<p> An assemblage of coral skeletons of the species Desmophyllum cristagalli were obtained from the top of Orphan Knoll, 550 km northeast of Newfoundland, from depths of approximately 1600 and 1800 m. The corals were observed for their macro- and micro-boring assemblages, and the boring morphologies documented, using binocular and scanning electron microscopy. Samples of coral were embedded with resin and etched for examination of the micro-boring assemblages.</p> <p> The largest volumetric amount of skeletal material removed was by sponges forming Entobia. This further corroborates the assumption that various species of boring sponges are distributed ubiquitously throughout a wide bathymetric range.</p> <p> Four distinct fungal forms were found, two tubular forms interpreted to be hyphal filaments and two bulbous forms interpreted to be sporangia. These forms were described on the basis of their shape, size, mode of branching, association with other structures and texture. They were then compared to other micro-boring assemblages found in previous studies from various bathymetric ranges. Some forms described in this study were found to be similar to forms described in other studies. Other forms in this study were not found to be documented. It is suggested, therefore, that certain forms, as well as low ichnodiversity, may indicate deeper water environments.</p> <p> Destructive micritization structures were also found in resin casts as well as in thin section. The extent of micritization may indicate the intensity of the parameters at the substrate that affect carbonate dissolution, such as CaCO3 and CO2 concentrations, pH, temperature, and salinity. It would therefore indicate the ambient water conditions at the substrate. If a sufficiently large database could be obtained, as well as distinct separation of the parameters responsible for carbonate dissolution, micritization may be used in a mapping of the carbonate compensation depth through time and depth ranges.</p> / Thesis / Bachelor of Science (BSc)

No description available.

Submitted by Angela Maria de Oliveira (amolivei@uepg.br) on 2018-11-09T12:32:08Z No. of bitstreams: 2 license_rdf: 811 bytes, checksum: e39d27027a6cc9cb039ad269a5db8e34 (MD5) Joao Anesio Bednaz.pdf: 2675052 bytes, checksum: a7697f4fc05d22e6a28c2613712fc43a (MD5) / Made available in DSpace on 2018-11-09T12:32:08Z (GMT). No. of bitstreams: 2 license_rdf: 811 bytes, checksum: e39d27027a6cc9cb039ad269a5db8e34 (MD5) Joao Anesio Bednaz.pdf: 2675052 bytes, checksum: a7697f4fc05d22e6a28c2613712fc43a (MD5) Previous issue date: 2018-08-24 / Os animais desempenham funções no ecossistema, incluindo modificações no ambiente como dispersão de espécies, extinção de outras e alterações na superfície do solo denominadas como bioturbações. Este trabalho objetivou analisar os suínos como agentes zoogeomofológicos influenciadores das formas e processos superficiais em área de pastoreio, dentro do criadouro comum em Sistema de Faxinal. Os faxinais são áreas ocupadas por povos tradicionais cuja formação social é distinguida e caracterizada pelo modo de organização socioespacial, com uso comum da terra e dos recursos florestais, acessível na forma do chamado criadouro comunitário. Esta forma de organização mantém hábitos culturais como: a criação extensiva de animais soltos, com destaque para a criação de suínos. A criação e comportamento dos suínos sem planejamento ou manejo promovem degradações ambientais relacionadas aos recursos hídricos e aos solos. O ato de revolverem grandes quantidades de materiais para exumar raízes e encontrar comida provoca perturbações nos solos modificando o regime hidroerosivo. Neste sentido este trabalho monitorou a conduta dos suínos bem como a dinâmica bioerosiva dos solos do criador comunitário do Faxinal Paraná Anta Gorda, no Município de Prudentópolis/Pr. Os porcos percorreram em média 5.700 metros dentro do criadouro comum transitando por áreas de floresta e pastagens, em busca de alimentos disponíveis neste perímetro. As condições físicas das áreas condicionaram a dinâmica das bioturbações, assim como as taxas de remoção de solos. Nas áreas de maior permanência dos suínos foi encontrada média de erosão de 56 kg/m²/ ano, nos locais utilizados para chafurdamento a média de remoção foi de 130 kg/m²/ano. As áreas com bioturbações pontuais na pastagem tiveram perdas de 10,7 