Solen cylindraceus is an infaunal filter-feeding bivalve inhabiting the intertidal mud banks of many southern African estuaries. It is particularly abundant in the Kariega estuary (33°41'S; 26°42'E) where it reaches densities of 400m⁻² (192g shell-free dry wt. m⁻²). The Kariega is a permanently open, marine dominated estuary about 18km in length, and S. cylindraceus is most abundant in its middle and upper reaches. Some physical characteristics of the estuary (temperature, salinity, sediment and water turbidity) are described, and the possible role of these factors in determining the density and distribution of S. cylindraceus within the Kariega estuary, is discussed. The structure of the alimentary system, gills and labial palps of S. cylintfraceus is described, all of which showed no major variation from the "typical" eulamellibranchiate form. Solen cylintfraceus was found to be a euryhaline osmoconformer with a salinity tolerance range of 15-65%. When animals were removed from their burrows, osmotic equilibration of the haemolymph was rapid (1-2 hours). By contrast, in animals left undisturbed in their burrows, osmotic equilibration was retarded (72-204 hours). It is suggested that the observed decrease in the rate of change of haemolymph osmolarity for animals in their burrows is linked to the stability of the interstitial salinity. A temperature tolerance range of 5-44°C was determined for S. cylintfraceus (in situ), in which prolonged exposure to 5°C and 40-45°C (12-36 hours respectively) resulted in a decreased burrowing ability, coma and death. Animal burrowing responses were not affected by temperatures in the range 15-35°C. Field experiments were carried out over several tidal cycles, in which the measurement of crystalline style volume was used as a means of assessing extracellular digestive activity. No major variation in style volume was recorded and it appeared that S. cylindraceus did not exhibit any cyclical pattern of style dissolution and regeneration. It is suggested that S. cylindraceus feeds continuously from the water column during high tide and possibly within its burrow, at or below the water table, during low tide. At a suspensoid concentration of 5Omg l⁻¹, S. cylindraceus was found to filter water almost continuously (90-95% of the time). Time spent filtering dropped to 68% at 100mg l⁻¹ and 32% at 500mg l⁻¹. Filtration rates for summer collected animals (25°C) were 22.86 ± 4.36ml min.⁻¹, some 3ml min.⁻¹ greater than that recorded for winter (16°C) collected animals. Filtration rate may be expressed as a function of shell length by the equations: y=0.247x¹̇⁰⁶⁶ (winter) and y=0.758x⁰̇⁸²⁶ (summer). Solen cylindraceus was capable of acclimating its filtration rate to both high and low temperatures under laboratory conditions. Filtration rate exhibited a thermal optimum in the range 15-35°C, declining at higher and lower temperatures. Q₁₀ values of filtration decreased rapidly from greater than 4 to less than 2, when the thermal optimum was reached. Maximum rates generally occurred at approximately 5°C above the temperature to which the animal had been acclimated. Optimal filtration rates (19-23ml min.⁻¹) were recorded in the salinity range 15-45%. When subjected to abrupt changes in salinity, filtration rates were immediately depressed. The extent and duration of these decreased filtration rates were dependent upon the magnitude and direction of salinity change, and were always less in animals exposed to hyper- than hyposaline conditions. Animals exposed to increased temperature and simultaneous elevated or unchanged salinity, showed a slight increase in filtration rate followed by rapid acclimation. A decrease in both temperature and salinity resulted in an initial decrease in filtration rate and a longer acclimation period. The ability of S. cylindraceus to acclimate fully within a wide temperature and salinity range, and to filter maximally in hypersaline conditions may, in part, explain its unusually high abundance in the Kariega estuary, despite it being close to the southernmost limit of the animal's geographical distribution. No significant difference in flItration rate was recorded at suspensoid concentrations of 5-100mg 1⁻¹. However, at 250 and 500mg l⁻¹ filtration rates decreased significantly, and coincided with increased levels of pseudofaecal production. Solen cylindraceus retained particles down to 2.5-3.0µm with great efficiency (ca. 60-90% efficiency). Below this particle size, retention efficiency decreased rapidly and a net production of particles was recorded below 1.51µm. Particle retention was independent of temperature (15 and 25°C) and salinity (15 and 35%). Use was made of stable carbon isotope analyses (¹³C/¹²C ratios) in an attempt to determine the important food sources of S. cylindraceus within the Kariega estuary. The results obtained demonstrated an enrichment in δ¹³ values for S. cylindraceus from the upper (-27.9%) to the middle (-25%c) and lower (-21.6%o ) reaches of the estuary, with no seasonal variation apparent. The bivalve was substantially more depleted in ¹³C relative to the dominant aquatic macrophytes Zostera capensis (-9.1 to -15.6%o) and Spartina maritima (-12.5%o). The use of δ¹³ alone, however, to unequivocally "pin point" specific food sources of a filter feeder in a predominantly detritus based food web, is limited. It is suggested that in the Kariega estuary, riparian litter and other terrestrially derived vegetation contribute to the carbon pool. A possible contribution of ¹³C depleted food sources via chemoautotrophic and/or anaerobic pathways, to the diet of S. cylindraceus, is suggested.

