Master of Science / Department of Agronomy / Walter H. Fick / Kevin Price / Caucasian bluestem [Bothriochloa bladhii (Retz) S.T. Blake] is a perennial, C4 warm-season bunchgrass that was first introduced in 1929 from Russia as a potential forage crop in the Great Plains. Due to its invasiveness and tolerance of drought and grazing pressure, Caucasian bluestem can out-compete native prairie species. Research has shown that this species, when compared to native tallgrass species in the Flint Hills of Kansas causes decreased cattle weight gains because of its poor forage quality relative to tallgrass prairie species. Traditional methods of plant data measurements and mapping are costly and time consuming. Use of remotely sensed data to map and monitor the distribution and spread of this plant would be most useful in the control of this aggressive invader. Spectroradiometer data were collected over the 2009 growing season to determine if and when Caucasian bluestem was spectrally unique from native tallgrass prairie species. Observations were made from June through September as the plants were going into a senescent state. Reflectance data were measured approximately every two weeks or when clear/near clear sky conditions prevailed. Statistical analyses for differences in spectral characteristics were conducted to determine the optimal spectral bands, indices and timing for discriminating Caucasian bluestem from native tallgrass species. Difference in reflectance for spectral reflectance of bands 760 nm, 940 nm, 1,070 nm, and 1,186 nm were found to be statistically significant on the June 17th and June 30th sampling dates. The following band ratios and indices were found to be significantly different between Caucasian bluestem and native range on the June 17th collection date: Simple Ratio, Modified Normalized Difference Index, Normalized Phaeophytinization Index, Plant Index 1, Normalized Water Difference Index, Water Band Index, Normalized Difference Nitrogen Index, and the Normalized Difference Lignin Index. Findings of this study suggest that Caucasian bluestem can be spectrally discriminated from native tallgrass prairies of the Flint Hills in Kansas if the measurements are collected in mid to late June. Statistical analyses also showed differences between treatments for percent litter, grass, and forb basal cover.

Master of Science / Department of Animal Sciences and Industry / Dale A. Blasi / An experiment was conducted to evaluate the effects of dry matter intake (DMI) restriction on early receiving performance by steers in a drylot and subsequent grazing performance, feedlot performance, and carcass characteristics. During the backgrounding period, crossbred, weanling steers (n = 329; initial BW = 191± 5.52 kg ) were randomly assigned to 1 of 4 DMI levels corresponding to ad libitum, 2.50% of BW (2.50%), 2.25% of BW (2.25%), and 2.00% of BW (2.00%) for 62 d. During the subsequent grazing period, the same steers were randomly assigned to 13 paddocks to graze for 90 d. Paddocks were stocked at 281 kg live weight per hectare. Initial steer BW were similar on each pasture and each backgrounding treatment was equally represented within a paddock. During the feedlot period, steers were finished at a commercial feedlot and were assigned to 1 of 4 pens according to their rank in BW. Entire pens were harvested when average steer BW reached 545 kg. During the backgrounding period, ad libitum-fed steers had greater (P < 0.001) ADG and final BW than other treatments; steers fed at 2.50 and 2.25% of BW had similar ADG and final BW and were greater (P < 0.001) than steers fed 2.00% of BW. During the grazing period, compensatory gain was observed in restricted DMI treatments. Steers fed at 2.00% of BW had greater (P = 0.006) ADG than ad libitum-fed steers but an ADG similar to that of the other restricted DMI treatments. Steers fed ad libitum, 2.50% of BW, and 2.25% of BW had similar final BW and steers fed 2.00% of BW had lesser (P < 0.001) final BW than other treatments. During the feedlot phase, steers fed 2.00% of BW were on feed longer (P < 0.05) than other treatments. Growth compensation during grazing illustrated that restricted feeding immediately prior to pasture grazing can reduce backgrounding costs.