enquanto que a floresta 2,2 kg/m²/ano. Neste sentido os suínos devem ser considerados como uns dos principais agentes bioerosivos em área de faxinais, sendo responsáveis pela degradação do solo. O aumento das áreas de pastagem e do número de suínos, aliados a redução do ambiente com cobertura florestal e do criadouro comunitário podem ampliar os níveis de degradação, resultando na insustentabilidade ambiental deste sistema. / The animals develop functions on the ecosystems, including modifications on the environment, such as dispersion and extinction of species, and modifications on the surface of the soil, known as bioturbations. This work aims to analyze the swine as zoogeomorfological inflator agents on paestore area`s forms and process, inside of a “criadouro comum” from the Faxinal System. The faxinais are areas occupied by the traditional people, in which the social formation is distinguished and characterized by it`s socioespatial organization, gathering common use of the land and the forestall resources. This form of organization maintains cultural habits, such as extensive creation of spread animals, featured by the swine creation. The creation and behavior of the swine without planning or management causes environmental degradation on relation to hydric resources and soil. The act of revolving a large amount of materials to exhume roots and find food provokes disturbances on the soil, which modifies the hydroerosive regime. On this sense, this work monitored the swine conduct as well as the bioerosive dynamic of the soil of a “criadouro comum” of the Faxinal Paraná Anta Gorda in the city of Prudentópolis Pr. The porks ran an average of 5.700 meters inside of the “criadouro comum”, going trough forest and pasture areas on the search for the food available on these areas. The physical conditions of these areas conditioned the dynamic of the bioturbations, as well as the soil removal rates. On the more swine permanence areas, there were encountered rates of erosion of 56 kg/m²/year. On the places utilized for wallow, the rate of removal was of 130 kg/year. The areas with punctual bioturbation on the pasture, had losses of 10.7, while the forest areas had losses of 2.2 kg/ m²/year. On this sense, the swine must be considered as one of the most bioerosive agents on a faxinal area, being responsible for soil degradation. The increase, of the pasture areas and the number of swines, attached to “criadouro comunitário” and covered forestall environmental reduction may enlarge the degradation rates, resulting on the environmental unsustainability of this system.

Es procedeix a l'estudi dels patrons, taxes i formes d'erosió a les costes rocoses carbonatades de Mallorca a partir de la utilització de diferents tècniques instrumentals i experiències de laboratori i camp per tal de desentrellar la contribució dels agents i processos d'alteració i erosió en la formació del relleu litoral. Es dedica una particular atenció a la contribució dels organismes brostejadors (Patella rustica i Melaraphe neritoides) en la destrucció de les costes rocoses, així com també al paper dels biofilms que habitens sobre la roca o bé que ocupen la porositat de la roca i que estan relacionats bé amb les variacions de microtopografia o bé amb el control de la capacitat agressiva de l'aigua retinguda al conjunt de formes del karren litoral. El principal interès del treball realitzat és posar de manifest que a una escala temporal curta, diària, i considerant aquells processos més continus en la dinàmica erosiva de les costes rocoses, la component biològica juga un paper capital en l'erosió del rocam. / This document deals with the study of the rates; patterns and morphologies of erosion at carbonate rock coasts of Mallorca. The approaches takes into account different instrumental techniques and field an laboratory experiments in order to separate the contribution of the different agents and processes in rock coast landform development. The investigation has been specially oriented towards the analysis of the contribution of grazing organisms and the role of biofilms in the evolution of coastal karren forms. The main contribution of the work is to point up that at short time scales and taking in account the processes that operate in a continuous way on the rock coast erosion dynamics, the biological component is very important in the coast erosion.

