Conselho Nacional de Desenvolvimento Científico e Tecnológico / This work was developed with the aim of studying the efficiency of the models that express the relationship of density and diameter in evenaged stands of Pinus taeda L., managed in full density and highly stocked for a bioeconomic model of production. For this, were tested density and diameter models, using data derived from 50 permanent plots, measured annually until 18 age, allocated to settlements established in the spacing of 1.5x1.0 m, 2.5x 1.0 m, 1.5x2.0 m, 2.0x2.0 m, 1.5x3.0 m, 2.5x2.0 m, 2.0x3.0 m and 2.5x3.0 m, kept at full density, in Otacílio Costa, physiographic region of Santa Catarina State plateau.The results of the bioeconomic modeling of production, based on the density management diagram system for tall trees, we concluded that the models proposed by Reineke, Yoda, Zeide and Tang showed a good statistical precision, with a determination coefficient higher than 0,88 and coefficient of variation less than 1.25 % but the model of Tang was more accurate and efficient. The size of the diameter at the point where you start self-thinning in the population varies considerably with the density of trees established per hectare. For all plant spacings analyzed did not prove the value of the slope of -3/2 law of self-thinning proposed by Tang and is a result of the initial spacing. The limit of self-thinning in less dense populations, occurs with a diameter slightly less than in maximum density, which is directly proportional to the density of trees when the implementation of the forest. The model of self-thinning described the management zones of great individuals in the population, between 0.45 and 0.55 limits the maximum density of the forest. The annual increment in diameter between 6 and 7 years, when related to relative density (G/d0,5622) identified five areas of growth, defined by: excessive space - trees grow free of competition;free growth - where growth in diameter is maximum; increased competition- when the diameter increment begins to decrease; full density - when productivity of the stand is maximum; and imminent death - begins when individuals have died by excessive density and high competition. The Diagrams Density Management of the variables mean diameter, basal area, volume by the Population Density Index was efficient and to estimate thisis values with accuracy, showed an efficiency of 0.99. The lines demarcating the areas of competition allow us to project the trajectory of the diameter and density, with the definition of the clearcutting. The Normal, Lnnormal, Weibull and Gamma probability distributions, described accurately the variation of the density of the stands, but the Weibull function with two parameters was comparable to the adjustment by the simplicity and efficiency in the prognosis of frequencies for diameter in time. The shape of the trunk was adjusted by the polynomial of five degree, allowed the best estimates than other models. The net present value of R$ 15587.60 ha-1 was given to the management regime with three thinnings at 9, 13 and 18 years and cut at 22 years, higher than the other schemes simulated. A comparison of management regimes for the equivalence of planning horizons for 28 years, determined by the equivalent annual value, said the management regime with three thinned and cut at 22 years, such as greater economic efficiency, with a value of R$ 1380.10 ha-1. The internal rate of return in any simulated management regimes was highly attractive, with varying from 14.83 % per year in managed regimes stands with three thinning and cut to 22 years to 14.25 % per year in managed regimes stands with threethinning and cut to 18 years. The biggest benefit/costs ratio was 2.81, obtained in the management regime with three thinnings, at 9, 13 and 18 years and final cutting at 22 years. / Este trabalho foi desenvolvido com o objetivo de estudar a eficiência dos modelos que expressam a relação de densidade e diâmetro em povoamentos equiâneos de Pinus taeda L., manejados em densidade completa e altamente estocados, para obtenção de um modelo bioeconômico da produção. Para isso, foram testados modelos de densidadee diâmetro, através de dados originados de 50 parcelas permanentes, medidas anualmente até os 18 anos, alocados em povoamentos implantados em espaçamentos de 1,5x1,0 m, 2,5x1,0 m, 1,5x2,0 m, 2,0x2,0 m, 1,5x3,0 m, 2,5x2,0 m, 2,0x3,0 m e 2,5x3,0 m, mantidos em densidade completa, em Otacílio Costa, região fisiográfica do planalto do estado de Santa Catarina. Os resultados obtidos da modelagem bioeconômica da produção, baseada no Diagrama de Manejo da Densidade para sistema de alto fuste, permitiram concluir que os modelos propostos por Reineke, Yoda, Zeide e Tang apresentaram uma boa precisão estatística, com um coeficiente de determinação superior a 0,88 e um coeficiente de variação inferior a 1,25%, porém o modelo de Tang mostrou-se mais preciso e eficiente. A dimensão do diâmetro médio no ponto em que inicia o autodesbaste na população varia consideravelmente com a densidade de árvores implantadas por hectare. Para todos os espaçamentos de plantio analisados, não se comprovou o valor do coeficiente angular de -3/2 da lei de autodesbaste proposto por Tang, sendo uma decorrência do espaçamento inicial. O