Conserving structural and compositional diversity in Fescue Prairie requires reintroducing natural disturbances according to their historic regime. Fire is an important natural process that may be a source of spatial heterogeneity in Fescue Prairies. The effects of burning in all months of the year except January and February were evaluated in a Fescue Prairie in central Saskatchewan for 6 years following burning on 2 sites that had not been previously burned and 2 sites that had been burned 5 years earlier. Except for burning in March, burning reduced cover of litter (P<0.01) and <i>Festuca hallii </i> (Vasey) Piper (P=0.01) while increasing bare soil (P<0.01) for 1 to 5 years. Cover of <i>Elymus lanceolatus </i>(Scribn. & J.G. Sm.) Gould (P<0.01), graminoids (P=0.02), and species evenness (P=0.01) increased with burning frequency. Burning in late-summer reduced cover of graminoids (P=0.03), plants other than the dominant grasses (P=0.03), and total plant cover (P=0.02). Burning increased the spatial variance (s2) in litter cover (P<0.01) and bare soil (P<0.01) for 1 to 3 years. Aside from burning in early spring, burning reduced s2 in total standing crop (P=0.02) and <i>F. hallii</i> (P=0.01). Variability in the cover of <i>E. lanceolatus </i>(P<0.01) and graminoids (P=0.04) increased with burning frequency. Canonical correspondence analysis (CCA) indicated that pre-burn history had a dominant effect on plant community composition, explaining 13% of the variation (P<0.01). The cumulative effects of repeated burning, annual variability in weather, and exposure to temperature extremes may have caused a shift in the composition of the plant community. The first 4 ordination axes explained 22% of the variation in plant community composition after burning, indicating that many other environmental or site variables controlled community composition. A range of burning dates and frequencies should be reintroduced or maintained in Fescue Prairie to create a mosaic of plant communities in various stages of recovery after burning. A mosaic will increase the structural and compositional diversity in remnant Fescue Prairies.

Conserving structural and compositional diversity in Fescue Prairie requires reintroducing natural disturbances according to their historic regime. Fire is an important natural process that may be a source of spatial heterogeneity in Fescue Prairies. The effects of burning in all months of the year except January and February were evaluated in a Fescue Prairie in central Saskatchewan for 6 years following burning on 2 sites that had not been previously burned and 2 sites that had been burned 5 years earlier. Except for burning in March, burning reduced cover of litter (P<0.01) and <i>Festuca hallii </i> (Vasey) Piper (P=0.01) while increasing bare soil (P<0.01) for 1 to 5 years. Cover of <i>Elymus lanceolatus </i>(Scribn. & J.G. Sm.) Gould (P<0.01), graminoids (P=0.02), and species evenness (P=0.01) increased with burning frequency. Burning in late-summer reduced cover of graminoids (P=0.03), plants other than the dominant grasses (P=0.03), and total plant cover (P=0.02). Burning increased the spatial variance (s2) in litter cover (P<0.01) and bare soil (P<0.01) for 1 to 3 years. Aside from burning in early spring, burning reduced s2 in total standing crop (P=0.02) and <i>F. hallii</i> (P=0.01). Variability in the cover of <i>E. lanceolatus </i>(P<0.01) and graminoids (P=0.04) increased with burning frequency. Canonical correspondence analysis (CCA) indicated that pre-burn history had a dominant effect on plant community composition, explaining 13% of the variation (P<0.01). The cumulative effects of repeated burning, annual variability in weather, and exposure to temperature extremes may have caused a shift in the composition of the plant community. The first 4 ordination axes explained 22% of the variation in plant community composition after burning, indicating that many other environmental or site variables controlled community composition. A range of burning dates and frequencies should be reintroduced or maintained in Fescue Prairie to create a mosaic of plant communities in various stages of recovery after burning. A mosaic will increase the structural and compositional diversity in remnant Fescue Prairies.