Tick borne diseases of livestock are among the most important factors which have retarded economic development in many parts of Africa. Modification of the environment due to the expansion of agriculture into previously undeveloped areas, together with the introduction of livestock, has disturbed the natural balance which existed between ticks and indigenous hosts. Many tick species have adapted to domestic stock, and in some instances have spread over large areas which were previously uninfested. A number of tick species which are parasitic on domestic stock, eg. Amblyomma hebraeum Koch, 1844, Boophilus decoloratus (Koch, 1844), Rhipicephalus evertsi Neumann, 1897 and Rhipicephalus appendiculatus Netunann, 1901, are vectors of pathogenic micro-organisms. Unlike indigenous hosts however, domestic stock have no innate immunity to tickborne diseases. Intro., p. 1.

The Kirkwood Formation in the Bushman's River area of the Algoa Basin is characterised by a number of fining-upward cycles. These have been interpreted as indicating deposition in a dynamic aggrading meandering river system with the channel deposits (conglomerates grading upwards into sandstones) fining upward into the overbank deposits (mudrocks). Channel, channel-margin and overbank deposits were recognised. The three mudrock sequences logged comprise compound pedofacies sequences of multistorey, simple and cumulative palaeosols. Distinctive palaeopedological features, such as root traces and pedotubules, soil horizons and structures, mottles, and iron-rich and calcareous glaebules and calcareous hardpan lenses and layers were used to identify a number of palaeosols within the mudrock sequences. Each mudrock sequence comprises multistorey entisol, inceptisol, alfisol, ultisol, aridisol and vertisol profiles at different stags of pedogenic maturity. The entisols and inceptisols are relatively immature profiles formed close to the meandering river channel and are classified as channel-margin palaeosols. The ultisols, alfisols, aridisols and vertisols are more mature and formed at some distance from the channel. They are classified as proximal floodbasin or distal floodbasin palaeosols depending on their maturity, distance from the channel and grain-size. Slickensides, desiccation cracks, and iron-rich concretions occur, indicating multiple cycles of wetting and drying. A low water-table beneath the floodplain is indicated by both the prominent maroon-brown colouration of the mudstones, caused by oxidation during deposition, and the general lack of evaporites in the sequence. Calcretes comprising calcic and petrocalcic horizons are very common in the lower mudrock sequence, rare in the middle mudrock sequence and relatively common in the upper mudrock sequence. The calcretes generally consist a nodular zone which may, in some cases, be capped by a thin hardpan layer. The calcic palaeosols commonly show stages of carbonate accumulation which indicate at least 10 000 years of formation. The lack of calcrete formation in some of the profiles may indicate frequent flooding and high sediment accretion rates or a decrease in the influx of Ca²⁺-rich aeolian dust into the depositional basin. Clay alluviation is common in many of the profiles and soil structures are commonly well developed. The palaeosols are interpreted as having formed on an aggrading floodplain in a warm to hot (25-30°C), semi-arid climate with a low but seasonal rainfall (100-500mm per annum).

[Introduction] During 1965 the author, in looking for a project for a thesis to be submitted for the degree of Master of Science in Geology, decided to map a suitable region in order to gain experience in geological field work. The exact nature of the region itself was of no great importance, but since this study was to be conducted through Rhodes University, it was decided that an area, as near to Grahamstown as possible, would be the most suitable. With this in view, the geologist in charge of the Grahamstown Office of the Geological Survey was invited to suggest an area suitable for study, and if possible, to obtain financial assistance. He indicated the region which has been mapped and which will eventually form part of the proposed sheet 143. It is immediately adjacent to, and to the west of the 1:125,000 sheet 136 of Grahamstown completed by Mountain in 1940.