Master of Science / Department of Civil Engineering / David R. Steward / Konza Prairie is one of the few areas in the United States were natural landscape of the area is still intact. Human action on changing the landscapes in this area is limited and much of the land remains as native grassland. In spite of its natural existence, this area is not completely isolated from the rest of the world. Changes that are taking place in climate will eventually have the same effect to this region as well as other human populated areas. Increase in carbon concentration in the air has resulted to increase in temperature, this increase in temperature increases the evaporation from the sea, oceans and the ice capes. As the atmospheric water vapor changes the precipitation pattern also change. Changes in precipitation due to climate change will result to change in hydrology and hydraulics of the streams and groundwater flow regime. Precipitation provides surface runoff and groundwater infiltration, which recharge the cracked limestone aquifer present in the Konza area. The infiltration water moves trough the cracked rocks and eventually reach the creeks such as Kings Creek and flow to the Kansas River. Increase in precipitation will result to increase in surface runoffs and more groundwater recharge. Decrease in precipitation will result to decrease in both surface and groundwater. To examine changes in groundwater elevation as recharge change in Konza, a groundwater model was developed based on erosion impact calculator (EPIC) ecological model and SLIT groundwater model. EPIC model estimates the deep percolation (recharge) as 12% and total runoff to about 24% of the annual average precipitation. The annual average recharge values from EPIC were used in SPLIT to simulate results for the groundwater elevation at Konza prairie. Field wells elevation were use to calibrate the SPLIT results. By estimating the hydraulic permeability value to 0.546m/d the field well measurements and SPLIT simulated groundwater elevation results provide a good match. After calibration max and min recharge together with a 5-years moving average were used to examine the changes in groundwater elevation as recharge changes. Future study intends to use the calibrated Konza groundwater model and the forecasted climate data to simulate result for groundwater elevation as climate changes.

Doctor of Philosophy / Department of Agronomy / Jay M. Ham / Land management practices such as burning and grazing may affect evapotranspiration (ET) and water balance of the tall grass prairie in the Flint Hills of eastern Kansas. Experiments were designed to estimate or measure the water balance of a stock-watering pond, and compare energy balance parameters and ET between grazed and ungrazed prairies. The hydrology of the native tallgrass prairie also was compared with mature stand of eastern red cedar (Juniperus virginiana), a site that was formerly prairie but converted to woodland when prescribed burning was discontinued. Data were collected to encompass the seasonal and yearly changes in weather variability. A host of micrometeorological sensors were used to measure surface atmosphere exchange and water losses, including: eddy covariance towers on prairie and woodland sites, specialized through fall and stem flow equipment at the woodland site, and an instrumentation raft at the stock watering pond. Results of the stock pond study showed that, on average, evaporation accounted for 64% of the water loss, followed by seepage at 31%, cattle use at 3% and transpiration at 2%. Comparisons of grazed and ungrazed areas showed that grazing caused only small, 3 to 6 %, reductions in seasonal ET compared with ungrazed pastures despite large differences in vegetative cover. In the woodland study, the 50-yr-old cedar canopy intercepted 54% of the precipitation received, thus decreasing the amount of precipitation reaching the soil. Evapotranspiration from woodland and prairie sites were similar, but net carbon exchange was greater on the prairie. Thus, the apparent water use efficiency during he summer months was 3 times greater on the prairie. Net radiation at the woodland site was 100 W m-2 higher compared with the prairie. This caused an increase in the woodland sensible heat flux and midday Bowen ratios, yet woodland latent heat flux and ET was similar to the prairie during the study, factors that could affect regional climate. Land management decisions regarding grazing, prescribed burning, and stock pond design will impact the watershed hydrology and productivity of the tallgrass prairie.

