Hydrothermal vents are deep-sea hot springs. Vents are home to luxuriant assemblages of animals that colonize the warm venting fluids. High biomass is fed by microbes that use hydrogen sulphide and other reduced chemicals in the vent fluid as an energy source to fix inorganic carbon. Individual vents may persist for a few years to several decades. The specialized animals must find new vents, cope with changing fluid conditions and foster their offspring. The composition and structure of vent communities vary in space and time. My research at Axial Volcano, a seamount on the Juan de Fuca Ridge (JdFR) in the northeast Pacific, aims to find pattern in this variation and to propose viable hypotheses of the mechanisms driving the patterns. Axial is an ideal location as it supports mature vent fields (venting for over 15 years) and young, developing vents initiated by a volcanic eruption in 1998. Thus, I was able to study both temporal and spatial variation in vent communities at the same site and relate patterns of developing assemblages to patterns observed at longer-lived vents. Pattern detection is the first critical step in any community ecology study as it justifies and focuses the search for process. I have refined existing statistical methods and developed novel techniques to test for pattern in vent species distributions and abundances. I modified an existing null model approach and showed that species distributions among sixteen vents differ from random in a long-lived (>15 years) vent field. I also developed a novel null model to confirm that initial patterns of community assembly seven months following the Axial eruption differ from random recruitment of species and individuals to new vents. My description of the community response to the Axial eruption is the first quantitative report of patterns of vent colonization and succession. My work documents that new vents are colonized quickly (within months) and that initial assemblages are variable. However, rapid community transitions and species replacements within the first few years cause new assemblages to resemble mature vents by 2.5 years post-eruption. Three habitat factors correlate with the development of nascent vent assemblages: the recruitment timing of the tubeworm Ridgeia piscesae post-eruption, vent age and vent fluid hydrogen sulphide content. I also describe a new polynoid polychaete discovered colonizing the new vents in high densities. My major contribution to vent community ecology is revealing species patterns through extensive sampling and rigorous statistical methods. These patterns are a necessary step towards understanding the processes that structure vent communities: they direct future research effort towards the key species and generate hypotheses to be experimentally tested. My work also elucidates how vent species respond to habitat destruction and creation, which is critical information for effectively managing Canada's only hydrothermal vent Marine Protected Area on the JdFR. / Graduate

This work seeks to explore current ecological theory through application to communities inhabiting hydrothermal vents. This thesis aims to: (1) add to and synthesise knowledge of species and their distributions at the intra- and intersegment scale; and (2) evaluate vent community patterns and speculate on processes. Samples used are submersible grabs of low temperature (<60°C) tubeworm assemblages on basalt and sulphide surfaces. Species abundances and distributions on three segments of the Juan de Fuca Ridge (Axial, Cleft, and CoAxial) are described. Community descriptors such as species density, Simpson's and the Shannon-Wiener diversity indices, evenness, species richness, species abundance-distribution models, species percent-average relative abundance and density are used. Vent community structure is compared among segments using these descriptors, visual descriptions, pairwise correlations, Friedman tests of distributions, cluster and correspondence analysis, rarefaction, complementarity, a test for saturation, and Whittaker's beta diversity. Vent community composition on Axial, north Cleft, and CoAxial is similar at the segment and inter-segment scale. The limpet Lepetodrilus fucensis is the most abundant species at all sites. Differences among communities are best seen temporally, not spatially. Senescent communities can be distinguished from active vent assemblages. Pioneer communities, however, are statistically indistinguishable from intermediate communities when sampled two or more years post-eruption. Axial and Cleft species dispersion fits the core-satellite hypothesis. The exceptions are the polynoids Branchinotogluma sp., Lepidonotopodium piscesae, and Levensteiniella kincaidi, which are widespread and present in low local abundances. Both local and mesoscale regional mechanisms explain observed local diversity. Spatial isolation, not habitat differences, influences between-habitat diversity (beta diversity) on Axial, Cleft, and all three segments combined. Meiofauna are important for species richness estimates, identifying differences among structurally similar communities, and understanding input/output between vents and the deep-sea. Measurements such as species richness and diversity indices may be poor at distinguishing among vent communities because vents are species poor and uneven. The Michaelis-Menten, Jackknife 2, and Chao 2 nonparametric vent species richness estimators perform well with small samples. Vent communities should be compared to habitats of similar diversity and evenness as well as disturbance and productivity regimes. Candidate comparison communities include communities in early successional states, selected taxocenes such as carabid beetles on fungi, or high disturbance and/or low diversity systems like the rocky intertidal, organically polluted sediments and oxygen minimum zones below upwelling regions in the deep-sea. / Graduate