Carbonate platforms dominated by great thicknesses of ostensibly uniform, thick bedded, poorly fossiliferous and burrow-mottled, subtidal limestones are found throughout the Phanerozoic, yet are poorly understood. Deposited in the aftermath of the FrasnianFamennian extinction event, the Palliser/Wabamun Formation with a thickness of up to 600 m and an extent of about 600 000 km2 represents one of the largest of such platforms to have ever existed.<p>The architecture and palaeoenvironment of the western side of this epeiric platform is reconstructed based on lateral and vertical variations of microfacies properties. It can be divided into inner and outer ramp and shelf belts that remained submerged virtually throughout the life of the platform. A new subdivision of the Palliser Formation is proposed based on long-term vertical facies patterns in sections with differing facies reflecting relative sealevel dynamics. <p>Limestones of the Palliser Formation record a rather limited benthic community. The soft substrate and abundant bioturbation deterred the settlement of sedentary organisms. Frequent physical disturbance, including storms and bioturbation, augmented water turbidity causing smothering and destruction. For these reasons at least, reef growth was impeded. <p>Furthermore, within the apparent facies monotony, two thirds of non-skeletal grains are generated by obliteration processes. Microendolithic bioerosion and diminution destroyed a substantial proportion of the skeletal particles, mostly crinoids, resulting in small micritized grains. Pervasive bioerosion is linked to excess nutrient flux caused by perturbations to the regional biogeochemical cycle. The Ellesmerian Orogeny and increased colonization of land surfaces by deep-rooting gymnosperms are identified as the cause of mesotrophy. Early seafloor dissolution consumed a vast amount of aragonite skeletons, dasycladalean algae in particular, leaving ample amounts of micritic steinkerns, which in turn broke down into peloids and small intraclasts. These discoveries suggest that large areas of the giant ramp-shelf system were characterized by subtidal dasycladalean and crinoid meadows. Finally, established carbonate classifications are re-evaluated and a revised classification for allochthonous limestones for both outcrop and laboratory is introduced. Special emphasis is laid on clastmatrix relationships. <p>-----------------------------------------------<p> ZUSAMMENFASSUNG<br> Die Palliser/Wabamun Formation ist eine von zahlreichen phanerozoischen Karbonatplattformen, die sich durch große Mächtigkeiten vermeintlich monotoner, fossilarmer, bioturbierter Kalke auszeichnen. Diese 600 m mächtige Abfolge sedimentierte im Schatten des Aussterbeereignisses an der Frasne-Famenne Grenze über einen Zeitraum von 69 Ma. Sie gehört mit einer Ausdehnung von etwa 600.000 km2 zu einer der größten Plattformen dieser Art, die auf unserem Planeten entstanden. Mit Hilfe lateraler und vertikaler Trends mikrofazieller Parameter konnte das Ablagerungsmilieu und die Architektur das westlichen Teils dieser epikontinentalen Plattform rekonstruiert werden. Es werden innere und äußere Rampe und Schelf voneinander unterschieden, die praktisch durchgehend über die ganze Plattformentwicklung überflutet blieben. Die vorgestellte Neugliederung der Palliser Formation basiert auf langfristigen vertikalen Faziesmustern in Profilen mit unterschiedlicher Fazies. Faziesmuster spiegeln relative Meeresspiegelschwankungen wider. <p>Die Palliser Kalke sind durch einen eher eingeschränkten Benthos gekennzeichnet, wobei weiches Substrat und intensive Bioturbation die Besiedlung durch sesshafte Organismen erschwerten. Während häufige Bewegung, zum Beispiel durch Stürme und Bioturbation, zu weitreichender mechanischer Zerstörung von Kalkskeletten zur Folge hatte, führte Sedimentaufschlämmung zum Ersticken eines großen Teils des Benthos. Zumindest auf Grund dieser Umstände war das Riffwachstum auf der Palliser Plattform stark reduziert. <p>Etwa zwei Drittel der non-skeletal grains" entstanden durch Klastenalterationsprozesse. Mikroendolithische