The plant communities of the Andries Vosloo Kudu Reserve (6493 ha.; Acocks's Valley Bushveld) are classified using the Braun-Blanquet phytosociological technique. A checklist of the flowering plants is presented. The flora of the reserve displays an affinity to five phytochoria (Capensis, Karoo-Namib, Sudano-Zambezian, Afro-montane and Indian Ocean Coastal Belt 1, of which the Karoo-Namib and Indian Ocean Coastal Belt are the most extensive. The flora of the Valley Bushveld is regarded as adequately conserved, but a recommendation is made to consolidate the study area with adjacent farms. A small herbarium, containing specimens of all species listed in the checklist, has been established. The climate of the study area is classified according to the Koppen classification as Cfa. Simultaneous air temperature recording at four separate localities demonstrated that differences exist between local ities of marginally different altitude, aspect and slope. The phenology of 51 plant species, recorded at monthly intervals over a continuous period of 478 days, is presented. These species are classified according to their response to air temperature and rainfall for the initiation of new growth, flowering and fruiting. Species exhibited a spectrum of responses ranging from endogenous to exogenous control over phenological events. / KMBT_363 / Adobe Acrobat 9.53 Paper Capture Plug-in

The phenotypic plasticity of the cichlid Oreochromis mossambicus was examined in fresh and brackish water populations living in the eastern Cape near the southern extremity of the natural range. The primary goal of the study was to examine the wide variety of life-history styles which were expressed in relation to the varying environmental conditions. Four natural populations were studied during 1986 and 1987, and some experimental work under artificial conditions was carried out at the Rhodes University fishfarm during 1988. This comparative approach gave insights into the complex interactions between biotic and abiotic factors which are exploited by this hardy species. O. mossambicus displayed a wide variety of life-history tactics under harsh environmental conditions. The growth of juvenile fish was rapid and mortality of the young-of-the-year was high. Age at first onset of sexual maturity ranged from one to three years. Under harsh conditions growth rate was markedly reduced after maturity, which occurred during the second or third summer and growth disparity between the sexes was more pronounced. Longevity was found to vary from five to ten years for different populations. Under more favourable conditions (resembling the typical "floodplain" habitats of this species at lower latitudes) very rapid juvenile growth, and a high maximum mass of over 2300 g for both sexes was iii achieved. Sexual maturity under these conditions could be delayed until the third or fourth summer of life. O. mossambicus demonstrated an ability to withstand and even benefit from fluctuating water levels, and adjusted its life style according to the prevailing conditions: at times of high water level recruitment was low and sexual maturity delayed. During low water periods reproductive rates increased. Q. mossambicus showed a distinct preference for gentle gradients and well-vegetated habitats for feeding, nesting and refuge. During the study winter temperatures were near and at some times below the generally accepted lower limits for this species. This implies that these indigenous populations have a degree of cold tolerance. A further implication is that Q. mossambicus has the ability to thrive under adverse conditions which may partially explain its wide and successful colonisation of water bodies far from its natural range, and its obvious fisheries potential.

Several strategies used by the gobiid fish Caffrogobius caffer enabling it to colonise the harsh intertidal environment in the eastern Cape were studied. Q. caffer is opportunistic in its use of available food resources and feeds throughout the day with a slight tendency towards crepuscular peaks of feeding activity. The system of space utilisation used by C. caffer is so organised that all individuals of the population have access to patchily distributed food resources. Current velocities occurring over high tide limit the movements of Qo caffer and interpool movements occur only during the initial stages of a rising tide and in the final stages of an ebbing tide when the current velocities are below the tolerance limit of Q. caffer. The epifauna associated with the alga Ulva rigida, which forms an important constituent of the diet of the intermediate size classes of Q. caffer, was examined and it appears that this epifauna is an important source of food to Q. caffer. This report presents the results of the first detailed study of the biology of an intertidal fish in southern Africa.

[From Introduction]. It is the purpose of this thesis to examine some of the broad relationships which exist between different land use types and farming practises on the one hand and the geographical environment on the other. The enquiry is confined to the study of a number of farm units which have been selected within the area enclosed by the District of Albany and Bathurst situated in the south-East coastal belt of the Cape Province. The procedure adapted conforms broadly to that of a project carried out in South America by R.S. Platt of the University of Chicago and embodied in hls book "Latin America, Countryside and United Regions." Platt's project, he notes, consists of a collection of "small field studies in a frame of complex generalisations." It does not aim to give a complete geographical account of the regions he selected for investigation, but is an attempt to enlarge geograpical understanding of the respective areas.

It is widely believed that Acacia karroo Hayne trees compete with and negatively affect the production of grasses in their immediate surroundings. This perception formed as a result of observations by farmers of their veld as well as some research. The complexity of plant interactions in different conditions required that research be done to better explain the extent of grass and tree interactions. In response to this need, the effect of a range of Acacia karroo densities on grass production was tested in areas of differing rainfall. Results emphasize the complexity of such plant interactions but highlight rainfall as influencing the effect of tree density on grass production and grass quality. Study sites were chosen with significant differences in tree density and rainfall and this was found to be concomitant with significant differences in grass yield and crude protein (p < 0.001). Grass yield (mass produced) was negatively correlated with tree density (p < 0.001) and was affected by rainfall. High tree density reduced grass yield at the lower rainfall sites, but a reduction in yield was only recorded during dry conditions in the higher rainfall areas. By contrast, the crude protein content of grasses was positively correlated to tree density (p < 0.001). However, as with yield, rainfall played an important part in the strength of the correlation. Soil properties such as temperature (-) (p = 0.011), phosphorus content (+) (p = 0.006), calcium concentration (+) (p = 0.005), acid saturation (-) (p = 0.018), and soil pH (+) (p = 0.008) were also significantly correlated to tree density. Tree density enriched the soil, subsequently benefitting grass quality. The study sites investigated, had different plant communities and environmental conditions with Cradock, Kubusi Drift and Kei Mouth having unique communities and conditions while Adelaide and Cathcart could not be separated in either. Cradock was the driest site and the vegetation was dominated by both grasses and karroid shrubs, differing in pattern and process from the other sites. The Kubusi Drift site had high tree densities, to the extent that the trees altered the grass species composition. The effects of rainfall on grass yield and quality exceeded that of community composition differences.

