Because of the scarcity of information for bird communities at the major mainland rivers of southeastern Alaska, the main objective of this study was to provide baseline information including distribution, status, and habitat associations of breeding birds. I conducted a meta-analysis of all known reports (including the current study) conducted at major mainland rivers during the breeding season. I described bird species composition, distribution, abundance estimates, status, habitat associations, and guild membership for all birds recorded at 11 major mainland rivers. Based on incidental observations, 170 species were recorded by all studies. Of these, 134 species were known or suspected to breed, accounting for 50% of all birds known from Alaska and 80% of all birds known from southeastern Alaska. In addition, I provided information on species of management concern as well as management implications and recommendations. I used point counts to survey birds within deciduous riparian vegetation at 6 major mainland rivers during 2000-2002. I compared bird species composition, abundance, richness, and diversity among four main vegetation types of deciduous riparian vegetation: shrubland, young deciduous forest, mature deciduous forest, and mixed deciduous-coniferous forest. Species richness was similar among all habitat types; however, relative abundance and diversity of birds was highest in mixed forest stands. Mature forests had the greatest number of species associated with the Canadian interior. I also used point counts to compare bird species composition, abundance, richness, and diversity among 6 major mainland rivers consisting of three trans-mountain and three coastal rivers. Latitude, connectivity, and availability of mature and mixed forests were the major factors thought to cause differences in bird communities among rivers. Contrary to our predictions, coastal rivers had higher bird species richness, diversity, point abundance , and point richness than trans-mountain rivers. Of the 10 species associated with the Canadian interior recorded during point counts, 8 occurred at both trans-mountain and coastal rivers.

There are around 10,426 bird species around the world. Recognizing the bird species for an untrained person is almost impossible either by watching or listening them. In order to identify the bird species from their sounds, there is a need for an application that can detect the bird species from its sound. Time-frequency domain analysis techniques are used to implement the application. We implemented two time-frequency domain feature extraction methods. In feature extraction, a signature matrix which consist of extracted features is created for bird sound signals. A database of signature matrix is created with bird chirps extracted features. We implemented two feature classification methods. They are auto-correlation feature classification method and reference difference feature classification method. An unknown bird chirp is compared with the database to detect the species name. The main aim of the research is to implement the time-frequency domain feature extraction method, create a signature matrix database, implement two feature classification methods and compare them. At last, bird species were identified in the research and the auto-correlation classification method detects the bird species better than the reference difference classification method.

Macroecology is the study of relationships between organisms and the environment at large spatial and temporal scales. This field of research examines patterns in species abundance, distribution and diversity. Understanding patterns in species distribution and richness can contribute significantly to our knowledge of community assembly and macroecological patterns, as well as to the effective conservation of threatened species and habitats. Although there have been a plethora of studies on birds in India over the years, there is a critical need to accurately delineate species distributions and understand patterns of richness. The focus of this study was to understand the factors (abiotic and biotic) that influence the distribution and composition of bird species in the Western Ghats, as well as to explore patterns in their geographic range sizes. The objectives of this study were addressed at the scale of the entire Western Ghats using a combination of field surveys, secondary data collection and species distribution modeling. The specific approaches to address these questions and the findings are outlined below. Chapter 2: Bird species in the Western Ghats – Patterns in composition and richness Fine-scale data on species presence and abundance are essential for exploring patterns in species distribution and richness. Despite the fact that birds have been extensively studied in the Western Ghats, systematic data collection and compilation of information over the entire mountain range has not been carried out, especially for the purpose of testing macroecological questions. This chapter describes patterns in bird species presence, abundance, composition and richness within the Western Ghats. The study area, site selection protocol and the sampling technique have also been described in detail. This dataset establishes a baseline of information about birds in the Western Ghats and subsets of this larger dataset will be used to address various questions in the following chapters. Chapter 3: Predicting bird species distribution in the Western Ghats Detailed knowledge of species’ ecological and geographical distributions is fundamental for conservation, as well as for understanding ecological and evolutionary determinants of spatial patterns of biodiversity. However, occurrence data for a vast majority of species are sparse, resulting in information about species distributions that is inadequate for many purposes. Species distribution models attempt to provide detailed predictions of distributions by relating presence or abundance of species to environmental predictors. In this chapter, we describe the