The genus Castor is represented in Eurasia by Castor fiber, North America by C. canadensis, and has been in North America since the late Miocene. This study aims to assess whether morphology of Miocene-Pliocene C. californicus and extant C. canadensis are distinctly different. Specimens of Castor were compared using geometric morphometrics on cranial material and linear measurements of postcranial material. Species occurrence data were compared with past and future climate data to assess Castor distribution in North America through time. Results show that C. canadensis is highly variable in both cranial and postcranial morphology and C. californicus falls largely within the range of variation seen within the extant species. Past distributions match fossil occurrences of Castor, suggesting confidence in projected models. Morphological and distribution similarities between the two species suggest that they can be treated as ecological analogs, though evaluation of whether they are conspecific will require more data.

During the latter half of the 20th century until present day there has been an unprecedented rise in global annual mean temperatures accompanied by rising sea levels and a decrease in Northern Hemisphere snow cover, which if it continues will lead to widespread disruption of climate patterns, ecosystems, and present-day landscapes. It is therefore of critical importance to establish an expanded network of paleoclimate records across the globe in order to better assesses how the global climate system has changed in the past, that we may create a metric by which to address modern change. Herein is presented a7,700 years record of Holocene climatic and environmental variability in Sermilik Valley, located on Ammassalik Island, SE Greenland. This objective of this study is to determine the timing of major Holocene climate transitions as expressed in the physical, elemental, and geochemical parameters preserved in the 484 cm sediment record of Lower Sermilik Lake. Major transitions observed in this study include the deglaciation of Sermilik Valley, the onset and termination of the Holocene Climatic Optimum, the transition into neoglacial conditions, and the Little Ice Age.

The chalicothere Moropus was a rare perissodactyl present in the Great Plains region of North America through much of the Miocene. A temporal gap in named species of Moropus is present in the early Hemingfordian North American Land Mammal Age. This gap is filled by specimens currently referred to as Moropus sp. from the Batesland Formation in southwest South Dakota, and unnamed specimens of Moropus in the Runningwater Formation in northwestern Nebraska. A comparison of the fossils of Moropus nsp. from the Batesland Formation with those of previously described chalicothere species from the Greats Plains region, such as Moropus elatus, Moropus hollandi, Tylocephalonyx skinneri, Moropus merriami, and Moropus matthewi, in addition to Metaschizotherium bavaricum and Metaschizotherium fraasi from the Miocene of Southern Germany, illustrates that there are substantial differences in morphology between equivalent skeletal elements. Based on these findings, I propose that the specimens of Moropus from the Batesland Formation belong to a new species.

Three new types of arthropod from Cambrian intertidal lithofacies of the Elk Mound Group and Lodi Member of Wisconsin, and the Potsdam Group of Quebec are described. These arthropods are preserved ventrally in three dimensions – allowing detailed characterization of morphology. Arenocaris inflata, from the Furongian Elk Mound Group and St. Lawrence Formation, is the earliest occurrence of a phyllocarid. Mosineia macnaughtoni, a large (>10 cm long) euthycarcinoid arthropod, also occurs in Elk Mound strata. Mictomerus melochevillensis represents a new family of early euthycarcinoids, and is a large (8–10+ cm long) arthropod with eleven pairs of homopodous, uniramous limbs. Phylogenetic analyses and reviews of Paleozoic phyllocarid systematics are presented, using morphology-based characters from Cambrian to Recent taxa. Resulting cladograms place Arenocaris inflata into a systematic context, and reveal that the families Ceratiocarididae and Caryocarididae, as traditionally defined, are paraphyletic. Caryocarididae is elevated to subordinal rank (Caryocaridina n. suborder), resulting in two monophyletic suborders. Emended diagnoses are integrated into this analysis, and result in synonymy of 30 Cambrian – Silurian caryocaridids and ceratiocaridids into 11 pre-existing species. The phyllocarid Arenocaris inflata from the Elk Mound Group of Wisconsin and the large enigmatic arthropod Mictomerus melochevillensis from The Potsdam Group of Quebec are both directly associated with trace fossils. Direct association between these arthropods and their traces allows functional morphological details of the animal to be assessed, provides a framework for understanding how arthropods can be sand-cast in three-dimensions, and helps provide insight about subaerially-produced traces from the Potsdam Group.

Compound specific isotope analysis of lake sediments is a powerful tool in deciphering evidence of changing climatic and paleoenvironmental conditions through time. Isotopic analysis of Lake El’gygytgyn pilot sediment cores, PG1351 and LZ1029, have contributed increased insight into paleoenvironmental interpretations regarding conditions of permanent ice cover and water column anoxia at the lake over the past 250 kyr. Bulk sediment δ15N was measured as a proxy for denitrification and a possible indicator for water column anoxia intensity. However, it appears that insufficient quantities of water column nitrate to fuel denitrification make its correlation with anoxia intensity ineffective. In pilot core LZ1029, compound-specific δ13C of alkanes, fatty acids, and alcohols were analyzed to determine the changing sources of organic matter as well as the source of a strong negative isotopic shift in the bulk sediment δ13C (-26‰ to -33‰) over the past 50 kyr. Results indicate that the majority of alkanes, fatty acids, and alcohols are long chain compounds consistent with a terrestrial plant origin, with increased aquatic contribution during the local last glacial maximum (LLGM). Among the compound classes examined, only the mid chain fatty acids display a strong LLGM depletion (δ13C = -43‰). Short chain fatty acids exhibit an LLGM depletion (δ13C = -35‰) similar to bulk sediment δ13C, while the δ13C trend of long chain alkanes, fatty acids, and alcohols differ from the bulk sediment δ13C trend, suggesting an autochthonous source of bulk isotope depletion. Evidence of methane cycling exists only in the presence and isotopic value of diplopterol (LLGM δ13C = -93.4‰), a biomarker for aerobic oxidation of methane. Two compounds indicative of archaeal lipids were present at considerable concentrations during the LLGM (394 and 668 µg/g TOC), but without the extreme negative δ13C associated with methanogenesis and methanotrophy. These results suggest insufficient generation of methane in the lake to have derived from such a large anaerobic archaeal methanogen community suggesting that archaea are not acting entirely as methanogens. Furthermore, it appears unlikely that a significant anoxic layer existed in the water column of Lake El’gygytgyn during the past ~50kyr. The results of this work will be applied to ongoing investigations on the newest cores from Lake El’gygytgyn, which represent the past 3.5 Myr.