limite de autodesbaste de populações menos densas ocorre com um diâmetro médio pouco inferior ao obtido em máxima densidade, sendo este diretamente proporcional à densidade de árvores quando da implantação da floresta. O modelo de autodesbaste descreveu as zonas de manejo ótimo dos indivíduos na população entre limites de 0,45 e 0,55 da densidade máxima da floresta. O incremento corrente anual em diâmetro entre o 6º e o 7º ano, quando relacionado com a densidade relativa (G/d0,5622), permitiu identificar cinco zonas de crescimento, definidas por: espaço excessivo as árvores crescem livre de concorrência; crescimento livre o incremento em diâmetro é máximo; aumento da competição o incremento em diâmetro começa a diminuir; estoque completo a produtividade do povoamento é máxima; e iminente mortalidade começa a haver morte de indivíduos por densidade excessiva e alta concorrência. Os Diagramas de Manejo da Densidade com as variáveis diâmetro médio, área basal e volume por Índice de Densidade do Povoamento foram eficientes e permitiram estimar estes valores com acurácia, para uma eficiência igual a 0,99. As linhas que delimitam as zonas de concorrência permitem projetar a trajetória do diâmetro por densidade populacional, com definição da idade de corte final. As distribuições de probabilidade Normal, Ln-normal, Weibull e Gama descreveram com acurácia a variação da densidade dos povoamentos, masa função de Weibull, com dois parâmetros, foi compatível pela simplicidade ao ajuste e à eficiência na prognose das frequências por diâmetro no tempo. A forma de tronco ajustada pelo polinômio do 5º grau apresentou melhores estimativas que outros modelos. O Valor Presente Líquido de R$ 15.587,60 ha-1 foi determinado para o regime de manejo com três desbastes, aos 9, 13 e 18 anos e corte final aos 22, superior aos demais regimes simulados. A comparação dos regimes de manejo pela equivalência dos horizontes de planejamento em 28 anos, determinado pelo Valor Anual Equivalente, indicou o regime de manejo com três desbastes e corte final aos 22 anos como o de maior eficiência econômica, com um valor de R$ 1.380,10 ha-1. A taxa interna de retorno em qualquer dos regimes de manejo simulados foi altamente atrativa, tendo variado de 14,83 % a.a. em povoamentos manejados com três desbastes e corte final aos 22 anos a 14,25 % a.a. em povoamentos manejadoscom dois desbastes e corte final aos 18 anos. A maior razão benefício sobre os custos foi de 2,81, obtida no regime de manejo com três desbastes, aos 9, 13 e 18 anos e corte final aos 22.

Stand carrying capacity is the maximum size of population for a species under given environmental conditions. Site resources limit the maximum volume or biomass that can be sustained in forest stands. This study was aimed at estimating and determining the carrying capacity in loblolly pine. Maximum stand basal area (BA) that can be sustained over a long period of time can be regarded as a measure of carrying capacity. To quantify and project stand BA carrying capacity, one approach is to use the estimate from a fitted cumulative BA-age equation; another approach is to obtain BA estimates implied by maximum size-density relationships (MSDRs), denoted implied maximum stand BA. The efficacy of three diameter-based MSDR measures: Reineke's self-thinning rule, competition-density rule and Nilson's sparsity index, were evaluated. Estimates from three MSDR measures were compared with estimates from the Chapman-Richards (C-R) equation fitted to the maximum stand BA observed on plots from spacing trials. The spacing trials, established in the two physiographic regions (Piedmont and Coastal Plain), and at two different scales (operational and miniature) were examined and compared, which provides a sound empirical basis for evaluating potential carrying capacity. Results showed that the stands with high initial planting density approached the stand BA carrying capacity sooner than the stands with lower initial planting density. The maximum stand BA associated with planting density developed similarly at the two scales. The potential carrying capacity in the two physiographic regions was significantly different. The value of implied maximum stand BA converted from three diameter-based MSDR measures was similar to the maximum stand BA curve obtained from the C-R equation. Nilson's sparsity index was the most stable and reliable estimate of stand BA carrying capacity. The flexibility of Nilson's sparsity index can illustrate the effect of physiographic regions on stand BA carrying capacity. Because some uncontrollable factors on long-term operational experiments can make estimates of stand BA carrying capacity unreliable for loblolly pine, it is suggested that the stand BA carrying capacity could be estimated from high initial planting density stands in a relatively short period of time so that the risk of damages and the costs of experiments could be reduced. For estimating carrying capacity, another attractive option is to choose a miniature scale trial (microcosm) because it shortens the experiment time and reduces costs greatly. / Master of Science