usage of Maxent, a species distribution modelling technique based on presence-only data, to predict the distributions of bird species within the Western Ghats. For this purpose, we put together primary locations of bird species presence along with a published dataset. Using a number of important environmental layers, predicted species distribution maps were derived for 98 bird species, including 13 endemics, in the Western Ghats. Additionally, we calculated predicted range sizes for each of these species and obtained percentage contributions of important environmental predictors to each species’ distribution. This is the first study to develop species distribution models for bird species within the Western Ghats. Chapter 4: Patterns of range size among bird species Understanding large-scale patterns of variation in species geographic range size is fundamental to questions in macroecology and conservation biology. In general, range is believed to be influenced by a combination of environmental factors, evolutionary history and biotic interactions, mediated by species specific traits. These patterns need to be examined even for well-studied taxa like birds, especially within biodiversity hotspots faced by persistent degradation due to anthropogenic activities such as the Western Ghats. In this chapter, we use a dataset of 98 bird species within the Western Ghats to examine trends in range sizes, measured as latitudinal extent of occurrence and predicted range size from species distribution models. We show a significant relationship between latitude and range size for these bird species, supporting Rapoport’s rule. As far as we know, this relationship has never been tested at such low latitudes for birds. We also find that species traits such as body size, mean abundance and diet do not seem to show any discernable effect on patterns of range size. Additionally, we found that widely-used bird species range maps (in this case, from BirdLife International) are inaccurate representations of species ranges in comparison to the predicted species distribution maps that were derived in the previous chapter. We quantitatively demonstrated that these expert-drawn maps need to re-evaluated, especially since they are used to make conservation decisions. This is the first study to quantify species range sizes of birds within the Western Ghats and assess such range maps that are used to determine conservation status of species. Chapter 5: Environmental predictors of bird species distribution One of the major goals in ecology is to understand patterns and processes that determine species diversity. The drivers of global species richness gradients have been studied, especially in the case of birds, in terms of contemporary and historical factors. Such broad scale processes may not always reflect the processes affecting richness and distribution at smaller scales. Therefore, understanding the factors that influence individual species distributions is the first step towards this larger goal. In this chapter, we examined the environmental predictors that contributed to the predicted distribution of bird species observed in the Western Ghats, using the variable importance contribution values derived in Chapter 3. We found that a large proportion of the 98 bird species studied were influenced by normalized differential vegetation index, annual precipitation and elevation. The predictors did not differ among birds of different diet guilds and body size classes. Using Prinicipal components analysis, we observed that all 98 bird species are spread out across the environmental ordination space depicted by the PC axes 1 and 2. These axes are governed by measures of habitat heterogeneity and water-energy related variables, consistent with other tropical studies. The insectivorous guild seemed to occupy a variety of environmental niches across this space and other guilds seemed to be nested within the insectivorous guild. Similarly, larger sized birds were spread across the entire environmental ordination space, with species of smaller sizes nested within. This is the first step in trying to understand environmental predictors acting on birds in the Western Ghats. Further detailed studies need to be carried out to come to definite conclusions. Chapter 6: Relative roles of floristics and vegetation structure on bird species composition On the basis of the hierarchical model of habitat selection, it is known that birds select suitable habitats based on vegetation structure (physiognomy) at coarse biogeographic scales, and plant species composition (floristics) at more local scales. This chapter examines the relative influence of tree species composition and vegetation structure on bird species composition in the Western Ghats. These relationships were specifically assessed across the entire Western Ghats, within regions of the Western Ghats as well as within specific forest types. We found that floristics had a strong association with bird species composition across the Western Ghats and within evergreen and mixed deciduous habitat types. This association seems to be independent of the structural variation in the region. There was a decrease in association strength from the southern to the northern Western Ghats, in terms of both floristics and structure. We did not find an association between vegetation structure and insectivore composition, whereas phytophage composition did show a stronger association with floristics than structure. This is the first study at the scale of the entire Western Ghats to test the relative roles of floristics and vegetation structure. Taken as a whole, this dissertation examines large-scale macroecological questions regarding species distribution, range size and patterns of composition using primary data at the scale of the Western Ghats. The findings of this study have established a foundation that will help further our understanding of species distribution and richness in the Western Ghats, and aid in the decision making for conservation strategies in the future.