Master of Science / Department of Agronomy / Walter H. Fick / Rough-leaved dogwood (Cornus Drummondii Meyer) is one of the most invasive woody plants in rangeland of Kansas. Reduced prescribed burning due to drought and urban encroachment probably has contributed to its spread. Herbicides are commonly recommended for control of rough-leaved dogwood, but minimal data exists for recommendation development. Ten herbicide treatments were applied in June during late flowering at two locations in northeast Kansas in 2005 and 2006. Each treatment was replicated three times in a randomized complete block design with individual plot sizes of 3 x 3 m. Herbicides were applied with hand sprayers in 1017 L/ha solution. Visual evaluations of defoliation were made about 1 and 12 months after treatment (MAT) and mortality was estimated about 1 year after treatment (YAT). Defoliation 1 MAT varied among herbicides with significant location by year and herbicide by year interactions. Treatments providing greater than 70% defoliation 12 MAT both years were triclopyr + 2,4-D (1.12 + 1.06 kg ae/ha), triclopyr + fluroxypyr (1.83 + 0.61 kg ae/ha), picloram + fluroxypyr (0.41 + 0.41 and 0.82 + 0.82 kg ae/ha), and picloram + 2,4-D + triclopyr (0.66 + 2.44 + 2.44 kg ae/ha). Rough-leaved dogwood is difficult to control with a single herbicide application, but treatments exist that will substantially reduce stands. Tebuthiuron pellets (Spike 20P) are another control measure recommended for rough-leaved dogwood. Treatments of 4.4 kg ai/ha (3/4 oz per 100 square feet) tebuthiuron pellets were applied in December 2004. A visual estimate of control indicated tebuthiuron reduced dogwood cover by 65% compared to a 3% decrease on untreated plots. Dogwood density was reduced by 2.2 stems/m2 (P<0.08). Total woody plant cover increased on untreated plots by 6.2 percentage units, but was decreased by 20.9 percentage units on tebuthiuron treated plots. Other woody plants decreased in both treated and untreated plots. Shading by a large elm tree likely caused variation between replications including increases in cover and density of rough-leaved dogwood on treated plots. Tebuthiuron is a photosynthetic inhibitor that often is not effective on shaded plants. Tebuthiuron pellets applied at 4.4 kg ai/ha was an effective control option for unshaded rough-leaved dogwood.

Master of Science / Department of Agronomy / Walter H. Fick / Patch-burn grazing is a relatively new concept in terms of rangeland management. While numerous benefits have been associated with this system, in the tallgrass prairie of Kansas, cattle production and sustainability of rangeland are critical. In 2006, 253 ha at the KSU Bressner Range Research Unit in Woodson County, Kansas were subjected to spring patch-burn grazing (using one-third portions) and traditional full-burn grazing. Each treatment within the split-block design was replicated four times for 3 years. The objectives were to evaluate whether livestock performance would be compromised under this grazing system, to monitor the health of the rangeland, and to observe the usefulness of this tool as a potential control of the invasive plant sericea lespedeza [Lespedeza cuneata (Dumont) G. Don]. In regards to cattle performance, burn treatments had no significant difference in average daily gain (p≥0.10) in any of the 3 years. On average, cattle utilized 61% of the current year’s forage production in patch-burned portions, which was higher (p≤0.10) than that of unburned (30%) and full-burn (41%) portions. Results of the botanical composition show forb and woody plant composition did not differ between treatments, however differences (p≤0.10) were present in grass composition. Total annual grasses increased 19.1 percentage units under patch-burn and 2.1 units under full-burn, while total perennial grasses decreased 18.4 and 1.1 units, respectively. When evaluated by treatment area (one-third portion), results indicated that the 3-year cycle did allow enough time for recovery. At 2 years after treatment (2-YAT), no significant difference in composition(p≥0.10) was present between initial patch-burn portions and the full-burn pastures. Finally, in only 1 year of the study did cattle statistically consume a greater percentage of sericea lespedeza plants (p≤0.10) in the patch-burned portions (92%) than in full-burned pastures (35%). Biomass utilization did not differ (p≥0.10) between burn treatments. Surprisingly, there was a trend for the number of plants in the sampled areas of the patch-burn portions to decrease throughout the cycle. However, at the conclusion of the 3-year cycle, sericea densities did not differ(p≥0.10) between treatments. Patch-burn grazing shows promise as a potential management tool for Kansas land managers.