abstract: Olympus Mons is the largest volcano on Mars. Previous studies have focused on large scale features on Olympus Mons, such as the basal escarpment, summit caldera complex and aureole deposits. My objective was to identify and characterize previously unrecognized and unmapped small scale features to understand the volcanotectonic evolution of this enormous volcano. For this study I investigated flank vents and arcuate graben. Flank vents are a common feature on composite volcanoes on Earth. They provide information on the volatile content of magmas, the propagation of magma in the subsurface and the tectonic stresses acting on the volcano. Graben are found at a variety of scales in close proximity to Martian volcanoes. They can indicate flexure of the lithosphere in response to the load of the volcano or gravitation spreading of the edifice. Using Context Camera (CTX), High Resolution Imaging Science Experiment (HiRISE), Thermal Emission Imaging System (THEMIS), High Resolution Stereo Camera Digital Terrain Model (HRSC DTM) and Mars Orbiter Laser Altimeter (MOLA) data, I have identified and characterized the morphology and distribution of 60 flank vents and 84 arcuate graben on Olympus Mons. Based on the observed vent morphologies, I conclude that effusive eruptions have dominated on Olympus Mons in the Late Amazonian, with flank vents playing a limited role. The spatial distribution of flank vents suggests shallow source depths and radial dike propagation. Arcuate graben, not previously observed in lower resolution datasets, occur on the lower flanks of Olympus Mons and indicate a recent extensional state of stress. Based on spatial and superposition relationships, I have constructed a developmental sequence for the construction of Olympus Mons: 1) Construction of the shield via effusive lava flows.; 2) Formation of the near summit thrust faults (flank terraces); 3) Flank failure leading to scarp formation and aureole deposition; 4) Late Amazonian effusive resurfacing and formation of flank vents; 5) Subsidence of the caldera, waning volcanism and graben formation. This volcanotectonic evolution closely resembles that proposed on Ascraeus Mons. Extensional tectonism may continue to affect the lower flanks of Olympus Mons today. / Dissertation/Thesis / Masters Thesis Geological Sciences 2015

A series of physical models consisting of three different diameter pipes at the same 4% slope were studied at the Utah Water Research Laboratory (UWRL). Various combinations of air flow and head on the pipe were used to determine the effect of pipe diameter, head, and air flow on the behavior of air bubbles introduced into the pipes and to determine the venting capacity of the pipes. It was determined that neither bubble velocity nor bubble length changes with pipe diameter or head changes within the range tested. It was also determined that bubble velocity and length will increase with increased air flow. Bubble velocity also increased with increasing bubble length consistent with prior research. Overall the venting capacity of a pipe is dependent upon having a large enough pipe to prevent slug flow. A procedure was developed to aid in the sizing of submerged vent piping during the design of pipelines based on the data collected during this study and utilizing prior research.

Ce projet a pour but d'examiner l'influence à grande échelle des vents en provenance des noyaux actifs de galaxies (NAG) sur l'enrichissement en métaux du milieu intergalactique. Un modèle semi-analytique de vents de NAG anisotropes est implémenté dans des simulations à N-corps suivant un modèle cosmologique ACDM. Les NAG évoluent dans une boîte de dimension (128 h-1Mpc)3, à partir d'un décalage vers le rouge initial z = 24 jusqu'à un décalage vers le rouge final de z = 0, en utilisant une fonction de luminosité observationnelle des quasars afin d'obtenir leur distribution en décalage vers le rouge et en luminosité, et un modèle analytique pour décrire l'expansion de leurs vents. Le volume enrichi en métaux par les vents anisotropes de la population cosmologique de NAG est calculé et analysé au cours du temps de Hubble. Le volume enrichi obtenu est relativement faible à z > 2.5 et croît rapidement par la suite jusqu'à z = 0. Les vents de NAG enrichissent une fraction de 0.65 - 1.0 du volume cosmologique total à l'époque actuelle, pour différentes valeurs des paramètres de notre modèle. Les vents plus anisotropes enrichissent préférentiellement les régions de faible densité, et ce, de façon plus accentuée à haut décalage vers le rouge et moindre à faible décalage vers le rouge.

