Faculty of Science School of Animal ,Plant and Enviromental Sciences 9707660j McConnachie@ukzn.ac.za / Characteristics comprising an energy budget of Pseudocordylus melanotus melanotus were investigated in terms of limiting factors for the lizard’s distribution, and included measures of thermal biology, metabolism and digestive physiology. Pseudocordylus m. melanotus is rupicolous, is limited to the temperate parts of southern Africa and exhibits adaptations to these environments. The distribution of the lizard was modelled using two different climate envelope modelling techniques, which are discussed in terms of the energy budget and factors limiting the distribution of the lizard. Pseudocordylus m. melanotus exhibits wide thermal tolerances. It can withstand body temperatures (Tb) below freezing, but freezing of body water, which occurs at ca. -5 ºC, is lethal. The lower critical minimum temperature was a surprisingly high measure for an apparently cold-adapted lizard. In the laboratory, the selected body temperature (Tsel) was ca. 30 ºC. In the wild, lizards thermoregulate by shuttling between hot and cold microclimates, modifying body postures and regulating activity times, and select Tbs of ca. 29 ºC in summer and ca. 26 ºC in winter. Energy expenditure was measured over a range of temperatures. MR increases significantly with increasing temperature. Body mass and metabolic rate (MR) were significantly related at 30 ºC only. The lack of significance at other temperatures can be ascribed to the small range of body mass over which measures were made. Energy gain in P. m. melanotus was quantified in terms of prey capture rates and by measuring aspects of digestive physiology. Prey capture attempts were 35 % successful and occurred ca. every 2 h 30 min in the field. Lizards consumed a variety of arthropods, but mainly ate beetles. Digestive rate and appetite increased significantly with increasing temperature. Apparent digestive efficiency (ca. 94 %) and apparent assimilation efficiency (ca. 87.2 %) were not affected by temperature. iii Two energy budgets were calculated; one based mainly on laboratory measures and the other based on the thermal profile of lizard body temperature measured in the field. The laboratory energy budget was approximately half that of the field energy budget. The field energy budget, however, provided a more realistic view of energy expenditure since it covered almost the entire thermal range experienced by lizards. Between 592.53 kJ (field energy budget) and 940.06 kJ (laboratory energy budget) per year is required for a standard 30 g P. m. melanotus to remain in energy balance. This means that lizards need to consume between 73.55 and 116.69 g of mealworms per year, which equates to ca. 740 to 1200 average mealworms. Any energy gained over and above these requirements can be allocated to growth, reproduction and storage. In terms of the lizard’s distribution, it will be excluded from areas where the available thermal environment and prey abundance do not allow the lizard to maintain a positive energy balance. The lizard is apparently at the cold limit of its range, so predicted climate change in southern Africa is unlikely to have a significant effect on the distribution of the lizard. Suitable crevices are essential to the lizard’s survival, particularly during periods of extreme cold when lizards may experience Tbs where they are incapacitated, or effectively ‘comatose’, while in retreats.

No description available.

No description available.

No description available.

Thesis (Ph. D.)--University of Wisconsin--Madison, 1963. / Typescript. Vita. eContent provider-neutral record in process. Description based on print version record. Includes bibliographical references (leaves 120-124).

Thesis. 1977. M.S.--Massachusetts Institute of Technology. Dept. of Meteorology. / Microfiche copy available in Archives and Science. / Bibliography : leaves 38-39. / M.S.

No description available.

No description available.

No description available.

Soil moisture and ground thaw state are both indicative of a hillslopes ability to transfer water. In cold regions in particular, it is widely known that the wetness of surface soils and depth of ground thaw are important for runoff generation, but the diversity of interactions between surface soil moisture and ground thaw themselves has not been studied. To fill this knowledge gap, detailed shallow soil moisture and thaw depth surveys were conducted along systematic grids at the Baker Creek Basin, Northwest Territories. Multiple hillslopes were studied to determine how the interactions differed along a spectrum of topological, typological and topographic situations (T³ template). Results did not show a simple relationship between soil moisture and ground thaw as was expected. Instead, correlation was a function of wetness such that the correlation between soil moisture and ground thaw improved with site wetness. To understand why differences in soil moisture and ground thaw state arose, water and energy fluxes were examined for these subarctic study sites to discern the key processes controlling the patterns observed. Results showed that the key control in variable soil moisture and frost table interactions among the sites was the presence of surface water. At the peatland and wetland sites, accumulated water in depressions and flow paths maintained soil moisture for a longer duration than at the hummock tops. These wet areas were often locations of deepest thaw depth due to the transfer of latent heat accompanying lateral surface runoff. Although the peatland and wetland sites had large inundation extents, modified Péclet numbers indicated that the relative influence of external and internal hydrological processes at each site were different. Continuous inflow from an upstream lake into the wetland site caused advective and conductive thermal energies to be of equal importance to ground thaw. The absence of continuous surface flow at the peatland and valley sites led to the dominance of conductive thermal energy over advective energy for ground thaw. A quantitative explanation for the shallow soil moisture-ground thaw patterns was provided by linking hydrological processes and hillslope storage capacity with the calculated water and energy fluxes as well as the modified Péclet number. These results suggest that the T³ template and the modified Péclet number could be very useful parameters for differentiating landscape components in modeling soil moisture and frost table heterogeneity in cold regions.