications and updating project details to them should be formulated. Shortage of land and closing of ward boundaries to prevent access to land by neighbouring wards is among community complexities to be explored. Communities in rural lands do share land and in most cases boundaries are known but invisible by an outsider to identify. It is important to strengthen and maintain every success in communal lands as that may form core of the project. Successes on grazing management by locals is far from being the improvement in veld but there are rather various factors to the successes of grazing projects in rural areas. Examples of successes based on Okhombe project are reduction in stock theft, improved in relationship between community and locals institutions, a reduction in stock mortalities during winter and improved animal health. Veld improvement is among successes but there are accomplishments phases to fulfil before focusing on improvement of species composition. / Thesis (M.Sc.)-University of KwaZulu-Natal, Pietermaritzburg, 2005.