Bioerosion und Komponentenverkleinerung verwandelten einen großen Teil der Bioklasten, hauptsächlich Krinoiden, zu kleinen mikritischen Körnern. Durchdringende Bioerosion steht in Verbindung mit Nährstoffüberschuss, ausgelöst durch Veränderungen im regionalen biogeochemischen Zyklus. Diese Mesotrophie steht in Zusammenhang mit der Ellesmere Orogenese und zunehmender Besiedlung von Landgebieten durch tiefwurzelnde Gymnospermen. Zusätzlich hinterließ die früh einsetzende Auflösung einer großen Anzahl aragonitischer Kalkskelette, insbesondere Dasycladaleen, am Meeresboden eine beträchtliche Anzahl mikritischer Steinkerne. Diese zerbrachen zu Peloiden und mikritischen Intraklasten. Die Ergebnisse dieser Arbeit legen nahe, dass weite Bereiche des riesigen Palliser Rampen-Schelf Systems durch subtidale Dasycladaleen-Krinoiden-Wiesen charakterisiert waren. Die detaillierte Untersuchung der riesigen Palliser Plattform zeigt, dass etablierte Karbonatklassifikationen nicht ausreichen, um die Fazies treffend zu beschreiben. Die revidierte Version für allochthone Kalke kann sowohl im Aufschluss als auch im Labor angewendet werden und legt besonderes Augenmerk auf Klasten-Matrix-Beziehungen. <br>-----------------------------------------------</br>RÉSUMÉ<br> Les plateformes carbonatées, dominées par d'épaisses couches de calcaires subtidaux prétendument uniformes, en bancs épais, peu fossilifères et creusés de terriers se rencontrent à travers tout le Phanérozoïque. Pourtant, ces plateformes sont encore mal comprises. Déposée à la suite de l'extinction en masse du Frasnien-Famennien, la Formation Palliser/Wabamun, de 600 m d'épaisseur maximale et s'étendant sur environ 600 000 km2, représente une des plus vastes plateformes carbonatées ayant jamais existées. L'architecture et les paléoenvironnements de la marge occidentale de cette plateforme peu profonde ont été reconstruits en se basant sur les variations latérales et verticales des microfaciès. Elle est constituée d'une rampe interne et externe ainsi que d'une plateforme au sens strict, qui sont restées pratiquement ennoyées durant toute la durée de vie de cette plateforme. Basée sur la distribution verticale à long-terme des faciès, avec les changements de faciès reflétant les dynamiques du niveau marin, une nouvelle subdivision de la Formation Palliser est proposée. Les calcaires de la Formation Palliser renferment une communauté benthique plutôt restreinte. Le substrat meuble et le phénomène de bioturbation intense ont empêché la colonisation des organismes sédentaires. Les perturbations physiques et fréquentes comme les tempêtes et la bioturbation ont augmenté la turbidité de l'eau de mer, entraînant asphixie et destruction. Au moins pour ces raisons le développement des récifs a été entravé. De plus, malgré l'apparente monotonie des faciès, les deux tiers des grains "non squelettiques" sont issus de processus de destruction. La bioérosion et diminution par microorganismes endolithiques a détruit une proportion substantielle des particules squelettiques, principalement de crinoïdes, résultant en de petits grains micritiques. Cette bioérosion étendue est liée à un excès de nutriments généré par des perturbations d'ordre géographique du cycle biochimique. L'orogénèse Ellesmere et le développement de la colonisation terrestre par les gymnospermes à racine profonde sont proposés comme cause de cette mésotrophie. La dissolution précoce du sédiment des fonds marins a consummé une importante proportion des squelettes aragonitiques, en particulier des algues dasycladales, laissant de grande quantité de steinkerns micritiques, qui à leur tour ont été décomposés en péloides et petits intraclastes. Ces découvertes suggèrent qu'une large partie de ce système de vaste rampe plateforme était caractérisée par des prés subtidaux à dasycladales et crinoïdes. Enfin; la classification longtemps établie des carbonates a été réévaluée et une classification révisée des calcaires allochtones pour leur étude à l'affleurement et en laboratoire est proposée. Un intérêt partulier est porté sur les relations grains-matrices.