Diss. (sammanfattning). Uppsala : Sveriges lantbruksuniv. / Härtill 5 uppsatser.

Le changement climatique entraîne un changement profond des régimes thermiques qui a des conséquences importantes sur la croissance des plantes. Toutefois, les effets des températures extrêmes ont été peu étudiés chez espèces fourragères pérennes, et ne sont pas intégrés dans les modèles actuels de prairie. Or ces modèles ont un rôle central à jouer dans le développement de stratégies d'adaptation au changement climatique. Dans ce contexte, cette thèse a cherché à approfondir la compréhension de la réponse de la luzerne et de la fétuque élevée à une large gamme de températures et à améliorer la prise en compte de ces réponses dans le modèle STICS. Un volet de travail expérimental en conditions contrôlées n'a pas permis de mettre en évidence de différences de réponses entre génotypes d'origine contrastées (tempéré/Méditerranéen) pour différents processus de croissance et de développement des deux espèces. En revanche, il est apparu des différences significatives entre les réponses normalisées de différents processus ; l'allongement des tiges en particulier se distinguant nettement des autres processus chez la luzerne. La réponse de la photosynthèse à la température a montré une forte acclimatation à la température de croissance, aussi bien pour des génotypes d'origine tempérée que méditerranéenne. Ces derniers sont apparus plus sensibles aux températures élevées du fait d'une désactivation de la RUBISCO. La modification des formalismes de réponse à la température de la croissance et du développement dans STICS (fonction non linéaire, pas de temps) a eu un fort impact sur le cumul de temps thermique pour des scénarios climatiques « futurs ». Toutefois, du fait de l'effet majoritaire d'autres stress conjoints durant les périodes de stress thermique et d'une rapide saturation du rayonnement intercepté, l'impact sur les projections de production fourragère s'est avéré limité pour les conditions testées. Au total ces résultats suggèrent que les productions fourragères pourraient être moins exposées au stress thermique que les cultures annuelles. / Due to climate change, the global average temperature is expected to rise with marked impacts on crop growth. However, extreme temperature's impacts on perennial herbaceous species have received much less attention to date than other crops and aren’t currently implemented in grassland models. Yet, grassland models have a major role to play in defining adaptation options to global warming’s impacts on agriculture. In this context, our aims were to characterise the responses to temperature of two major perennial forage species over a large range of growth temperature and to improve the temperature response formalisms in the STICS model. First, experiment conducted in growth chambers indicated that the responses of normalized developmental rates did not differ between genotypes from contrasting thermal areas (Mediterranean/temperate) within each species. On the other hand, the responses to temperature of normalised rates differed significantly between several of the physiological processes studied; in particular stem elongation rate differed from other developmental rates in alfalfa. Irrespective of cultivar origin, a remarkable acclimation of photosynthesis to growth temperature was observed. Mediterranean genotypes displayed a greater sensitivity of the maximum rate of Rubisco carboxylation to elevated temperatures. Modifications of temperature formalism in the STICS crop model (implementation of nonlinear response, hourly/daily step of response) had a significant impact on thermal time accumulation and crop development for future climate scenarios. However, due to larger effects of water stress and to saturation of radiation interception, impacts on projections of grassland dry matter production were limited. This result suggests that grassland production could be less exposed to thermal stress than annual crops.

