The renewable natural resources (RNR) of Senegal form the basis of socioeconomic development for the population of this Sahelian country of West Africa. However, because of the current maladapted policies and practices, these resources are in a crisis that has led to a major degradation of the socioeconomic and environmental conditions. To solve these socioeconomic and environmental problems, many solutions have been developed and implemented. However, because of their disregard for both socioeconomic and traditional cultural values, these attempted solutions have often contributed to the problems. To resolve this crisis, Senegal needs to adopt a new integrated management approach along with new integrated natural resources tenure and decentralization policies. Considering the country's limited economic means, the urgent need for policy change, and the necessary progressive implementation of a new policy, this new natural resources policy will need to be developed incrementally. It should take into account the existing policies and examine different alternatives from the current ecological, economic, and sociocultural situation.

In desert grasslands of the southwestern United States, Prosopis velutina (mesquite), an N-fixing legume, has proliferated from historic drainage locations into more xeric grassland plains. This expansion is forming a more heterogenous soil nutrient topography in grasslands, N-pools are becoming localized under mesquite canopies, yet the rate and extent of this sequestration remains relatively unknown. Repeated prescribed burning has been used to control Prosopis distribution, but effects of fires on grassland soil nutrient distribution and aboveground plant biomass are also largely unknown. I examined recent research concerning P. velutina natural history, emphasizing characteristics that contribute to range expansion. I also evaluated Prosopis management practices---which include herbicide treatment, prescribed burning, grazing reduction, and mechanical removal---and management goals---which involve complete removal, no removal, and limited removal. Of these, limited removal is the most beneficial, using an herbicide application followed by periodic prescribed burning. In 1997 I established a study area at Fort Huachuca Military Reservation in southeastern Arizona, selecting two adjacent sites with similar soil composition and topography but different fire histories. I examined spatial and seasonal changes in composition and distribution of available soil N and litterfall. My results indicated these were more spatially and temporally heterogenous on sites with low fire frequency and high P. velutina stand development. In 1998 I selected nine sites at Fort Huachuca on two upland surfaces located < 1 km apart, with similar soil physical characteristics and fire frequencies ranging from 0 to 5 fires/decade. I evaluated relationships between fire frequency, soil nutrient status (pH, available P, organic C, total N, and available N), and aboveground plant biomass, including that of the non-native Eragrostis lehmanniana (Lehmann lovegrass). Soil pH and ammonium significantly decreased with increased fire frequency on one surface, and available P significantly decreased with increased fire frequency on the other surface. Available P and pH were significantly different between the 2 surfaces, but aboveground biomass was similar. Soil nutrient status and biomass were not related, suggesting plant-available soil nutrients may not control plant distribution or recovery following fire. E. lehmanniana biomass was negatively correlated with native grass and forb biomass, and tended to increase with increasing fire frequency. Surface litter and E. lehmanniana biomass were correlated, and may increase fire frequency, an important consideration when implementing grassland fire management practices.

The effects of land-use history in the Animas Mountains, New Mexico (AM) and the Sierra los Ajos, Sonora (SLA) were studied in relation to morphological and soil chemical characteristics, radial and basal area growth, and forest structure. Litter depth, organic matter, total nitrogen, CEC, and exchangeable cations were greater in the AM than in the SLA, apparently as a result of differences in fire frequency and other land uses. Seasonal precipitation (October-January) was reconstructed for the AM. Annual precipitation (July-July) was reconstructed for the SLA. July PDSI was reconstructed for both mountain ranges. Fires in the AM were preceded by relatively wet conditions two years before the fire year. No significant climate-fire relationship was found in the SLA. Tree radial and basal area increase differed between mountain ranges, suggesting that annual growth was influenced by differences in land-use history. Fire suppression activities and other land uses in the AM apparently have produced an increase in tree density and dominance of shade-tolerant but less-fire resistant species (i.e. Douglas-fir, southwestern white pine, pinyon pine). More frequent fires and logging activities in the SLA apparently have produced lower tree densities and dominance by shade-intolerant ponderosa pine.