Carbonate platforms dominated by great thicknesses of ostensibly uniform, thick bedded, poorly fossiliferous and burrow-mottled, subtidal limestones are found throughout the Phanerozoic, yet are poorly understood. Deposited in the aftermath of the FrasnianFamennian extinction event, the Palliser/Wabamun Formation with a thickness of up to 600 m and an extent of about 600 000 km2 represents one of the largest of such platforms to have ever existed.<p>The architecture and palaeoenvironment of the western side of this epeiric platform is reconstructed based on lateral and vertical variations of microfacies properties. It can be divided into inner and outer ramp and shelf belts that remained submerged virtually throughout the life of the platform. A new subdivision of the Palliser Formation is proposed based on long-term vertical facies patterns in sections with differing facies reflecting relative sealevel dynamics. <p>Limestones of the Palliser Formation record a rather limited benthic community. The soft substrate and abundant bioturbation deterred the settlement of sedentary organisms. Frequent physical disturbance, including storms and bioturbation, augmented water turbidity causing smothering and destruction. For these reasons at least, reef growth was impeded. <p>Furthermore, within the apparent facies monotony, two thirds of non-skeletal grains are generated by obliteration processes. Microendolithic bioerosion and diminution destroyed a substantial proportion of the skeletal particles, mostly crinoids, resulting in small micritized grains. Pervasive bioerosion is linked to excess nutrient flux caused by perturbations to the regional biogeochemical cycle. The Ellesmerian Orogeny and increased colonization of land surfaces by deep-rooting gymnosperms are identified as the cause of mesotrophy. Early seafloor dissolution consumed a vast amount of aragonite skeletons, dasycladalean algae in particular, leaving ample amounts of micritic steinkerns, which in turn broke down into peloids and small intraclasts. These discoveries suggest that large areas of the giant ramp-shelf system were characterized by subtidal dasycladalean and crinoid meadows. Finally, established carbonate classifications are re-evaluated and a revised classification for allochthonous limestones for both outcrop and laboratory is introduced. Special emphasis is laid on clastmatrix relationships. <p>-----------------------------------------------<p> ZUSAMMENFASSUNG<br> Die Palliser/Wabamun Formation ist eine von zahlreichen phanerozoischen Karbonatplattformen, die sich durch große Mächtigkeiten vermeintlich monotoner, fossilarmer, bioturbierter Kalke auszeichnen. Diese 600 m mächtige Abfolge sedimentierte im Schatten des Aussterbeereignisses an der Frasne-Famenne Grenze über einen Zeitraum von 69 Ma. Sie gehört mit einer Ausdehnung von etwa 600.000 km2 zu einer der größten Plattformen dieser Art, die auf unserem Planeten entstanden. Mit Hilfe lateraler und vertikaler Trends mikrofazieller Parameter konnte das Ablagerungsmilieu und die Architektur das westlichen Teils dieser epikontinentalen Plattform rekonstruiert werden. Es werden innere und äußere Rampe und Schelf voneinander unterschieden, die praktisch durchgehend über die ganze Plattformentwicklung überflutet blieben. Die vorgestellte Neugliederung der Palliser Formation basiert auf langfristigen vertikalen Faziesmustern in Profilen mit unterschiedlicher Fazies. Faziesmuster spiegeln relative Meeresspiegelschwankungen wider. <p>Die Palliser Kalke sind durch einen eher eingeschränkten Benthos gekennzeichnet, wobei weiches Substrat und intensive Bioturbation die Besiedlung durch sesshafte Organismen erschwerten. Während häufige Bewegung, zum Beispiel durch Stürme und Bioturbation, zu weitreichender mechanischer Zerstörung von Kalkskeletten zur Folge hatte, führte Sedimentaufschlämmung zum Ersticken eines großen Teils des Benthos. Zumindest auf Grund dieser Umstände war das Riffwachstum auf der Palliser Plattform stark reduziert. <p>Etwa zwei Drittel der non-skeletal grains" entstanden durch Klastenalterationsprozesse. Mikroendolithische