Abstract The main objective of this thesis was to investigate tree - cover crop interactions and soil response in an intercropping system, in which leguminous and grass cover crops were used with birch (red-leaved Downy birch Betula pubescens L.). The investigation was conducted in two field experiments in a nursery and in a greenhouse experiment. In the latter the effect of the water supply in interspesific competition was also investigated. The cover crops in the field experiments were perennial clovers Trifolium pratense L., T. repens L. and T. hybridum L. and annual clovers T. incarnatum L., T. resupinatum L. and T. subterraneum L. and perennial Festuca rubra L. The height, stem diameter, leaf area and nutrient status of the birch were determined, as well as soil nutrient status and microbial characteristics. The cover crops in the greenhouse experiment were T. repens, T. subterraeum and F. rubra. The biomass, height, leaf area, leaf area index, specific leaf area and N concentration of the birch, the biomass and N concentration of the cover crops were measured, and soil N and microbial characteristics, as well. Bare ground was the control in all the experiments. The perennial clovers and Festuca strongly decreased the birch growth and nutrient status, but the annual clovers sown in midsummer in the field experiments provided about as good growth as bare ground. In the greenhouse experiment all cover crops were effective competitors with the birch. The soil NO3-N was, in general, the highest on bare ground and second highest in the annual clover plots. Though there were, in general, only minor differences in the soil nutrient concentrations between the treatments, there were significant differences in the tree growth and nutrient concentrations. The interspecific competition in this kind of intercropping system is mainly belowground. The growth reduction in the birch was mainly due to competition for nitrogen but water seems to play an important role in regulating the competitive interaction between the birch and cover crops. The competition for these resources seems to be most crucial at the beginning of the growing season. The microbial biomasses and soil respiration were greatest in the Festuca and perennial clover treatments, which may indicate that microorganisms together with these cover crops may seriously compete with birch for nutrients. Intercropping system is complex and comprises both negative and positive influences. In order to minimize negative competition effects, the cover crop should be non-competitive or the ground should be kept free of vegetation at the beginning of the growing season. By improving soil microbial characteristics, the vegetative ground covers make this cropping system one possiblity towards sustainability in the long-term.

Previous field research show, that late successional species correspond with soil development, especially with organomineral A horizon developing. Aim of my thesis was to compare plant growth on different types of soil from spoil heaps. They were taken from three chronosequencies of different age - nonrecultivated and soils recultivated with planting of alder (Alnus) and spruce (Picea). In my research were used 7 types of plants - Arrhenatherum elatius, Centaureae jacea, Festuca rubra, Lychnis flos-cuculi, Lotus corniculatus, Plantago major, Trifolium medium. Spontaneous sites support more species than reclaimed ones this difference increase with plot age without distinct difference between early and late succession species. Key words Succession, recultivation, germination alder (Alnus), spruce (Picea), Arrhenatherum, Centaureae, Festuca, Lychnis, Lotus, Plantago, Trifolium.