Master of Science / Department of Civil Engineering / David R. Steward / Konza Prairie is one of the few areas in the United States were natural landscape of the area is still intact. Human action on changing the landscapes in this area is limited and much of the land remains as native grassland. In spite of its natural existence, this area is not completely isolated from the rest of the world. Changes that are taking place in climate will eventually have the same effect to this region as well as other human populated areas. Increase in carbon concentration in the air has resulted to increase in temperature, this increase in temperature increases the evaporation from the sea, oceans and the ice capes. As the atmospheric water vapor changes the precipitation pattern also change. Changes in precipitation due to climate change will result to change in hydrology and hydraulics of the streams and groundwater flow regime. Precipitation provides surface runoff and groundwater infiltration, which recharge the cracked limestone aquifer present in the Konza area. The infiltration water moves trough the cracked rocks and eventually reach the creeks such as Kings Creek and flow to the Kansas River. Increase in precipitation will result to increase in surface runoffs and more groundwater recharge. Decrease in precipitation will result to decrease in both surface and groundwater. To examine changes in groundwater elevation as recharge change in Konza, a groundwater model was developed based on erosion impact calculator (EPIC) ecological model and SLIT groundwater model. EPIC model estimates the deep percolation (recharge) as 12% and total runoff to about 24% of the annual average precipitation. The annual average recharge values from EPIC were used in SPLIT to simulate results for the groundwater elevation at Konza prairie. Field wells elevation were use to calibrate the SPLIT results. By estimating the hydraulic permeability value to 0.546m/d the field well measurements and SPLIT simulated groundwater elevation results provide a good match. After calibration max and min recharge together with a 5-years moving average were used to examine the changes in groundwater elevation as recharge changes. Future study intends to use the calibrated Konza groundwater model and the forecasted climate data to simulate result for groundwater elevation as climate changes.

Doctor of Philosophy / Department of Agronomy / Jay M. Ham / Land management practices such as burning and grazing may affect evapotranspiration (ET) and water balance of the tall grass prairie in the Flint Hills of eastern Kansas. Experiments were designed to estimate or measure the water balance of a stock-watering pond, and compare energy balance parameters and ET between grazed and ungrazed prairies. The hydrology of the native tallgrass prairie also was compared with mature stand of eastern red cedar (Juniperus virginiana), a site that was formerly prairie but converted to woodland when prescribed burning was discontinued. Data were collected to encompass the seasonal and yearly changes in weather variability. A host of micrometeorological sensors were used to measure surface atmosphere exchange and water losses, including: eddy covariance towers on prairie and woodland sites, specialized through fall and stem flow equipment at the woodland site, and an instrumentation raft at the stock watering pond. Results of the stock pond study showed that, on average, evaporation accounted for 64% of the water loss, followed by seepage at 31%, cattle use at 3% and transpiration at 2%. Comparisons of grazed and ungrazed areas showed that grazing caused only small, 3 to 6 %, reductions in seasonal ET compared with ungrazed pastures despite large differences in vegetative cover. In the woodland study, the 50-yr-old cedar canopy intercepted 54% of the precipitation received, thus decreasing the amount of precipitation reaching the soil. Evapotranspiration from woodland and prairie sites were similar, but net carbon exchange was greater on the prairie. Thus, the apparent water use efficiency during he summer months was 3 times greater on the prairie. Net radiation at the woodland site was 100 W m-2 higher compared with the prairie. This caused an increase in the woodland sensible heat flux and midday Bowen ratios, yet woodland latent heat flux and ET was similar to the prairie during the study, factors that could affect regional climate. Land management decisions regarding grazing, prescribed burning, and stock pond design will impact the watershed hydrology and productivity of the tallgrass prairie.

My thesis focuses on different ranch management practices and the effects they may have on bird communities in northern Texas. Specifically, I address the potential differences between ranches enrolled in the Audubon Conservation Ranching program versus ranches that are not enrolled. Each research chapter is meant to be an individual publication. Chapter 2 compares and contrasts two different ways to survey bird communities in prairie systems. Chapter 3 compares ranch management techniques, prairie health indicators, and bird communities. This study aims to observe any mechanistic drivers of rangeland health that may elicit a positive bird assemblage response.