Bioerosion und Komponentenverkleinerung verwandelten einen großen Teil der Bioklasten, hauptsächlich Krinoiden, zu kleinen mikritischen Körnern. Durchdringende Bioerosion steht in Verbindung mit Nährstoffüberschuss, ausgelöst durch Veränderungen im regionalen biogeochemischen Zyklus. Diese Mesotrophie steht in Zusammenhang mit der Ellesmere Orogenese und zunehmender Besiedlung von Landgebieten durch tiefwurzelnde Gymnospermen. Zusätzlich hinterließ die früh einsetzende Auflösung einer großen Anzahl aragonitischer Kalkskelette, insbesondere Dasycladaleen, am Meeresboden eine beträchtliche Anzahl mikritischer Steinkerne. Diese zerbrachen zu Peloiden und mikritischen Intraklasten. Die Ergebnisse dieser Arbeit legen nahe, dass weite Bereiche des riesigen Palliser Rampen-Schelf Systems durch subtidale Dasycladaleen-Krinoiden-Wiesen charakterisiert waren. Die detaillierte Untersuchung der riesigen Palliser Plattform zeigt, dass etablierte Karbonatklassifikationen nicht ausreichen, um die Fazies treffend zu beschreiben. Die revidierte Version für allochthone Kalke kann sowohl im Aufschluss als auch im Labor angewendet werden und legt besonderes Augenmerk auf Klasten-Matrix-Beziehungen. <br>-----------------------------------------------</br>RÉSUMÉ<br> Les plateformes carbonatées, dominées par d'épaisses couches de calcaires subtidaux prétendument uniformes, en bancs épais, peu fossilifères et creusés de terriers se rencontrent à travers tout le Phanérozoïque. Pourtant, ces plateformes sont encore mal comprises. Déposée à la suite de l'extinction en masse du Frasnien-Famennien, la Formation Palliser/Wabamun, de 600 m d'épaisseur maximale et s'étendant sur environ 600 000 km2, représente une des plus vastes plateformes carbonatées ayant jamais existées. L'architecture et les paléoenvironnements de la marge occidentale de cette plateforme peu profonde ont été reconstruits en se basant sur les variations latérales et verticales des microfaciès. Elle est constituée d'une rampe interne et externe ainsi que d'une plateforme au sens strict, qui sont restées pratiquement ennoyées durant toute la durée de vie de cette plateforme. Basée sur la distribution verticale à long-terme des faciès, avec les changements de faciès reflétant les dynamiques du niveau marin, une nouvelle subdivision de la Formation Palliser est proposée. Les calcaires de la Formation Palliser renferment une communauté benthique plutôt restreinte. Le substrat meuble et le phénomène de bioturbation intense ont empêché la colonisation des organismes sédentaires. Les perturbations physiques et fréquentes comme les tempêtes et la bioturbation ont augmenté la turbidité de l'eau de mer, entraînant asphixie et destruction. Au moins pour ces raisons le développement des récifs a été entravé. De plus, malgré l'apparente monotonie des faciès, les deux tiers des grains "non squelettiques" sont issus de processus de destruction. La bioérosion et diminution par microorganismes endolithiques a détruit une proportion substantielle des particules squelettiques, principalement de crinoïdes, résultant en de petits grains micritiques. Cette bioérosion étendue est liée à un excès de nutriments généré par des perturbations d'ordre géographique du cycle biochimique. L'orogénèse Ellesmere et le développement de la colonisation terrestre par les gymnospermes à racine profonde sont proposés comme cause de cette mésotrophie. La dissolution précoce du sédiment des fonds marins a consummé une importante proportion des squelettes aragonitiques, en particulier des algues dasycladales, laissant de grande quantité de steinkerns micritiques, qui à leur tour ont été décomposés en péloides et petits intraclastes. Ces découvertes suggèrent qu'une large partie de ce système de vaste rampe plateforme était caractérisée par des prés subtidaux à dasycladales et crinoïdes. Enfin; la classification longtemps établie des carbonates a été réévaluée et une classification révisée des calcaires allochtones pour leur étude à l'affleurement et en laboratoire est proposée. Un intérêt partulier est porté sur les relations grains-matrices.