Soil contamination with heavy metals is a problem of worldwide concern that is still unsolved. The analysis of the current methods of soil cleaning from heavy metals leads to conclusions that phytoremediation, i.e. soil decontamination by using plants, is one of the best methods. Although this method has not received a wide application yet and possibilities of its application are still being analysed, it is one of the most prospective soil cleaning methods due to low cost and a rather efficient cleaning of the upper surface of soil. Three kinds of Poaceae f. Species – Lolium perenne L., Poa pratensis L. and Festuca pratensis Huds. – have been chosen in this work for decontaminating soil from heavy metals. These plants were grown under artificial laboratory conditions in soil which was once and periodically contaminated with heavy metals. It was established that it is the Lolium perenne L. that most efficiently removes heavy metals and cleans soil. The results of experiments show that the selected grassy plants (perennial ryegrass, meadow-grass and fescue-grass) efficiently clean soil from heavy metals (copper, lead, manganese, zinc, nickel and chromium) when soil contains both low (0.5–120 mg/kg) and high (up to 6,850 mg/kg) concentrations of these metals. Since the perennial ryegrass absorbs the highest amount of heavy metals, the highest efficiency of soil cleaning is achieved using this plant, therefore, 80 % of the mixture of grassy plants was formed of the perennial... [to full text] / Dirvožemio tarša sunkiaisiais metalais visame pasaulyje aktuali problema iki šiol neturinti universalaus sprendimo. Išanalizavus šiuo metu dirvožemio valymui nuo sunkiųjų metalų naudojamus metodus, galima daryti išvadą, kad vienas iš tinkamiausių metodų yra fitoremediacija, t. y. dirvožemio valymas nuo teršalų panaudojant augalus. Šis metodas dar nėra plačiai taikomas ir jo taikymo galimybės dar tik tiriamos tačiau, tai vienas iš perspektyviausių dirvožemio valymo metodų dėl sąlyginai mažos kainos ir pakankamai efektyvaus paviršinio dirvožemio sluoksnio išvalymo. Atliktų eksperimentinių tyrimų rezultatai parodė, kad pasirinkta žolinė augalija – daugiametė svidrė, pievinė miglė ir tikrasis eraičinas efektyviai valo dirvožemį nuo sunkiųjų metalų (vario, švino, mangano, cinko, nikelio, chromo) tiek esant mažoms (0,5–120 mg/kg), tiek didelėms (iki 6850 mg/kg) jų koncentracijoms dirvožemyje. Kadangi daugiausiai sunkiųjų metalų iš dirvožemio sorbuoja svidrė, tai dirvožemio valymo efektyvumas naudojant šią žolinės augalijos rūšį yra efektyviausias ir sudarant žolinės augalijos mišinį 80  sudarė svidrė ir po 10  eraičinas bei miglė. Eksperimentų metu tirta, kaip sunkiųjų metalų mišinio koncentracijos kenkia hidrobiontams ir nustatyta, kad didinat sunkiųjų metalų mišinio koncentracijas (iki 0,198–5,12 g/l) jos tampa letalinėmis vaivorykštiniam upėtakiui, tuo pačiu ir kitiems vandens gyvūnams.Pagal tyrimų su vaivorykštiniais upėtakiais rezultatus nustatyta, kad stebimų... [toliau žr. visą tekstą]

Dirvožemio tarša sunkiaisiais metalais visame pasaulyje aktuali problema iki šiol neturinti universalaus sprendimo. Fitoremediacija – naujai atsirandanti ir daug žadanti technologija nestipriai užterštiems dirvožemiams valyti naudojant augalus. Šiame darbe dirvožemio valymui nuo sunkiųjų metalų pasirinktos trys žolinės augalijos rūšys – svidrė, miglė ir eraičinas, augintos modelinėmis laboratorinėmis sąlygomis vieną kartą ir periodiškai sunkiaisiais metalais teršiamuose dirvožemiuose. Nustatyta, kad geriausiai sunkiuosius metalus iš dirvožemio sorbuoja ir didžiausias dirvožemio valymo efektyvumas pasiekiamas naudojant svidrę. Augalijai sorbuojant sunkiuosius metalus iš dirvožemio dalis šių metalų (Cu, Cr, Zn, Pb, Mn, Ni) koncentruojasi žolinės augalijos šaknyse, tačiau didžioji dalis ten susikaupiančių metalų (iki 90 proc.) pereina į antžeminę augalo dalį. Didžiausios sunkiųjų metalų koncentracijos augale (iki 60 proc. ir daugiau) ir atitinkamai efektyviausias dirvožemio valymas nustatytas svidrės atveju, todėl sudarant mišinį didžiąją dalį (80 proc.) jame sudaro svidrė. Sunkiųjų metalų mišinys daro neigiamą įtaką vaivorykštinio upėtakio jauniklių fiziologiniams ir morfologiniams rodikliams, todėl didinat sunkiųjų metalų mišinio koncentracijas (iki 0,198–5,12 g/l) jos tampa letalinėmis vaivorykštiniams upėtakiams, tuo pačiu ir kitiems vandens gyvūnams. Pagal tyrimų su vaivorykštiniais upėtakiais rezultatus nustatyta, kad stebimų upėtakių, gyvenusių akvariume, užterštame 5,7... [toliau žr. visą tekstą] / Soil contamination with heavy metals is a problem of worldwide concern that is still unsolved. The analysis of the current methods of soil cleaning from heavy metals leads to conclusions that phytoremediation, i.e. soil decontamination by using plants, is one of the best methods. Although this method has not received a wide application yet and possibilities of its application are still being analysed, it is one of the most prospective soil cleaning methods due to low cost and a rather efficient cleaning of the upper surface of soil. Three kinds of Poaceae f. Species – Lolium perenne L., Poa pratensis L. and Festuca pratensis Huds. – have been chosen in this work for decontaminating soil from heavy metals. These plants were grown under artificial laboratory conditions in soil which was once and periodically contaminated with heavy metals. It was established that it is the Lolium perenne L. that most efficiently removes heavy metals and cleans soil. The results of experiments show that the selected grassy plants (perennial ryegrass, meadow-grass and fescue-grass) efficiently clean soil from heavy metals (copper, lead, manganese, zinc, nickel and chromium) when soil contains both low (0.5–120 mg/kg) and high (up to 6,850 mg/kg) concentrations of these metals. Since the perennial ryegrass absorbs the highest amount of heavy metals, the highest efficiency of soil cleaning is achieved using this plant, therefore, 80 % of the mixture of grassy plants was formed of the perennial... [to full text]