Master of Science / Department of Agronomy / Walter H. Fick / Rough-leaved dogwood (Cornus Drummondii Meyer) is one of the most invasive woody plants in rangeland of Kansas. Reduced prescribed burning due to drought and urban encroachment probably has contributed to its spread. Herbicides are commonly recommended for control of rough-leaved dogwood, but minimal data exists for recommendation development. Ten herbicide treatments were applied in June during late flowering at two locations in northeast Kansas in 2005 and 2006. Each treatment was replicated three times in a randomized complete block design with individual plot sizes of 3 x 3 m. Herbicides were applied with hand sprayers in 1017 L/ha solution. Visual evaluations of defoliation were made about 1 and 12 months after treatment (MAT) and mortality was estimated about 1 year after treatment (YAT). Defoliation 1 MAT varied among herbicides with significant location by year and herbicide by year interactions. Treatments providing greater than 70% defoliation 12 MAT both years were triclopyr + 2,4-D (1.12 + 1.06 kg ae/ha), triclopyr + fluroxypyr (1.83 + 0.61 kg ae/ha), picloram + fluroxypyr (0.41 + 0.41 and 0.82 + 0.82 kg ae/ha), and picloram + 2,4-D + triclopyr (0.66 + 2.44 + 2.44 kg ae/ha). Rough-leaved dogwood is difficult to control with a single herbicide application, but treatments exist that will substantially reduce stands. Tebuthiuron pellets (Spike 20P) are another control measure recommended for rough-leaved dogwood. Treatments of 4.4 kg ai/ha (3/4 oz per 100 square feet) tebuthiuron pellets were applied in December 2004. A visual estimate of control indicated tebuthiuron reduced dogwood cover by 65% compared to a 3% decrease on untreated plots. Dogwood density was reduced by 2.2 stems/m2 (P<0.08). Total woody plant cover increased on untreated plots by 6.2 percentage units, but was decreased by 20.9 percentage units on tebuthiuron treated plots. Other woody plants decreased in both treated and untreated plots. Shading by a large elm tree likely caused variation between replications including increases in cover and density of rough-leaved dogwood on treated plots. Tebuthiuron is a photosynthetic inhibitor that often is not effective on shaded plants. Tebuthiuron pellets applied at 4.4 kg ai/ha was an effective control option for unshaded rough-leaved dogwood.

Master of Science / Department of Agronomy / Walter H. Fick / Patch-burn grazing is a relatively new concept in terms of rangeland management. While numerous benefits have been associated with this system, in the tallgrass prairie of Kansas, cattle production and sustainability of rangeland are critical. In 2006, 253 ha at the KSU Bressner Range Research Unit in Woodson County, Kansas were subjected to spring patch-burn grazing (using one-third portions) and traditional full-burn grazing. Each treatment within the split-block design was replicated four times for 3 years. The objectives were to evaluate whether livestock performance would be compromised under this grazing system, to monitor the health of the rangeland, and to observe the usefulness of this tool as a potential control of the invasive plant sericea lespedeza [Lespedeza cuneata (Dumont) G. Don]. In regards to cattle performance, burn treatments had no significant difference in average daily gain (p≥0.10) in any of the 3 years. On average, cattle utilized 61% of the current year’s forage production in patch-burned portions, which was higher (p≤0.10) than that of unburned (30%) and full-burn (41%) portions. Results of the botanical composition show forb and woody plant composition did not differ between treatments, however differences (p≤0.10) were present in grass composition. Total annual grasses increased 19.1 percentage units under patch-burn and 2.1 units under full-burn, while total perennial grasses decreased 18.4 and 1.1 units, respectively. When evaluated by treatment area (one-third portion), results indicated that the 3-year cycle did allow enough time for recovery. At 2 years after treatment (2-YAT), no significant difference in composition(p≥0.10) was present between initial patch-burn portions and the full-burn pastures. Finally, in only 1 year of the study did cattle statistically consume a greater percentage of sericea lespedeza plants (p≤0.10) in the patch-burned portions (92%) than in full-burned pastures (35%). Biomass utilization did not differ (p≥0.10) between burn treatments. Surprisingly, there was a trend for the number of plants in the sampled areas of the patch-burn portions to decrease throughout the cycle. However, at the conclusion of the 3-year cycle, sericea densities did not differ(p≥0.10) between treatments. Patch-burn grazing shows promise as a potential management tool for Kansas land managers.