The development of coral reefs is largely restricted to areas within the tropics where favourable conditions for both coral and reef growth prevail. There is, however, a continuum from these typical, accretive reefs in the tropics to marginal, non-accretive, coral-dominated reef communities which occur at higher latitudes. High-latitude reefs function similarly in many regards to their tropical counterparts and are regulated by similar processes to a varying degree. In this study, the major biological and physico-chemical processes were assessed which directly or indirectly prevent the continued persistence of reefal frameworks and thus hinder reef accretion on high-latitude reefs in the iSimangaliso Wetland Park. These reefs have a high diversity of hard and soft corals with significant reef coverage, yet little evidence of any biogenic accretion has been observed. The scleractinian coral, Acropora austera, is one of the few corals which may be responsible for reef framework production. It exhibits a gregarious growth pattern, forming large, monospecific stands with an interlocking framework characteristic of the early stages of reef accretion. The framebuilding potential of A. austera and the continued persistence of such frameworks were thus determined by in situ monitoring of coral growth, mortality, bioerosion and several physico-chemical parameters. Growth rate and mortality of A. austera branches were measured at three sites of differing stand size and apparent age. This was achieved by repeated image analysis and by staining branches with the vital stain, Alizarin Red S. Both measures of growth yielded a similar linear extension rate of 24.5 mm/yr (n = 467), comparable to related species at similar latitudes. Mean branch mortality was as high as 50%, with clear differences manifested between each A. austera stand. Branch extension rates and branch mortality were inversely related between sites. Small, young stands exhibited significantly faster coral growth rates, lower mortality and a net increase in overall branch length over the study period, whilst the opposite was true of larger, more developed stands. In addition, bioerosion was determined at each site to assess its potential for carbonate removal and its destabilizing effect on reef frameworks. Bioerosion intensity was recorded as “percentage area damage” within cross-sections and “frequency of occurrence” of bioeroding organisms in coral rubble fragments (n = 120). The level of bioerosion was found to be substantial (up to 11.5% loss in weight of coral fragments over the 12-month study period) and was found to decrease significantly with a reduction in size of each A. austera stand. Aragonite saturation state is considered a major factor that limits the geographical range of coral reefs globally. Although previously thought to be limiting in Maputaland, mean ΩArag values of 4.40±0.29 were measured on the reefs in summer and 4.33±0.21 in winter and thus would not have limited reef development. Past studies have noted the turbulence on South African east coast reefs and its adverse effect on reef development. This was corroborated in this study with the measurement of considerable sediment re-suspension (0.17 g cm⁻² day⁻¹) and regular damage to both living coral and the reef framework caused by large swells. These results lead to the theory that Acropora austera stands senesce with increasing size and age. Although large coral frameworks are found on the Maputaland reefs, they do not persist in the long term. High rates of sediment re-suspension prevent infilling of the interstitial spaces and eventual cementation, while high levels of bioerosion lead to framework instability over time. Rough seas further hamper accretion by physical removal of both living coral and the coral-derived framework, thus removing recent growth. This process is suspected to cause an imbalance in the carbonate budget of these marginal reefs, ultimately favoring carbonate removal over carbonate deposition. / Thesis (M.Sc.)-University of KwaZulu-Natal, Westville, 2011.