Individuals of Tapirus polkensis from the Gray Fossil Site exhibit an excellent level of preservation. Intact skulls collected from the site were arranged in a rough age class system separated into 7 categories based on the teeth present and amount of dental wear. Such an eruption series is useful for general comparisons amongst the individual tapirs; however, the classes do not represent an age in years due to a lack of data on living tapirs. Consequently, it is possible that some age classes may contain several years of a tapirs life, or comparatively only a few months. In this study I placed ages on individuals of T. polkensis from the Gray Fossil Site based on age data taken from The Baird's Tapir Project of Costa Rica (Baird's Tapirs, T. bairdii) ranging from several months to 7 years in age. As eruption data is only useful for aging tapirs up to 7 years, this study also took dental wear into account for adult tapirs. After aging all possible individuals in the sample, the sample was compared to other studies on perissodactyl population structure. Interestingly, the T. polkensis sample is remarkably similar to modern populations of Diceros bicornis.

The analysis of samples recovered from the abdominal area of an articulated tapir from the Late Miocene (4.5-7 million bp) Gray Fossil Site, revealed a rich palyno-fauna comprised of about 94% egg/oocyst-like structures and 6% pollen and other palynomorphs. In addition, a tight grouping of six hickory nuts (Carya) was recovered from the same area suggesting that the samples represent the abdominal contents. The analysis of a sample from immediately outside the tapir produced a sample with 98% pollen and less than 0.5% egg-like structures. The size, shape, and general morphology of egg-like structures were analyzed with light and scanning electron microscopy and were compared to a variety of intestinal eggs found in extant ungulates, and in particular the Perissodactyla. We also compared the fossil structures to the numbers and kind of intestinal parasites recovered from fecal samples from the Baird's tapir (Tapirus bairdii) in Costa Rica and from samples collected from the Lowland tapir (T. terrestris) from Ecuador to assess their similarity to our fossil sample. Based on these data we discuss what role parasites may have played in the biology of T. polkensis during the late Miocene-early Pliocene.

The preservation and quantity of fossil tapirs (Tapirus polkensis) from the Gray Fossil Site in northeastern Tennessee provides a unique opportunity for comparison and analysis of skeletal characters. Intraspecifically, modern tapirs show little to no variation in the morphology of the sagittal crest. However, several different morphologies exist within the sample at Gray. No osteological evidence of sexual dimorphism exists for tapirs, and no correlation between crest shape and sex was observed. Several juveniles display well defined crests, while some adults have only minor thickening of the temporal ridges; therefore, no distinct correlation between age and crest state could be established. Three different patterns of wear exist within the sample, but there is no correlation between these and crest morphology. No cranial pathologies were found to be associated with a crest type. Despite some correlations the sagittal crest could simply be a variable character due to intraspecific variation.

Excavations of ACb-3 Cave have uncovered the remains of at least seven individual Megalonyx jeffersonii, providing the most complete ontogenetic sequence of the taxon from one locality. Four individuals representing four distinct age classifications (infant, juvenile, subadult, and adult) were described and examined for pathologies. Cranial and major forelimb elements depict a change in morphology (trending from robust and stout to gracile and elongate) through ontogeny. Pathologies on the scapula and radius of the adult ground sloth indicate a potential attack from fighting or mating. The presence of infant and juvenile sloths suggests this cave may have been used as a maternity den. The nearly complete adult (RMM 5353) was compared to Megalonyx from other North American localities. Bivariate plots, created using linear measurements, showed that RMM 5353 was within the size range of M. jeffersonii and further supports the concept of Megalonyx chronospecies.

The paleobiology of the Pleistocene North American giant short-faced bear, Arctodus simus, has eluded paleontologists for decades. Its more gracile form has led past researchers to myriad intepretations of the locomotion and feeding ecology of this species. While earlier studies have focused on craniodental morphology and simple postcranial indices, it is forelimb morphology that represents a direct compromise between locomotor and foraging behavior. The study here uses traditional and 3D landmark morphometrics to more completely compare the 3-dimensional shape of the major forelimb elements and their muscle attachment sites between A. simus, extant ursids, and other carnivorans. Results herein agree well with previous studies and provide additional evidence for reduced abductor/adductor and supinator/pronator musculature, more restricted parasagittal motion, increased stride length, and lighter and more packed distal elements. Forelimb skeletal morphology therefore supports the hypothesis that A. simmus represents a bear in the early stages of cursorial evolution.