Thesis (MSc)--Stellenbosch University, 2015. / ENGLISH ABSTRACT: Forests in South Africa have had a long history of human utilization and disturbance, and are under threat from a variety of anthropogenic land use changes. Foremost of these are deforestation and forest degradation, impacting the species native to these forests. The aims of this study were to determine changes in the distribution of forest dependent bird species according to the South African Bird Atlas Project; to relate these changes to changes in land-use; to identify links between these changes; to determine the extent, location and causes of the decline of each forest dependent bird species; and to identify current risks to forest dependent bird species in South Africa. Range data on 57 forest dependent bird species from SABAP1 (1987-1992) and SABAP2 (2007-present) were analyzed. Of these, 28 species were found to have declining ranges. Thirty sites across South Africa were identified as being most at risk, with all having experienced a loss of more than 10 of the 57 forest dependent bird species between SABAP1 and SABAP2. The range change data of the 28 species with decreasing ranges were correlated with data on changes in land cover over the same time period to infer relationships between changes in land use and change in bird ranges. Occupancy modelling was done to determine which land cover types affect extinction and initial presence. Individual species characteristics were analyzed to determine links between characteristics and response to land use change. A pan-European trait-based risk assessment framework was applied to all 57 species to identify habitats and species most at risk, as well as the most important threats to species persistence. Results showed that natural vegetation decreased in 67% of sites, while plantations and cultivation increased in 50% of sites. Occupancy modelling showed extinction likelihood to increase with plantations in some species, while plantations mitigated extinction likelihood in other species. Urbanization and cultivation likewise mitigated extinction likelihood in some species. Natural vegetation was replaced by cultivation, while cultivation was replaced by urbanization. The number of species lost increased with a loss of natural vegetation. Twenty two of the thirty sites experienced deforestation of indigenous forests between 2000 and 2013/2014; changes in natural vegetation in these sites can be attributed primarily to deforestation, and a loss of plantations. While most at-risk sites were in the Eastern Cape, there was no geographic grouping of species loss or of land use change. Most species lost were birds of prey or insectivores, and species characteristics and habitat preferences determined the sites from which they were lost. The Cape parrot (Poicephalus robustus), rufous-chested sparrowhawk (Accipiter rufiventris) and the migratory Eurasian golden oriole (Oriolus oriolus) suffered the largest declines in range size and are thought to be most at risk. Montane forests were found to be more at risk than other forest types. The major risks facing montane forests were increased abundance of small predators, increased fire suppression, increased soil management, removal of deadwood and reduced diversity of tree species. These threats are all products of plantation forestry and local harvesting. Nesting risk was higher than foraging risk for all species, indicating that nesting habitat should be better preserved. Half of South Africa’s forest dependent bird species have declining ranges, with the loss of these species most prominent in the Eastern Cape. Natural vegetation loss, comprising mostly recent deforestation; increased cultivation and urbanization; and changes in plantation cover are thought to be the main factors determining these declines. Montane forests in particular should be better protected to preserve forest dependent species, and the negative effects of plantation forestry and local harvesting should be mitigated. / AFRIKAANSE OPSOMMING: Suid-Afrika het ‘n lang geskiedenis van woude wat deur mense gebruik, asook vernietig word. Woude word bedreig deur verskeie menslike grondgebruike en veranderinge. Die mees vooraanstaande van die bedreigings is ontbossing en woud verdunning, wat ‘n invloed het op inheemse spesies in die woude. Die doelwitte van hierdie studie was om veranderinge in die verspreiding van bos-afhanklike voëlspesies vas te stel volgens die Suid Afrikaanse Voël Atlas Projek; om hierdie veranderinge te verband met veranderinge in grondgebruik; om verbande tussen hierdie veranderinge te identifiseer; om die mate, die plek en die oorsake van die agteruitgang van elke bos-afhanklike voëlspesies te bepaal; en om huidige risiko's aan die bos afhanklik voëlspesies in Suid-Afrika te identifiseer. Trefwydte inligting oor 57 woud afhanklike voëlspesies van SABAP1 (1987-1992) en SABAP2 (2007-nou) was geanaliseer. Daar was gevind dat vanuit die spesies, het 28 ‘n afneming in trefwydte ervaar. Dertig terreine in Suid-Afrika was uitgeken as dié met die hoogste risiko, met ‘n verlies van meer as 10 van die 57 woud afhanklike voëlspesies tussen SABAP1 en SABAP2. Die trefwydte inligting van die 28 spesies met ‘n afnemende trefwydte was gekorreleer met inligting oor veranderinge in grond bedekking oor dieselfde tydperk om verhoudings tussen veranderinge in grond gebruik en veranderinge in voël trefwydtes aftelei. Besetting modellering was gedoen om te bepaal watter tipes grond bedekking beinvloed uitwissing en aanvanklike teenwoordigheid. Kenmerke van individuele spesies is ontleed om verbande tussen kenmerke en reaksie op verandering in grondgebruik te bepaal. ‘n Pan-Europese eienskap gebaseerde risiko-analise raamwerk is toegepas op die 57 spesies om die spesies en habitatte met die grootse risiko te identifiseer, asook die belangrikste bedreigings vir spesies-volharding. Die resultate het gewys dat natuurlike plantegroei het verminder in 67% van terreine, terwyl plantasie en verbouing vermeerder het in 50% van terreine. Besetting-modellering het gewys dat waarskynlikheid van