Rangeomorphs are a distinct group of millimeter- to meter-scale soft-bodied macrofossils that are restricted to the latter half of the late Neoproterozoic Ediacaran Period (635Ma- 542Ma). These fossils represent an extinct higher level taxon characterized by a modular construction based on a single architectural unit: the centimeter-scale, chevron-shaped rangeomorph element which displays several orders of self-similar branching. These elements could be arranged in a variety of different ways, constituting the wide array of gross morphologies found within the Group Rangeomorpha. The largest and most diverse assemblage of rangeomorph fossils in the world is found at Mistaken Point, on the Avalon Peninsula of Newfoundland, Canada, where these organisms are preserved within their original, in situ paleocommunities. Multibranched rangeomorphs are typified by bush-, comb- and network-shaped fossils which display multiple rangeomorph-bearing structures attached to an untethered basal stolon or central attachment point. Multibranched, comb-shaped rangeomorphs are endemic to Mistaken Point, and are represented by fossils displaying multiple parallel struts emerging along one side of an elongate, curved pedicle rod. Morphological and taphonomic evidence suggests that, in life, this organism had two rows of struts, each bearing a rangeomorph frondlet, arranged in an alternating pattern along the curved, tubular pedicle rod. Biometric analyses imply that the struts were added to both ends of the pedicle rod throughout the organism’s lifetime, with later inflation of the rangeomorph frondlets. Each comb-shaped rangeomorph locality likely represents a different age cohort within the organism’s lifecycle, providing rare evidence for spatfall reproduction in Ediacarans, which is similar to that found in modern macrobenthic organisms with pelagic larvae. Network-shaped multibranched rangeomorphs, represented by symmetrical to asymmetrical net-like fossils, are also endemic to Mistaken Point. This genus is reconstructed as having a symmetrical arrangement of flexible, rangeomorph-bearing leaflets that were, in part, neutrally buoyant with respect to the seawater. This flexible leaflet structure is unique, and shared only with a rare, previously undescribed, Ediacaran frond-like organism. It is suggested that the enigmatic leaflet structures shared by these two morphologically distinct taxa represent a new type of rangeomorph branching architecture, and therefore constitute a new type of rangeomorph. / Thesis (Master, Geological Sciences & Geological Engineering) -- Queen's University, 2008-07-25 11:01:43.469

Hiemalora is a genus residing within the Ediacaran biota. There are just two species of Hiemalora; H. stellaris and H. pleiomorphus. Despite there being only being two species within this genus, there is a degree of morphological variability within each species which is highlighted by the description of specimens around the world. The distribution of Hiemalora is global with specimens observed in Newfoundland, England, the White Sea (Russia), Siberia, Australia and arctic Norway. There has been a great deal of evolution in the interpretation of Hiemalora since the original description of H. stellaris in 1980. This genus was first interpreted as a hydrozoan with a central polyp with radiating tentacles (Fedonkin. 1981). It is now thought to be associated with rangeomorphs as a holdfast-type basal attachment structure. It is most closely linked to the frondose taxon Primocandelabrum which is abundant in Mistaken Point, Newfoundland and Charnwood Forest, England.The Hiemalora fossils in this study are sampled from the Indreelva member on the Digermulen Peninsula, Finnmark, Norway during fieldwork in both 2015 and 2018 at two close although separate localities. The most abundant species in this study is H. stellaris which makes up the majority of the 9 samples. There is also the potential inclusion of a H. pleiomorphus specimen, which features a distinctly different morphology to the other Hiemalora samples. One specimen has been identified with what is possibly a Primocandelabrum frond attachment. It is almost certainly a rangeomorph although the affinity is up for debate. One of the main aims for this study is to, as accurately as possible, determine the taxonomic affinity for all the specimens. This is subsequently an aim to better understand the biota inhabiting the Ediacaran seafloor in Digermulen Peninsula and the palaeoenvironment. This study is the first incorporating Hiemalora concerning material from the Digermulen Peninsula since the work done since Farmer et al. in 1992 and the first study in this area solely focused on the genus. Due to this, the study is allowing for the comparison between modern interpretations of Hiemalora from other Ediacaran locations around the world. Due to the Digermulen Peninsula residing on Baltica during the Ediacaran, comparisons to Avalonian assemblages such as in Newfoundland and England may highlight polarising features or many similarities. / Hiemalora är ett släktskap av skivformad organism som levde under Ediacaraperioden. Ediacara är den sista perioden under Neoproterozoicum och varade från 631Ma till 541Ma. Fossil av Hiemalora är globalt distribuerade, med exemplar från Ryssland, England, Kanada, Australien och Norge. Hiemalora-exemplaren i denna studie hittades på Digermulen-halvön i Finnmark, Norge. Det har funnits många olika tolkningar av Hiemalora kring dess livsstil, utseende och dess affinitet inom fylogenien eftersom den först beskrevs 1980. Den enkla uppbyggnaden hos fossilen, med bara en central skiva och "tentakelliknande" strålar som härrör från det i ett plan lämnar mycket för tolkning. Hiemalora är relativt sällsynt i jämförelse med andra skivformade fossil. Andra discoidala Ediacarafossil som Aspidella är mycket vanligare. Det finns två arter av Hiemalora som har beskrivits. Dessa är H. stellaris och H. pleiomorphus. Det finns subtila morfologiska skillnader mellan dessa arter. Skillnader mellan dessa arter inkluderar parallella ribbliknande strukturer på ytan av den centrala skivan av H. pleiomorphus och tätt packade strålar. Den centrala skivan i H. pleiomorphus är också proportionellt större i jämförelse med dess centrala skiva. H. stellaris har associerats med fronds. Fronds är en förgreningsstruktur som är orienterad vertikalt i vattenkolumnen. Dessa strukturer är associerade med olika skivliknande organismer och kan ha en stor morfologisk variation. Den frond som är associerad med H. stellaris är Primocandelabrum. Denna frond ses vid H. stellaris fossil i Newfoundland, Kanada. En frond som är tveksamt identifierad som Primocandelabrum ses vid en av Hiemalora-exemplaren i denna studie. Betydelsen av detta skulle vara att det är det första Hiemalora-exemplaret utanför Newfoundland och Charnwood Forest, England som bevaras med en Primocandelabrum frond. Det finns nuvarande idéer att Primocandelabrum och H. stellaris kan vara samma organism fast med olika delar bevarade. Faktumet att båda delarna är bevarade på Digermulen halvön kan ge mer inblick i detta. Sju av Hiemalora-exemplaren i denna studie är H. stellaris, medan en art möjligen är H. pleiomorphus. Provet med fronden är sannolikt H. stellaris, men på grund av att den är dåligt bevarad runt centralskivan är dess affinitet inte säker. En dålig bevarandekvalitet hos många exemplar i studien gjorde det svårt att identifiera den. Bevarandet ger dock en inblick i miljön som Hiemalora bodde i och under vilka förhållanden den dog. Många platser runt om i världen har exceptionellt bevarade fossil som kan ha varit möjligt på grund av en händelse där alla blev bevarade samtidigt, likt de vulkaniska askadepositionerna i Newfoundland. Detta var inte fallet i Digermulen-halvön, där organismer sannolikt inte begravdes omedelbart efter döden.