Critical properties of the NaCI-H₂0 fluid system are of fundamental interest to a variety of geochemical applications including fluid inclusion studies, numerical modeling of hydrothermal systems, and development of theoretical models for two·component fluid systems. Although many workers have expressed interest in NaCl·H₂0 fluid critical properties, most studies have been limited to small compositional ranges with little agreement among data sets at higher salinities. Critical densities are recorded in only one of these reports, and no studies have determined the locations of NaCl-H₂0 critical isochores (PT projections of critical densities). Furthermore, no studies to date have determined critical properties of NaCl·H₂0 solutions in excess of room temperature saturation (26.4 wt.% NaCl). / Master of Science

Prokaryotic microorganisms, which are at the base of deep-sea hydrothermal vent food webs, adapt rapidly to environmental fluctuations. This study aimed at comparing bacterial communities in contrasting hydrothermal habitats to better understand compositional adaptations to local conditions. I first used small subunit (SSU) ribosomal RNA (rRNA) gene sequences to compare mat-forming bacterial communities associated with iron oxides at two hydrothermal vent sites on the Tonga Arc, southwest Pacific. Operational taxonomic units (OTUs), defined at 97% sequence similarity, were affiliated to a great diversity of autotrophic and heterotrophic groups. Metabolically diverse Gammaproteobacteria dominated the sample from Volcano 19, collected at 992 m depth. The sample from Volcano 1, collected at 197 m depth, was dominated by iron-oxidizing bacteria from the class Zetaproteobacteria. The depth of the sampling sites was proposed to explain clone library dissimilarities. In the following studies, I compared bacterial communities associated with the vestimentiferan tubeworm Ridgeia piscesae, a foundation species at the Juan de Fuca Ridge. Samples of the polychaete were collected from tubeworm habitats in contrasting flow regimes that influenced temperature and hydrogen sulphide concentrations. Free-living bacteria were analyzed using both sequencing and 454 pyrosequencing of the SSU rRNA gene. Statistical analyses suggested a predictable pattern of bacterial community composition for the two habitats, with higher proportions of sulphur and hydrogen oxidizers in High Flow and more heterotrophic groups in Low Flow environments. Temperature, available energy for metabolism, and stability of the habitat were suggested to explain these distinctive bacterial communities. Symbiotic assemblages were investigated using the same sequencing methods together with catalyzed reporter deposition-fluorescence in situ hybridization (CARD-FISH). Gammaproteobacteria dominated all sequence libraries, followed by Epsilonproteobacteria. CARD-FISH confirmed the co-occurrence of these groups within R. piscesae trophosomes. Statistical analyses indicated distinctive membership and structure of trophosome assemblages between sampling sites. Analysis of R. piscesae juvenile showed distinctive structural properties when compared to adult individuals, but similar membership, within sampling sites. These results suggested that the composition of trophosome assemblages might be affected by specific physical and chemical conditions at each vent site and that a selection process might occur during R. piscesae’s development. / Graduate / 0410 / 0416 / 0329 / nathalieforget@gmail.com

Ventilation might seem like a minor consideration, but when properly installed, it can extend the life of the attic and roof structure of residential buildings saving huge dollar amounts in repair costs. The objectives of this study were to measure the wind effect on different kind of vents as the problem is vents are subjected to high wind loads during hurricane events and often fail causing water penetration and damage of interior contents. Wind-induced failure of rooftop vents during a hurricane may result in large openings in the roof that will allow water to penetrate into the building, a puncturing of the roof membrane, again allowing water infiltration, and detached rooftop vents can pose considerable threats as windborne debris. The study found that water infiltration through a vent system is dependent upon the differential pressure as well as the vent mechanism. For vents experiencing higher differential pressures, vent covers can be used during storms to reduce water infiltration. Active controls can also be designed to close the vents automatically as differential pressure increases based on the wind speed and wind angle of attack.

Vestimentiferan tubeworms of the eastern Pacific Ocean are often keystone species in vent communities. These polychaetes are host to intracellular Gammaproteobacteria symbionts. In this association, the siboglinid worms supply their symbionts with the compounds necessary to chemosynthesis while the sulfide oxidizing bacteria provide their host with the organic molecules necessary for their metabolism. The adult worms lack a digestive system and are therefore completely dependent on their symbionts for their nutrition. Given the obligate nature of the association for the host, it is surprising that the symbionts are not transmitted from parents to offspring but are acquired de novo from the environment at each generation. In other known cases of horizontally acquired mutualism (e.g. Rhizobium-legumes, dinoflagellates- corals), obtaining symbionts from the environment benefit the hosts by allowing for a degree of partner choice. According to the partner choice hypothesis, tubeworms that associate with the best-adapted partner(s) to a specific range of habitat conditions are in turn better adapted to this environment. Of course, this hypothesis assumes that there is diversity within the symbiotic partners. Phylogenetic analyses on the other hand seemed to indicate that nearly all species of vent tubeworms of the eastern Pacific were associated with the same species of symbionts: Candidatus Endoriftia persephone. However, these studies focussed on a few molecular markers. In this thesis, I used in situ hybridization and next generation sequencing to characterize the symbiont diversity at the species and strain level, as well as within individual hosts and across host species. I found that the intra-host symbiont populations are likely composed of multiple strains or lineages of the same bacterial species, that the symbiont populations separated by mid-ocean ridge discontinuities are vicariant, and that other factors such as local environmental conditions or host specificity might participate in shaping the genetic make-up of these populations. / Graduate / 0410 / 0307 / 0715 / 0306 / 0416 / 0329