uitwissing vermeerder met plantasies in sommige spesies, terwyl plantasies die waarskynlikheid van uitwissing verminder het in ander spesies. Verstedeliking en verbouing het ook die waarskynlikheid van uitwissing verminder in sommige spesies. Natuurlike plantegroei was vervang deur verbouing, terwyl verbouing vervang is deur verstedeliking. Die aantal spesies verlies het vermeerder met die vermindering van natuurlike plantegroei. Twee en twintig van die dertig terreine het ontbossing van inheemse woude ervaar tussen 2000 en 2013/2014. Veranderinge in natuurlike plantegroei in die terreine is meestal as gevolg van ontbossing, en ‘n vermindering van plantasies. Terwyl meeste van die hoë risiko terreine in die Oos-Kaap was, was daar geen geografiese groepering van spesies vermindering of grondgebruik veranderinge nie. Meeste spesies wat vernietig is was roofvoëls of insectivore, en spesies kenmerke en habitat voorkeure het die terreine bepaal waaruit hulle verloor is. Die grootpapegaai (Poicephalus robustus), rooiborssperwer (Accipiter rufiventris) en die trekvoël Europese wielewaal (Oriolus oriolus) het die grootste vermindering in trefwydte grootte ervaar, en is vermoedelik die grootste risiko. Bergwoude was gevind om meer in gevaar te wees as ander woud tipes. Die grootste risikos wat bergwoude beïnvloed het was ‘n toenemende hoeveelheid klein roofdiere, verhoogde vuur onderdrukking, verhoogde grondbestuur, verwydering van dooie hout, en verlaagde diversiteit van boomspesies. Hierdie bedreigings is almal as gevolg van plantasie bosbou en plaaslike oes. Nes-risiko was hoër as kos soek risiko vir alle spesies, wat aandui dat nes-habitat beter bewaar moet word. Die helfte van Suid-Afrika se woud-afhanklike-voëlspesies het dalende trefwydtes, met die verlies van hierdie spesies mees opvallend in die Oos-Kaap. Die verlies van natuurlike plantegroei, as gevolg van onlangse ontbossing; verhoogde verbouing en verstedeliking; en veranderinge in plantasie bedekking is vermoedelik die hooffaktore wat die vermindering van voëlspesies veroorsaak. Veral bergwoude moet beter beskerm word om woud-afhanklike-voëlspesies te bewaar en die negatiewe invloede van plantasiebosbou en plaaslike oes te verminder.

Human enlargement and improvements are following the same scheme as the world economies, and have through the same structure damaged and overexploited the ecological systems of the Earth, leading to a progressive degradation of its biodiversity. The need for protected areas is now inescapable when it comes to restoring species while sustainable development and prevention of damaging the environment have become prominent. It is now even more important that the awareness of an actual impact of our living standards on the ecological system has been established. International and more specifically European Laws have been established to prevent and ensure species of all kinds to live in a safe and preserved environment, but the question of their effectiveness has to be asked and nuanced. In that respect the issue of bird species is one of major importance as they are victims of human development. Within the European member states the problem is coming from the inability of some of them to implement and apply European laws because all do not agree on the terms of a protection provided for bird species.

The song thrush (Turdus philomelos) was introduced to New Zealand from Britain during the mid 19th century and has become one of the most common terrestrial bird species in New Zealand. In this study, I surveyed a range of life-history traits in New Zealand song thrushes for comparison with traits of British thrushes. Clutch size, egg size and nest size have decreased, while the nestling period is shorter and the incubation period longer. This combination of changes suggests birds are investing less energy into each reproductive bout. Birds also appear unable to raise large broods, as nestling starvation is common in New Zealand, which suggests that food is limiting. I experimentally tested the ability of song thrushes to incubate enlarged clutches and broods, but productivity was not higher for enlarged broods and natural 3- and 4-egg clutches produced similar numbers of fledglings. Thus reduced clutch size may be an adaptation to the local environment. Differences in female incubation behaviour, with 3- and 4-egg clutches receiving higher levels of incubation and more visits per hour than 5-egg clutches, also suggest New Zealand thrushes have difficulty coping with clutches as large as those in Britain. The decrease in clutch size between New Zealand and Britain is in the direction and magnitude expected based on the change in latitude, which supports the hypothesis that factors affecting foraging time and food availability, such as daylength, temperature and rainfall, may be selecting for smaller clutches. Egg size was also found to have decreased in New Zealand, though this may be the result of smaller adult size. Hatchling mass was related to egg volume, but I found no effect of egg volume or clutch size on hatching success. However, nests containing more pointed eggs (i.e., abnormally-shaped eggs), had lower survival and hatching/fledgling success. Data from the national nest record database and my study both suggest that differences in song thrush productivity are the result of differential survival of nestlings. Nestling mortality due to starvation was common at Kowhai Bush, but rare in Britain, so either adult condition or food availability may be lowering reproductive success in New Zealand. High rates of nest failure (>65%) could also affect clutch size, but the strong directional selection imposed by food limitation during the nestling period suggests that increases in food supply would result in increased reproductive success even with the same levels of nest failure. When comparing clutch size throughout New Zealand, I found a significant, positive relationship with rainfall, which further suggests that food limitation may be the main factor driving changes in life-history traits of song thrushes in New Zealand.