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Made available in DSpace on 2016-12-08T16:24:12Z (GMT). No. of bitstreams: 1 PGCA13MA094.pdf: 3484558 bytes, checksum: 4cd098df0840598b9d19fbbc52ff65cb (MD5) Previous issue date: 2013-03-04 / Coordenação de Aperfeiçoamento de Pessoal de Nível Superior / In regions with subtropical climates the use of grasses intercropping, mainly oats and ryegrass is a common practice, but the advantages and disadvantages of the intercrop in relation the pure culture are little studied. The aim of this work was to evaluate the production and persistence of black oat and ryegrass swards grown pure or intercropping throughout the period of use. The three treatments were randomly distributed into a complete blocks design with four replicates. The pre-grazing sward heights were 23, 20 and 17 cm for the oat, intercrop and ryegrass treatments, respectively. The post-grazing sward heights were 40% lower than their initial heights. Morphogenesis was performed using the technique of marked tillers and dynamic and tiller population density (TPD) was determined with three 10-cmdiameter PVC rings per paddock. The treatments did not differ in the final length of intact leaf, being the October month with the lowest values (approximately 48% lower). The phyllochron was not affected by the form of cultivation, but the values were increasing with the advance of the growing season. The leaf elongation rate decreased over the months, with a mean value of 0.14 cm / perfilhos.graus-day for all treatments. The advance of the growing season resulted in increases stem elongation rates, with the species in the intercrop having smaller increments in the final months. In general, rates of senescence reduced with the advance of months, and the intercrop increased tissue death of ryegrass. The intercrop TPD remained constant until October, inverting the participation of species in August. Of the tillers existing on the last grazing cycle, approximately 0, 40 and 20% were first generation of the oats, ryegrass and intercrop, respectively. Thus, despite of ryegrass pure have the lowest rate of tillering, it have the same rate of population stability of the intercrop. The pure ryegrass had the highest average net accumulation rates by offering smaller forage losses by senescence. Tiller size/density compensation was observed in the three plant communities (treatments) according to the self-thinning rule. In addition, no relationships were found when each species was analysed individually in the intercrop treatment. For the species tested the developmental stage of sward has greater influence on the morphogenesis and structural characteristics than cultivation form. The morphological similarity of oats and ryegrass enables evaluated traits similar in both forms of cultivation. When intercropping the temperate climate grasses under rotational grazing, size/density compensation mechanisms occurred throughout the grazing cycles and in the plant communities the same form of the pure cultivation to maintain the leaf area index and production capacities of the pastures / Em regiões de clima subtropical a utilização de gramíneas em consórcio, principalmente aveia e azevém, é uma prática comum, porém as vantagens e desvantagens do consórcio em relação ao cultivo puro são pouco estudadas. Objetivou-se avaliar a produção e a longevidade de pastos de aveia-preta e azevém anual cultivados puros ou em consórcio ao longo de todo o período de utilização. Os três tratamentos foram distribuídos em blocos completos casualizados com quatro repetições, tendo cada unidade experimental 98 m2. As alturas em pré-pastejo foram 23, 20 e 17 cm para a aveia, o consórcio e o azevém, respectivamente, sendo rebaixados em 40% destas alturas. A morfogênese foi realizada por meio da técnica de perfilhos marcados e a dinâmica e densidade populacional de perfilhos (DPP) em três anéis de PVC com 10 cm de diâmetro por piquete. Os tratamentos não diferiram quanto ao comprimento final de folha intacta, ficando o mês de outubro com os menores valores (aproximadamente 48% menor). Não houve efeito da forma de cultivo para os valores de filocrono, porém os valores foram crescentes com o avanço da estação de crescimento. A taxa de alongamento foliar reduziu com o passar dos meses, com valor médio de 0,14 cm/perfilho.grau-dia para todos os tratamentos. O avanço da estação de crescimento resultou em incrementos nas taxas de alongamento de colmos, com as espécies no consórcio apresentando menores incrementos nos meses finais. De maneira geral, as taxas de senescência foliar reduziram com o avanço dos meses, sendo que o consórcio aumentou a morte de tecidos do azevém. O consórcio manteve a DPP constante até outubro, apresentando uma inversão na participação das duas espécies em agosto. Dos perfilhos existentes no último ciclo de pastejo, aproximadamente 0, 40 e 20% eram da primeira geração no azevém, na aveia e no consórcio, respectivamente. Com isso, apesar de apresentar a menor taxa de aparecimento de perfilhos o azevém cultivado puro apresentou o mesmo índice de estabilidade populacional do consórcio. O azevém cultivado puro apresentou as maiores taxas de acúmulo líquido médio por apresentar as menores perdas de forragem por senescência. A compensação tamanho/densidade de perfilhos foi observada segundo a lei do auto-desbaste (self thinning rule) nos três tratamentos quando levados em conta todos os perfilhos da área, não sendo encontrada relação quando as espécies foram analisadas individualmente no consórcio. Para as espécies testadas o estádio de desenvolvimento dos pastos apresenta maior influência sobre as características morfogênicas e estruturais do que a forma de cultivo. A semelhança morfológica da aveia e do azevém possibilita características morfogênicas e estruturais parecidas em ambas as formas de cultivo. É sugestivo que em consórcios entre gramíneas de clima temperado sob lotação intermitente os mecanismos de compensação tamanho/densidade operem na comunidade de plantas e ao longo dos ciclos de pastejo da mesma forma que nos cultivos puros, mantendo elevado índice de área foliar e capacidade produtiva