Doctor of Philosophy / Department of Agronomy / Clenton E. Owensby / Most rangelands are harvested at some point during the year and removal of plant leaf area and biomass alters a host of ecosystem processes including gas exchange. An experiment was conducted in 2005 and 2006 to study the effects of clipping tallgrass prairie at different dates on water vapor and CO2 fluxes. A portable, non-steady-state chamber was designed to measure CO2 and water vapor fluxes from small plots in less than 40 s. A combination of sunlit and shaded readings allowed measurements of net carbon exchange (NCE) and ecosystem respiration (RE); by summing NCE and RE, gross canopy photosynthesis (GCP) was calculated. Throughout the two-year study, the chamber had a minimal effect on microclimate, i.e., average chamber temperature increased 2.9° C, while chamber pressure increased only 0.3 Pa during measurements, and photosynthetically active radiation attenuation was 10%. The immediate effect of all clipping treatments was a loss of leaf area that led to reductions in GCP, NCE, and RE and in most cases decreased water vapor flux. Further patterns of carbon flux were governed by the amount of water stress during canopy development, while water vapor flux rates varied with water availability. Canopies that developed during periods of low water stress quickly increased carbon flux rates following precipitation after a mid-season drought. However, flux rates of canopies, which developed during the mid-season drought, responded considerably slower to subsequent water availability. A separate experiment was conducted from June-October of 2006 to estimate GCP, leaf area index (LAI), and total aboveground biomass with a hyperspectral radiometer. Indices such as the Normalized Difference Vegetation Index and the Simple Ratio were used to estimate LAI and biomass had poor correlations with measured values. However, GCP was significantly correlated to all six indices derived in this study. While GCP measured from June-October was significantly correlated with all indices, removal of the senesced canopy scans recorded during October greatly increased the relationship.

Doctor of Philosophy / Department of Biology / Kimberly A. With / The loss, fragmentation, and degradation of grasslands have resulted in widespread declines in grassland birds. Nest predation is the leading cause of avian reproductive failure; therefore minimizing nest predation can lessen the severity of bird declines. Snakes are important predators of bird nests, but little is known about how snakes may enhance predation risk. To address this issue, I studied the habitat use, movement behavior, population genetic structure, and connectivity of snakes in the grasslands of northeastern Kansas. I addressed the connectivity of eastern yellowbelly racer (Coluber constrictor flaviventris) populations by using a landscape genetics approach at a broad scale (13,500 km2). I also radio-tracked the yellowbelly racer and Great Plains ratsnake (Pantherophis emoryi) at Konza Prairie Biological Station to understand their spatial ecology while simultaneously evaluating nest survival in grassland birds. Individual racers had limited dispersal (<3 km), but substantial admixture occurred within 30 km and populations were in migration-drift equilibrium and had high allelic diversity; therefore, racers must be abundant and continuously distributed for gene flow to be fluid throughout the region. Racers may be more likely to encounter bird nests, as they had more frequent movements and traversed greater distances on average than ratsnakes, which exhibited long periods of inactivity between directed movements. As for grassland birds, nest survival rates decreased with increasing shrubs and decreasing vegetation height. Discriminant function analysis revealed that successful nests were likely to occur in tall vegetation but reduced shrub cover, whereas higher shrub cover characterized snake habitats. Because snakes often use shrubs, nests in areas of increased shrubs may be at higher risk of predation by snakes. Targeted removal of shrubs may increase nest success by minimizing the activity of predators attracted to shrubs. Although predator removal is often a strategy for protecting bird populations, it may not be feasible in this instance, especially since snakes are a native component of the grassland community. Efforts to reduce snake predation on grassland bird nests should therefore focus on managing habitat within grasslands (i.e., shrubs) that influence snake activity, as no natural or anthropogenic habitat barriers currently limit snake movement across the landscape.