It is generally assumed that when natural habitat is converted to human-dominated cover such area is “lost” to its native species. Extinctions will ensue. The literature generally assumes that species are extirpated as natural area is reduced, following the well-known species-area relationship (SAR). However, SARs have consistently over-estimated species losses resulting from conversion of natural habitat to human-dominated land covers. We hypothesize that the overestimation occurs because these area-based models assume that converted habitat is “lost”, eliminating all species. However, in the real world, conversion of natural land cover to human-dominated cover frequently produces new land covers, different from the original habitat, but not necessarily completely inhospitable to biodiversity. We evaluated the responses of total avian richness, forest bird richness and open habitat bird richness to remaining natural area within 991 quadrats, each 100 km2, across southern Ontario. Total bird species richness does not follow SAR predictions; rather, the number of bird species peaks at roughly 50% natural land cover. The richness of forest birds does follow the usual SAR power-law as a function of forested area. In contrast, richness of birds that prefer open-habitat does not increase monotonically with either natural- or human-dominated land cover. However, we can partition human-dominated land cover into an “available human-dominated” component and “lost” habitat. Richness of open-habitat species relates to the amount of available human-dominated cover. Distinguishing three habitat types (natural, available human-dominated, and lost) permits accurate predictions of species losses in response to natural habitat conversion.

Livestock grazing impacts more land than any other use. Yet knowledge of grazing impacts on native fauna is scarce. This thesis takes a predictive approach to investigating the effects of livestock grazing on Australian woodland birds, employing some novel methodological approaches and experimental designs. These include methods of analysis to handle zero-inflated data and the application of Bayesian statistics to analyse predictions based on expert opinion. The experimental designs have enabled impacts of grazing to be separated from the frequently confounding effects of other disturbances, and to consider the effect of grazing on habitat condition in the context of different surrounding land uses. A distinguishing feature of many datasets is their tendency to contain a large proportion of zero values. It can be difficult to extract ecological relationships from these datasets if we do not consider how these zeros arose and how to model them. Recent developments in modelling zero-inflated data are tested with the aim of making such methods more accessible to mainstream ecology. Through practical examples, we demonstrate how not accounting for zero-inflation can reduce our ability to detect relationships in ecological data and at worst lead to incorrect inference. The impact of grazing on birds was first examined through the elicitation of a priori predictions from 20 Australian ecologists. This expert knowledge was then used to inform a statistical model using Bayesian methods. The addition of expert data through priors in our model strengthened results under at least one grazing level for all but one bird species examined. This study highlights that in fields where there is extensive expert knowledge, yet little published data, the use of expert information as priors for ecological models is a cost effective way of making more confident predictions about the effect of management on biodiversity. A second set of a priori predictions were formulated using a mechanistic approach. Habitat structure is a major determinant of bird species diversity and livestock grazing is one mechanism by which structure is altered. Using available information on the vegetation strata utilised by each species for foraging and the strata most affected by grazing, predictions of the impact of grazing on each bird species were formulated. We found that foraging height preference was a good predictor of species’ susceptibility to grazing. This approach is a starting point for more complex predictive models, and avoids the circularity of post hoc interpretation of impact data. The confounding of grazing with tree clearing was addressed by examining the impact of pastoral management on birds in sub-tropical grassy eucalypt woodland in Southeast Queensland, where land management practices have made it possible to disentangle these effects. Changes in bird species indices were recorded across woodland and riparian habitats with and without trees across three levels of grazing, replicated over space and time. Tree removal had a dramatic influence on 78% of the bird fauna. 65% of species responded significantly to changes in grazing level and the abundance of 42% of species varied significantly with habitat, level of clearing and grazing. The impact of grazing on birds was most severe in riparian habitat. Finally, the extent to which landscape context and local habitat characteristics influence bird assemblages of riparian habitats in grazed landscapes is addressed. Over 80% of bird species responded significantly to changes in local riparian habitat characteristics regardless of context, while over 50% of species were significantly influenced by landscape context. The influence of landscape context increased as the surrounding landuse became more intensive. These results suggest that it is not enough to conserve riparian habitats alone but conservation and restoration plans must consider landscape context. The ability to predict which bird species will be most affected by grazing will facilitate the transformation of this industry into one that is both profitable and ecologically sustainable. Results from this thesis suggest that any level of commercial grazing is detrimental to some woodland birds. Habitats with high levels of grazing support a species-poor bird assemblage dominated by birds that are increasing nationally. However, provided trees are not cleared and landscape context is not intensively used, a rich and abundant bird fauna can coexist with moderate levels of grazing, including iconic woodland birds which are declining elsewhere in Australia.