Des changements de croissance ont été documentés pour le XXe siècle dans de nombreuses régions en Europe. Cependant une évaluation exhaustive des changements de productivité, à une large échelle géographique, avec une analyse de leur hétérogénéité spatiale et de la diversité interspécifique de la réponse fait encore défaut. L'objectif de cette thèse est d'évaluer les changements récents de la productivité forestière aux échelles nationale, régionale et locale en France, et de rechercher leurs causes environnementales, à partir d'une approche de modélisation statistique de l'accroissement en surface terrière du peuplement (∆G) et d'indicateurs environnementaux. Nous avons utilisé les données de l'inventaire forestier national français pour 8 espèces dont la niche écologique et la distribution diffèrent (Fagus sylvatica, Quercus robur, Quercus petraea, Quercus pubescens, Picea abies, Abies alba, Pinus sylvestris and Pinus halepensis), observées en peuplements purs et réguliers. Nous présentons les facteurs ayant un effet sur la productivité des différentes espèces à l'échelle nationale, ainsi que des cartes de productivité suggérant que l'aire de distribution des espèces n'est pas toujours limitée par les conditions environnementales. Entre 1980 et 2005, nous observons des tendances positives, modales ou non significatives de la productivité pour toutes les espèces à l'exception des deux espèces méditerranéennes dont la productivité a diminué, soulignant ainsi la variabilité interspécifique de ces changements. Nous observons également de fortes variations spatiales des changements de productivité, autant dans leur intensité que dans leur signe, aux échelles régionale et intra-régionale. Ces résultats remettent en question la pertinence d'une évaluation moyenne à large échelle et soulignent le caractère contextuel des estimations. Nous mettons en évidence le rôle du réchauffement climatique récent dans les tendances observées. Ce travail fournit une évaluation plurispécifique et multi-échelle de la réaction de la productivité des espèces arborées à un environnement changeant. Nous avons souligné le caractère spécifique des changements de productivité et leur caractère contextuel, du fait de différences dans l'autécologie des espèces et de variations spatiales des facteurs limitants. Une étude approfondie de l'effet des facteurs environnementaux et de leurs interactions complexes est nécessaire pour la prédiction de la productivité future des espèces. / Growth trends have been reported in many regions of Europe over the twentieth century. However, an integrated assessment of productivity changes, including focus on a wide geographical scale, analysis of spatial heterogeneity, and the inter-specific diversity of growth responses is still lacking. The aim of this Ph.D work was to assess recent changes in forest productivity on a national, regional and local scale in France, and to investigate their potential environmental causes, based on statistical modeling approaches of stand basal area increment (BAI), and environmental indicators. We used the French NFI data for 8 species of contrasted ecological niches and distributions (Fagus sylvatica, Quercus robur, Quercus petraea, Quercus pubescens, Picea abies, Abies alba, Pinus sylvestris and Pinus halepensis), taken in pure and even-aged stands. We identified the main factors influencing tree species productivity on a national scale, and produced productivity maps suggesting that species distribution ranges are not always limited by environmental conditions. Between 1980 and 2005, the productivity trends reported were positive, modal or non-significant for all species, except the two Mediterranean species for which productivity decreased, highlighting inter-specific differences in these changes. We observed strong variations of productivity changes, both in intensity and sign, on a regional and intra-regional scale. These results question the relevance of wide-scale average assessments and highlight their context-dependence. The role of recent climatic warming in featuring the BAI trends was highlighted. This work provides an enriched scale- and species-dependent assessment of tree species reaction to a changing environment. We emphasized the species- and context dependence of productivity changes, due to differences in species autecology and spatial variations in the limiting factors. Further focus on the effect of environmental factors and their complex interactions is needed for the prediction of species future productivity.

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