La germination et la croissance hétérotrophe sont des phases clés de l'établissement des plantes. Ils sont sous contrôle génétique et affectés par la température. L'objectif de cette thèse est d'analyser la variabilité inter et intra-spécifique de cinq espèces prairiales dans leurs réponses à la température pendant la germination et la croissance hétérotrophe. Trente-deux populations de Lolium perenne, Festuca arundinacea, Dactylis glomerata, Medicago sativa, et Onobrychis viciifolia ont été évalués sous huit températures constantes entre 5 et 40°C.La nouveauté de ce travail vient de la large gamme de températures et le nombre de populations évaluées. Il a été observé que, au sein des espèces, la réaction des populations à la température indique une forte variabilité et des différences significatives. La germination n'a été pas observée à 40°C pour l'une des 23 graminées, l'O. viciifolia germé peu tandis que les variétés de M. sativa ont été peu affectées. La croissance hétérotrophe des 32 populations a été affectée par la température. La croissance des axes fortement fiable ou nulle à 40°C. Les courbes de réponse du taux de croissance maximale de l'axe sont asymétriques en forme de cloche.Des interactions significatives entre la température et la population ont été détectées pour la germination et la croissance hétérotrophe. La germination des populations des sites froids est renforcée par des températures plus chaudes et limité par les plus froids et vice-versa pour les populations chaudes adaptées. / Germination and heterotrophic growth are key phases for plant establishment. They are under genetic control and affected by temperature. The objective of this thesis was to analyse the inter- and intra-specific variability of five pasture species in their responses to temperature during germination and heterotrophic growth.Thirty two populations of Lolium perenne, Festuca arundinacea, Dactylis glomerata, Medicago sativa, and Onobrychis viciifolia were evaluated under eight constant temperatures between 5 and 40°C.The novelty of this work comes from the wide range of temperatures and the number of populations evaluated. It was observed that, within species, the response of populations to temperature shows high variability and significant differences. No germination was observed at 40°C for any of the 23 grasses, O. viciifolia germinated poorly while M. sativa varieties were little affected. Heterotrophic growth of the 32 populations was affected by temperature. The growths of the axes were negligible or lethal at 40°C. Response curves of maximum growth rate of the axis were asymmetric bell-shaped.Significant interactions between temperature and population were detected for germination and heterotrophic growth. Germination of populations from cold sites is enhanced by warmer temperatures and limited by colder ones and vice versa for warm-adapted populations. This study demonstrates that genetic variability does exist within the five studied species. This variability could be exploited to breed new varieties adapted to the future climate.

Disputats. Rheinische Friedrick-Wilhelms-Universität, 2001. / Haves kun i elektronisk udg.