Fundação de Amparo à Pesquisa do Estado do Rio de Janeiro / Coordenação de Aperfeiçoamento de Pessoal de Nível Superior / A distribuição da biodiversidade está associada aos fatores espaciais, ambientais e biológicos. Esses fatores influenciam a dinâmica das comunidades biológicas, gerando diferenças na distribuição e na abundância de espécies em escalas local e regional, além de criarem variações nos processos populacionais e nos deslocamentos dos animais. Um exemplo é a variação na distribuição e estrutura das comunidades de aves em gradientes altitudinais. Entretanto, não há um consenso sobre o padrão de distribuição da biodiversidade nesses gradientes, sendo reconhecidos quatro padrões de distribuição altitudinal de aves. Nesse contexto, a presente tese teve como objetivo geral estudar algumas das respostas ecológicas das aves à altitude. No primeiro capítulo, avaliamos o conhecimento sobre as migrações altitudinais de aves por meio por meio de uma revisão da literatura científica. Encontramos 84 estudos, a maioria na região Neotropical. Nesses estudos, constatamos 380 espécies de aves que realizam essas migrações, sendo insetívoros e nectarívoros os principais grupos tróficos envolvidos. Esses estudos também mostram que fatores bióticos e abióticos podem interagir para explicar as migrações altitudinais. Os deslocamentos para altitudes mais elevadas podem ser explicados principalmente pela disponibilidade de recursos e o menor risco de predação. Enquanto que os deslocamentos para baixas altitudes podem se relacionar, principalmente, às limitadas oportunidades de forrageamento e à competição. No segundo capítulo analisamos a distribuição regional de beija-flores na Mata Atlântica, por meio do uso de mapas de distribuição de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. Encontramos variações na composição das espécies de beija-flores em relação à altitude, mas, um conjunto de fatores pode explicar essas variações na composição. Nossos resultados mostraram que além da variaçãovariação altitudinal, variação altitudinal, variação altitudinal, variação altitudinal, o efeito do componente espacial (latitude e longitude) e das variáveis ambientais correlacionadas a ele foram importantes na distribuição das aves nas áreas nas áreas estudadas. No terceiro capítulo, estudamos, estudamos a distribuição altitudinal das aves (e de beija-flores) de sub-bosque em cinco altitudes na Reserva Ecológica de Guapiaçu (170 e 370 m) contígua ao Parque Estadual dos Três Picos (570, 770 e 1.000 m), no estado do Rio de Janeiro. Coletamos dados bimestralmente (julho/2010 a junho/2011) e mensalmente (agosto/2011 a julho/2012). Utilizamos o método de captura-marcação-recaptura com dez redes de neblina (12 x 2,5 m, malha de 32 mm) expostas no sub-bosque por sete horas/dia em cada ponto amostral por campanha. Observamos também os beija-flores no sub-bosque, mensalmente, em transecções lineares (400 m de extensão). Capturamos 95 espécies de aves (53% endêmicas de Mata Atlântica), incluindo 10 espécies de beija-flores (oito endêmicos). Detectamos a maior riqueza em 770 m e a menor em 170 m de altitude. Não encontramos relação entre a riqueza das aves e a altitude. Entretanto, encontramos diferenças na composição, riqueza, abundância e na organização trófica das aves nas cinco altitudes amostradas, sendo 170 m, frequentemente, diferente das demais altitudes. Para os beija-flores amostrados com as duas metodologias (captura e observação; 13 espécies), não encontramos diferenças na composição e riqueza nas cinco altitudes. A diversidade e os elevados endemismos registrados na área ressaltam a importância da região para as aves da Mata Atlântica e para preservação dessas no estado do Rio de Janeiro. / Biodiversity distribution is associated to spatial, environmental and biological factors. These factors influence the biological communities dynamics, generating differences in species distribution and abundance, both locally and regionally, and also creating variation in population processes and in animal movements. Variation in the distribution and structure of bird communities in altitudinal gradients is an example, but, there is no consensus about the pattern of altitudinal distribution for birds, and four different patterns are recognized. In this context, the present thesis intended to evaluate the ecological responses of birds to altitude. In the first chapter, we reviewed the literature to evaluate the state of the art about birds altitudinal movements/migrations. We found 84 studies, most conducted in the Neotropics. In these studies, authors reported 380 species of birds performing some kind of altitudinal migration, with insectivores and nectarivores the main trophic