Doctor of Philosophy / Department of Agronomy / Walter H. Fick / Sericea lespedeza [Lespedeza cuneata (Dumont) G. Don] is a rangeland weed of the tallgrass prairie of Kansas. Experiments were carried out during the 2001-2003 growing seasons (June-November) to examine the relationship between sericea lespedeza and other rangeland species, wildlife use and dispersal of sericea lespedeza propagules, and the relationship between sericea lespedeza stubble height and utilization by livestock. Sericea lespedeza cover was positively correlated with violet lespedeza (r=0.25) [Lespedeza violacea (L.) Pers.], heath aster (Aster ericoides L.) (r=0.23) and total forb cover (r=0.56). Sericea lespedeza composition was negatively correlated with big bluestem (Andropogon gerardii Vitman.) (r=-0.27) and sideoats grama [Bouteloua curtipendula (Michx. Torr.)] (r=-0.27) composition and positively correlated with western ragweed (Ambrosia psilostachya D.C.) (r=0.21) and violet lespedeza (r=0.36) composition. Sericea lespedeza density was inversely related to forb species richness on all sites. On half the sites, there was also an inverse relationship between grass species richness and sericea lespedeza density. Grazed sites had less difference in grass species richness between high and low levels of sericea lespedeza densities. There was no evidence of an ecological threshold to the detriment of species richness with increasing sericea lespedeza density. Cattle digestive processes did not affect sericea lespedeza germination. Quail digestive processes enhanced germination of the few seeds that were excreted. Quail diet selection was investigated with a field study. Five out of 49 crops collected contained seed classified as sericea lespedeza. None of these seeds germinated in the greenhouse. Both cattle and quail could potentially disperse sericea lespedeza seed, but voluntary consumption appears to be low for both species, at least during November when other food is available. An height-weight table estimating percent utilization for various grazed and ungrazed heights of sericea lespedeza was constructed. Coefficient of determination values were greater than 0.85 between plant height and weight, indicating that the heightweight method was appropriate for estimating sericea lespedeza utilization. Using additional regression analysis, a chart for estimating forage from percent of plants grazed was constructed.

Doctor of Philosophy / Department of Agronomy / Stephen J. Thien / Range and Training Land Assessment (RTLA) and Land Rehabilitation and Maintenance (LRAM) are key components of The United States Army's Integrated Training Area Management (ITAM) Program, which outlines its commitment to support the sustainable use of military training lands. The primary purpose of the RTLA Program is to provide information and recommendations regarding the condition of training lands to range managers for scheduling of training areas and monitoring the effectiveness of rehabilitation projects. The goal of the LRAM component of ITAM is to reduce the long-term impacts of training on installations through the implementation of improvements to vegetation cover and repairs to landscape damage in disturbed areas. Fort Riley Military Installation, located in the largest remaining expanse of tallgrass prairie in the Flint Hills of northeastern Kansas, is a major training reservation, with seventy percent of its 40,434 ha used for mechanized maneuvers. A randomized complete block design composed of M1A1 tank traffic in a figure-eight pattern during wet and dry soil conditions was established in each of two soil types, a silty clay loam and a silt loam, and recovery of physical, chemical, and biological indicator variables was monitored from 2005 through 2007. In a second study, the effectiveness of LRAM procedures, including leveling, mulching, and reseeding, was evaluated following wheeled vehicle disturbance. The goals of this study were to identify disturbance indicators appropriate for assessing soil quality and, based on the status of these indicators, develop a method for modeling the stage and rate of ecological degradation and potential response to remediation. Disturbance increased significantly during wet compared to dry soil conditions, for increased traffic intensity, and for curve compared to straight-a-way areas in both soil types. The greatest impacts were on above and below ground community structure, providing an effective bioindicator of ecosystem health for military training land managers. Remediation of wheeled vehicle disturbance with leveling and mulching, but not reseeding, increased total vegetation production. The tallgrass prairie typically is considered to be among the most resilient of military training lands, but resiliency is dependent upon soil type and training conditions, and may require longer periods of recovery than previously thought.