Doctor of Philosophy / Department of Agronomy / Clenton E. Owensby / Most rangelands are harvested at some point during the year and removal of plant leaf area and biomass alters a host of ecosystem processes including gas exchange. An experiment was conducted in 2005 and 2006 to study the effects of clipping tallgrass prairie at different dates on water vapor and CO2 fluxes. A portable, non-steady-state chamber was designed to measure CO2 and water vapor fluxes from small plots in less than 40 s. A combination of sunlit and shaded readings allowed measurements of net carbon exchange (NCE) and ecosystem respiration (RE); by summing NCE and RE, gross canopy photosynthesis (GCP) was calculated. Throughout the two-year study, the chamber had a minimal effect on microclimate, i.e., average chamber temperature increased 2.9° C, while chamber pressure increased only 0.3 Pa during measurements, and photosynthetically active radiation attenuation was 10%. The immediate effect of all clipping treatments was a loss of leaf area that led to reductions in GCP, NCE, and RE and in most cases decreased water vapor flux. Further patterns of carbon flux were governed by the amount of water stress during canopy development, while water vapor flux rates varied with water availability. Canopies that developed during periods of low water stress quickly increased carbon flux rates following precipitation after a mid-season drought. However, flux rates of canopies, which developed during the mid-season drought, responded considerably slower to subsequent water availability. A separate experiment was conducted from June-October of 2006 to estimate GCP, leaf area index (LAI), and total aboveground biomass with a hyperspectral radiometer. Indices such as the Normalized Difference Vegetation Index and the Simple Ratio were used to estimate LAI and biomass had poor correlations with measured values. However, GCP was significantly correlated to all six indices derived in this study. While GCP measured from June-October was significantly correlated with all indices, removal of the senesced canopy scans recorded during October greatly increased the relationship.

Doctor of Philosophy / Department of Biology / Kimberly A. With / The loss, fragmentation, and degradation of grasslands have resulted in widespread declines in grassland birds. Nest predation is the leading cause of avian reproductive failure; therefore minimizing nest predation can lessen the severity of bird declines. Snakes are important predators of bird nests, but little is known about how snakes may enhance predation risk. To address this issue, I studied the habitat use, movement behavior, population genetic structure, and connectivity of snakes in the grasslands of northeastern Kansas. I addressed the connectivity of eastern yellowbelly racer (Coluber constrictor flaviventris) populations by using a landscape genetics approach at a broad scale (13,500 km2). I also radio-tracked the yellowbelly racer and Great Plains ratsnake (Pantherophis emoryi) at Konza Prairie Biological Station to understand their spatial ecology while simultaneously evaluating nest survival in grassland birds. Individual racers had limited dispersal (<3 km), but substantial admixture occurred within 30 km and populations were in migration-drift equilibrium and had high allelic diversity; therefore, racers must be abundant and continuously distributed for gene flow to be fluid throughout the region. Racers may be more likely to encounter bird nests, as they had more frequent movements and traversed greater distances on average than ratsnakes, which exhibited long periods of inactivity between directed movements. As for grassland birds, nest survival rates decreased with increasing shrubs and decreasing vegetation height. Discriminant function analysis revealed that successful nests were likely to occur in tall vegetation but reduced shrub cover, whereas higher shrub cover characterized snake habitats. Because snakes often use shrubs, nests in areas of increased shrubs may be at higher risk of predation by snakes. Targeted removal of shrubs may increase nest success by minimizing the activity of predators attracted to shrubs. Although predator removal is often a strategy for protecting bird populations, it may not be feasible in this instance, especially since snakes are a native component of the grassland community. Efforts to reduce snake predation on grassland bird nests should therefore focus on managing habitat within grasslands (i.e., shrubs) that influence snake activity, as no natural or anthropogenic habitat barriers currently limit snake movement across the landscape.