guilds involved. The studies also showed that there is a set of interacting biotic and abiotic factors that can explain altitudinal migration. Movement to higher altitudes can be explained mainly by resource availability and low predation risk, while migration to lower altitudes appears to be related to competition and limited foraging opportunities. In the second chapter, we analyzed the regional distribution of hummingbirds in the Brazilian Atlantic Forest, using distribution maps and information available in the literature. We found variation in hummingbird species composition in relation to altitude, but a set of factors could probably explain this variation. Our results showed that in addition to the altitudinal effect, the spatial component (latitude and longitude) related to environmental variables were also important to evaluate the birds distributions. In the third chapter, we investigated the altitudinal distribution of birds (and hummingbirds) in five altitudes in the Ecological Reserve of Guapiaçu (170 and 370 m) contiguous to the Três Picos State Park (570, 770 and 1000 m), Rio de Janeiro state. Data were collected bimonthly (July/2010 to June/2011) and monthly (August/2011 to July/2012), using a capture-marking-recapture method with mist-nets (12 x 2.5 m, 32 mm mesh) opened in the undergrowth for 7 hours/day in each sampling point. We also collected data on hummingbirds monthly using linear transects (400 m long). We captured 95 bird species (53% endemic to the Atlantic Forest), including 10 hummingbird species (8 endemic). The highest species richness was found at the altitude of 770 m and the lowest species richness was found at the altitude of 170 m. We did not find a relationship between altitude and bird richness. However, we found differences in the composition, richness, abundance and in the trophic organization of birds in the five altitudes, with the altitude of 170 m being different from the others in most cases. For hummingbirds sampled with both methods (captured with mist-nets and observed; 13 species), we did not find differences in the composition and in the richness in all five studied altitudes. Diversity and the higher endemism recorded for the area emphasize the importance of the region for the conservation of the Atlantic Forest birds in Rio de Janeiro state.

Fundação de Amparo à Pesquisa do Estado do Rio de Janeiro / Coordenação de Aperfeiçoamento de Pessoal de Nível Superior / A distribuição da biodiversidade está associada aos fatores espaciais, ambientais e biológicos. Esses fatores influenciam a dinâmica das comunidades biológicas, gerando diferenças na distribuição e na abundância de espécies em escalas local e regional, além de criarem variações nos processos populacionais e nos deslocamentos dos animais. Um exemplo é a variação na distribuição e estrutura das comunidades de aves em gradientes altitudinais. Entretanto, não há um consenso sobre o padrão de distribuição da biodiversidade nesses gradientes, sendo reconhecidos quatro padrões de distribuição altitudinal de aves. Nesse contexto, a presente tese teve como objetivo geral estudar algumas das respostas ecológicas das aves à altitude. No primeiro capítulo, avaliamos o conhecimento sobre as migrações altitudinais de aves por meio por meio de uma revisão da literatura científica. Encontramos 84 estudos, a maioria na região Neotropical. Nesses estudos, constatamos 380 espécies de aves que realizam essas migrações, sendo insetívoros e nectarívoros os principais grupos tróficos envolvidos. Esses estudos também mostram que fatores bióticos e abióticos podem interagir para explicar as migrações altitudinais. Os deslocamentos para altitudes mais elevadas podem ser explicados principalmente pela disponibilidade de recursos e o menor risco de predação. Enquanto que os deslocamentos para baixas altitudes podem se relacionar, principalmente, às limitadas oportunidades de forrageamento e à competição. No segundo capítulo analisamos a distribuição regional de beija-flores na Mata Atlântica, por meio do uso de mapas de distribuição de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. de mapas distribuição e informações disponíveis na literatura. Encontramos variações na composição das espécies de beija-flores em relação à altitude, mas, um conjunto de fatores pode explicar essas variações na composição. Nossos resultados mostraram que além da variaçãovariação altitudinal, variação altitudinal, variação altitudinal, variação altitudinal, o efeito do componente espacial (latitude e longitude) e das variáveis ambientais correlacionadas a ele foram importantes na distribuição das aves nas áreas nas áreas estudadas. No terceiro capítulo, estudamos, estudamos a