Bioerosion is a major process that affects the carbonate balance on coral reefs, and excavating sponges from the genus Cliona are some of the most important bioeroders on Caribbean reefs. The orange boring sponge, Cliona delitrix, is an abundant excavating sponge offshore southeast Florida that frequently colonizes dead portions of live stony corals, killing live coral tissue as it grows. With the recent decline in coral cover attributed to combined environmental and anthropogenic stressors, the increasing abundance of excavating sponges poses yet another threat to the persistence of Caribbean coral reefs. In the first part of this study, I explored distributional patterns of C. delitrix offshore southeast Florida and compared yearly sponge growth/corresponding coral tissue loss rates across habitats of different depths. C. delitrix densities and growth rates were significantly higher on the outer reef, where coral colonies also showed some of the fastest tissue retreat rates. More sponge individuals were found on sites with higher coral densities, likely resulting from the higher availability of preferred coral skeleton substrate. C. delitrix showed a clear preference for boulder stony coral species, which could alter the coral community composition in the future and allow an increase in branching and foliose species. The growth rates of C. delitrix offshore southeast Florida are slower compared to rates from other locations, likely a result of intense fouling of the coral-sponge interface by other spatial reef competitors. These results suggest that outer reef sites with high boulder coral density offshore southeast Florida are most vulnerable to C. delitrix colonization and may continue to suffer the greatest impacts of coral bioerosion. Excavating sponges are also strong competitors for space on coral reefs; able to colonize, excavate, and kill entire live stony corals. Despite the known negative effects of excavating sponges on stony corals very few studies have experimentally tested the competitive nature of this interaction. In the second part of this study, I examined the effect of manual removal of the excavating sponge, Cliona delitrix (Pang 1973), on tissue loss of the stony coral Montastrea cavernosa (Linnaeus 1767), and its possibility as a restoration technique. A total of 33 M. cavernosa colonies colonized by small C. delitrix sponges (up to 10 cm in diameter) were examined. Sponges were removed using a hammer and chisel from 22 of the affected colonies, and 11 colonies were left alone as controls. After sponge removal, the resultant cavities in the coral skeletons were filled to minimize future colonization by other bioeroders and promote coral tissue growth over the excavation. Cement was used as fill material on 11 of the colonies, and the remaining 11 cavities were filled with epoxy. Standardized photos of each colony were taken immediately, at 6 months and 12 months after sponge removal. Results show a significant reduction in coral tissue loss in colonies where sponge was removed, and both fill materials performed similarly reducing coral tissue loss. I also found that a majority of experimental corals showed no return of C. delitrix to the colony surface a year after removal. This study demonstrates that eliminating the bioeroding sponge competitor may promote recovery of the affected stony coral. Additionally, the sponge removal technique can be applied to any stony coral colonized by C. delitrix to preserve, or at least slow the loss of, remaining live tissue.

Sediments of Bohemian Forest lakes are important natural archives. Their sedimentary record covers postglacial history of the lakes as well as history of natural processes in a wider region. It also documents local settlements and changes in landscape management. The lake sediments have attracted the interest of the scientific community since the end of the 19th century. Despite of the fact that modern paleolimnological and paleoecological investigations were already performed in the second half of the 20th century in the Bohemian Forest Mts., the great potential of the lake sediments was not fully utilized in scientific research so far. The ultimate objective of this thesis is to deepen knowledge of these natural archives and support their utilization in future studies. Several specific objectives have been set to achieve the ultimate objective: i) to compare age of the Bohemian Forest lake sediments with the recent knowledge of local deglaciation at the end of the last ice age, ii) to integrate chronostratigraphic marker horizons as one of the tools of Late Glacial sediment dating, iii) to assess the role of bioerosion in chitinous subfossil freshwater invertebrate remains on the record representativeness, and iv) to prove the presumed dystrophic nature of the lakes during the Holocene using...

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