Les radars à synthèse d'ouverture (SAR) spatioportés ont fait la preuve de leur valeur inestimable dans l'observation des vents et des vagues océaniques mondiaux. Les images SAR acquises par plusieurs capteurs sont utilisées, notamment Sentinel-1 (S-1), Envisat/ASAR, Gaofen-3 et Radarsat-2. Cette thèse passe en revue les paramètres SAR couramment utilisés dans la première partie. Une série d'étapes d'étalonnage sont nécessaires pour obtenir un NRCS approprié et une évaluation du NRCS est effectuée pour le mode d'onde S-1 (WV). Il s'avère que WV est mal calibré et est donc recalibré pour obtenir un NRCS précis. Il a été démontré que la coupure de l'azimut est complémentaire du NRCS et peut expliquer l'impact de l'état de la mer sur l'extraction du vent. D'après les produits SAR entièrement polarimétriques disponibles, la coupure de l'azimut varie considérablement en fonction des polarisations. La transformation actuelle de la cartographie SAR est suffisante pour interpréter la coupure azimutale copolarisée, mais pas pour la polarisation croisée. Compte tenu des limites de l'imagerie SAR, un nouveau paramètre est proposé et défini en fonction du spectre croisé de l'image SAR, appelé MACS. La partie imaginaire de MACS est une quantité signée par rapport à la direction du vent. Compte tenu de cette dépendance, on s'attend à ce qu'un algorithme indépendant de récupération du vent en bénéficie. L'ampleur du MACS peut aider à estimer la fonction de modulation de la cartographie SAR. De plus, la MACS donne également des résultats prometteurs en ce qui concerne les études globales sur les vagues. Les signatures globales du MACS à différentes longueurs d'onde sont bien représentatives de la distribution spatiale et saisonnière des vents. Les MACS des vagues longues montrent des valeurs plus élevées sur les trajectoires des tempêtes alors que les vagues plus courtes sont principalement dans les vents des trader. Ces résultats devraient aider à évaluer les résultats du modèle et compléter les études ultérieures sur le climat spectral global des vagues. / Spaceborne synthetic aperture radar (SAR) has been demonstrated invaluable in observing the global ocean winds and waves. SAR images acquired by multiple sensors are employed, including Sentinel-1(S-1), Envisat/ASAR, Gaofen-3 and Radarsat-2. This thesis reviews the commonly used SAR parameters (NRCS and azimuth cutoff) in the first part. A series of calibration steps are required to obtain a proper NRCS and assessment of NRCS is carried out for S-1wave mode (WV). It turns out that WV is poorly calibrated and is thus re-calibrated to obtain accurate NRCS. Azimuth cut off is demonstrated to be complementary to NRCS and can account for the sea state impact on the wind retrieval. Based on the available fully polarimetric SAR products, azimuth cut off is found to vary greatly with polarizations. The present SAR mapping transformation is sufficient to interpret the co-polarized azimuth cut off, while not for the cross-polarization. With the limitations of SAR imaging in mind, a new parameter is proposed and defined based on the SAR image cross-spectra, termed as MACS. The imaginary part of MACS is found to be a signed quantity relative to the wind direction. Given this dependence, an independent wind retrieval algorithm is expected to benefit. The magnitude of MACS is able to aid for estimate of modulation function of SAR mapping. In addition, MACS also gives promising results regarding the global wave studies. The global signatures of MACS at various wave lengths are well representative of the winds distributions, spatially and seasonally. MACS of long waves shows greater values over the storm tracks while the shorter waves are mostly within the trader winds. These results are expected to help evaluate the model outputs and complement further studies of the global wave spectral climate. Data continuity in the coming 10 years shall extend the study towards longer duration.