The Cape Griffon Gyps coprotheres is listed as a globally threatened species, because of threats posed by, inter alia, poisoning, loss of habitat, food shortages, electrocutions and drowning in high-walled farm reservoirs. The Cape Griffon has undergone major decline in South Africa, including the Eastern Cape. A detailed investigation into the causes of this decline is vital to ensure the survival of the species. Since it is known that electrocution on powerlines is a major cause of mortality of the griffons in the Eastern Cape, the study therefore focuses on this factor in this region. More specifically, the overall aim or objective of the study is to investigate, through the application of appropriate GIS techniques, the use of descriptive, empirical, habitat and foraging models to prioritise powerline networks, in the Eastern Cape, for mitigation to prevent electrocution of Cape Griffons Gyps coprotheres. The specific activities associated with the aim of the study are: (1) to identify the broad habitat types used by the griffons, (2) to determine the mean daily foraging range size of the griffons, (3) to map currently active foraging ranges and to categorise them according to their role, and the level of importance, in the foraging and breeding behaviour of the species, (4) to map the locations of networks of griffon-unfriendly powerline networks, and (5) to integrate the outcomes of (1) to (4) above, to produce a spatially explicit product that ranks griffon-unfriendly powerline networks according to their priority level for mitigation. The study uses simple descriptive models to investigate, spatially, the threat posed by griffon-unfriendly powerlines to the Cape Griffon in the Eastern Cape. The modelling approach includes a broad level simple habitat model (Categorical model), a Maxent model, a spatial foraging model and an electrocution model. Several map outputs were produced from the analyses. Cape Griffon habitat was successfully modelled using ecological input variables: biome, vegetation xvii type (suitable/unsuitable for griffon foraging), stock (cattle, sheep and goats), and stock farming types (commercial & communal). All outputs were modelled through the use of a Geographical Information System (GIS). Opportunistic sightings data were used for the study, which means that the data were collected on a non-systematic basis and are therefore considered incomplete. In ecology and zoogeography, incomplete datasets are common. To address this issue, a maximum entropy (Maxent) model was applied to the available dataset. Maxent produces predictions or conclusions from incomplete information, and was therefore useful for this study. The mean daily foraging area (coverage) of the Cape Griffon in the study area was estimated from published and unpublished marked bird studies conducted elsewhere, but in generally similar habitats. Based on all the data from these studies, it was considered appropriate to use a circular area, with a radius of 40 km, to represent the mean daily foraging range of the Cape Griffon in the Eastern Cape. It was considered appropriate to use a roughly circular area to depict the daily foraging area, given that the Cape Griffon qualifies as a central place forager. This means that the griffons are tied to a permanent site (roosting site or breeding site) and forage within a certain area around that central site. The mean daily foraging area (40 km) was applied to the currently active griffon sites: breeding sites, regular roosting sites, seasonal/occasional sites, and roosting sites (uncertain status). This produced four foraging area types, which formed the basis for the creation of two spatial foraging area models: Hierarchical Foraging Area Model (HFAM) and the Intersecting Foraging Area Model (IFAM). The HFAM produced three outputs: Primary, Secondary and Tertiary Foraging Areas. These three areas represent the hierarchical presentation of the relative importance of the four foraging area types. The outcome was areas of different predicted griffon occurrence: ‘high’, ‘medium’, ‘low’. The IFAM is essentially an extension of the HFAM, which represents a single area where foraging ranges based on all four types are intersected. The outcome represents the area of predicted highest griffon occurrence. The identified areas of different predicted griffon occurrence, or density (outputs of the HFAM and IFAM), were individually intersected with the layer that indicates the locations of the ‘unsafe’ powerlines. These intersects are depicted in hierarchical mode and expressed according to a number of four risk categories (very high, high, medium and low). The final spatial output of the study is a map that identifies priority powerlines for mitigation against griffon mortality caused by electrocutions and collisions with powerline infrastructure.