distribuição altitudinal das aves (e de beija-flores) de sub-bosque em cinco altitudes na Reserva Ecológica de Guapiaçu (170 e 370 m) contígua ao Parque Estadual dos Três Picos (570, 770 e 1.000 m), no estado do Rio de Janeiro. Coletamos dados bimestralmente (julho/2010 a junho/2011) e mensalmente (agosto/2011 a julho/2012). Utilizamos o método de captura-marcação-recaptura com dez redes de neblina (12 x 2,5 m, malha de 32 mm) expostas no sub-bosque por sete horas/dia em cada ponto amostral por campanha. Observamos também os beija-flores no sub-bosque, mensalmente, em transecções lineares (400 m de extensão). Capturamos 95 espécies de aves (53% endêmicas de Mata Atlântica), incluindo 10 espécies de beija-flores (oito endêmicos). Detectamos a maior riqueza em 770 m e a menor em 170 m de altitude. Não encontramos relação entre a riqueza das aves e a altitude. Entretanto, encontramos diferenças na composição, riqueza, abundância e na organização trófica das aves nas cinco altitudes amostradas, sendo 170 m, frequentemente, diferente das demais altitudes. Para os beija-flores amostrados com as duas metodologias (captura e observação; 13 espécies), não encontramos diferenças na composição e riqueza nas cinco altitudes. A diversidade e os elevados endemismos registrados na área ressaltam a importância da região para as aves da Mata Atlântica e para preservação dessas no estado do Rio de Janeiro. / Biodiversity distribution is associated to spatial, environmental and biological factors. These factors influence the biological communities dynamics, generating differences in species distribution and abundance, both locally and regionally, and also creating variation in population processes and in animal movements. Variation in the distribution and structure of bird communities in altitudinal gradients is an example, but, there is no consensus about the pattern of altitudinal distribution for birds, and four different patterns are recognized. In this context, the present thesis intended to evaluate the ecological responses of birds to altitude. In the first chapter, we reviewed the literature to evaluate the state of the art about birds altitudinal movements/migrations. We found 84 studies, most conducted in the Neotropics. In these studies, authors reported 380 species of birds performing some kind of altitudinal migration, with insectivores and nectarivores the main trophic guilds involved. The studies also showed that there is a set of interacting biotic and abiotic factors that can explain altitudinal migration. Movement to higher altitudes can be explained mainly by resource availability and low predation risk, while migration to lower altitudes appears to be related to competition and limited foraging opportunities. In the second chapter, we analyzed the regional distribution of hummingbirds in the Brazilian Atlantic Forest, using distribution maps and information available in the literature. We found variation in hummingbird species composition in relation to altitude, but a set of factors could probably explain this variation. Our results showed that in addition to the altitudinal effect, the spatial component (latitude and longitude) related to environmental variables were also important to evaluate the birds distributions. In the third chapter, we investigated the altitudinal distribution of birds (and hummingbirds) in five altitudes in the Ecological Reserve of Guapiaçu (170 and 370 m) contiguous to the Três Picos State Park (570, 770 and 1000 m), Rio de Janeiro state. Data were collected bimonthly (July/2010 to June/2011) and monthly (August/2011 to July/2012), using a capture-marking-recapture method with mist-nets (12 x 2.5 m, 32 mm mesh) opened in the undergrowth for 7 hours/day in each sampling point. We also collected data on hummingbirds monthly using linear transects (400 m long). We captured 95 bird species (53% endemic to the Atlantic Forest), including 10 hummingbird species (8 endemic). The highest species richness was found at the altitude of 770 m and the lowest species richness was found at the altitude of 170 m. We did not find a relationship between altitude and bird richness. However, we found differences in the composition, richness, abundance and in the trophic organization of birds in the five altitudes, with the altitude of 170 m being different from the others in most cases. For hummingbirds sampled with both methods (captured with mist-nets and observed; 13 species), we did not find differences in the composition and in the richness in all five studied altitudes. Diversity and the higher endemism recorded for the area emphasize the importance of the region for the conservation of the Atlantic Forest birds in Rio de Janeiro state.