No description available.

The fossil record is the only direct source of data for studying modes (patterns) and rates of morphological change over geologic time periods. Determining modes is critical for understanding macroevolutionary processes, but just how modes can vary within a taxon, and why, have hitherto been largely understudied. To address this, I examined patterns of morphological change in the shell of the Mesozoic marine bivalve genus Buchia over its geographic and temporal range. Buchia was chosen as a test subject because it is abundant, well-preserved across a variety of facies, and is widely distributed across the Northern Hemisphere where the likelihood of multiple lineages is low. While the focus of this thesis is on evolutionary patterns, it is also necessary to address issues of taxonomy and geographical variation, making this research applicable to a wide-variety of fields. Previous to this study there was no protocol for measuring buchiid valves, nor was the genus studied in a quantitative manner. Throughout this research I used ten morphological characters to describe shell shape and size. Multivariate methods (principle component and canonical variate analyses) were employed to discriminate between species of Buchia and examine how morphological characters change through time and space within the genus. Evolutionary patterns were delineated using two well-established programs that discriminate between multiple modes of evolution. Overall, nearly 2000 specimens from eight geographical locations around the world were studied for this thesis. I found the genus Buchia was a useful tool for evolutionary studies as it can be studied quantitatively in space and time. Specically I have found that buchiid species can be delineated using morphometrics; the genus is restricted to the Northern Hemisphere; while the environment significantly affects morphology, there is no evidence of a latitudinal gradient; diversity and disparity within Buchia are not correlated; most evolutionary modes conformed to random walks or stasis; and modes and rates vary across the geographical range of the genus. Overall, I have found that the environment plays an important role in shaping both morphology and modes. / Science, Faculty of / Earth, Ocean and Atmospheric Sciences, Department of / Graduate

No description available.

No description available.

Trabalhos objetivando a tafonomia de concentrações fossilíferas geradas em ambiente glacial ou periglacial são raros. Neste contexto, a presente dissertação realizou a caracterização tafonômica das concentrações fossilíferas da Formação Cape Melville, Grupo Moby Dick (Mioceno Inferior), Ilha Rei George, Antártica, tendo em vista a elucidação da gênese destes depósitos. Para tanto, dados relativos a um total de 534 espécimes foram obtidos nas camadas ricamente fossilíferas das quatro seções, denominadas Pingüineira (PRS), Hard Ground (HGS), Chaminé (CS) e Lava Crag (LCS), levantadas no topo da Península Melville entre Janeiro e Fevereiro de 2003. Destas, apenas a Seção PRS apresenta duas camadas fossilíferas distintas (PRS-C1 e PRS-C2). A fauna estudada é composta predominantemente por restos de moluscos bivalves. Restos menos abundantes de corais, caranguejos, gastrópodes e braquiópodes, além de icnofósseis, também estão presentes. A análise da composição taxonômica, ecológica e tafonômica permitiu identificar semelhanças entre as Seções PRS e HGS e entre as Seções CS e LCS. As seções PRS e HGS são compostas dominantemente por bivalves depositívoros da infauna rasa (Ennucula frigida, Enncula musculosa e Yoldia peninsularis), preservados preferencialmente com as valvas articuladas fechadas, porém fora da posição de vida, indicando remobilização da fauna pré-soterramento; a ausência de sinais de fragmentação, abrasão e incrustação indica que os bioclastos não foram afetados por processos bioestratinômicos químicos, físicos ou biológicos intensos. A ocorrência de bioclastos piritizados na Seção HGS é interpretada como resultante da decomposição dos organismos soterrados, ainda vivos, em ambiente anóxico. As assembléias das seções CS e LCS são constituídas dominantemente por bivalves suspensívoros da infauna profunda (Neilo (N.) rongelii), preservados preferencialmente com as valvas desarticuladas, indicando que os processos bioestratinômicos físicos foram mais atuantes, sendo, no entanto, raros os sinais de fragmentação e de outras assinaturas tafonômicas como incrustação e bioerosão. As valvas desarticuladas estão preservadas preferencialmente com a convexidade voltada para baixo indicando que os bioclastos foram colocados em suspensão e redepositados fora da posição de maior estabilidade hidrodinâmica; a orientação em planta destes bioclastos exibe direção preferencial, indicando atuação de correntes fracas e um maior tempo de exposição na interface água/sedimento; a ocorrência de raros restos de bivalves suspensívoros escavadores da infauna profunda (Panopea (P.) cf. P. regularis) e de caranguejos (Antarctidromia inflata) preservados em posição de vida é indicativa de que esta comunidade foi soterrada in situ por sedimentos em suspensão que trouxeram a tanatocenose de valvas desarticuladas. Os resultados obtidos no presente estudo reiteram a importância dos estudos tafonômicos e paleoecológicos no entendimento da dinâmica deposicional do passado, contribuindo com um grande conjunto de dados úteis na caracterização de ambientes glaciais e periglaciais. / Works focusing on the taphonomy of fossil concentrations generated in glacial or periglacial environment are rare. In this context, the present dissertation carried out the taphonomic characterization of the fossil concentrations of the Cape Melville Formation, Moby Dick Group (Lower Miocene), King George Island, Antarctica, in order to elucidate the genesis of these deposits. To this end, data relative to a total of 534 specimens were obtained in the richly fossil layers of the four sections, called Pingüineira (PRS), Hard Ground (HGS), Chaminé (CS) and Lava Crag (LCS), investigated at the top of the Melville Peninsula between January and February of 2003. Of these, only the PRS Section presents two distinct fossil layers (PRS-C1 and PRS-C2). The studied fauna is mainly composed of remains of bivalve clams. Less abundant remains of corals, crabs, gastropods and brachiopods, as well as trace fossils, also are present. Taxonomic, ecological and taphonomic analyses allowed similarities to be identified between PRS and HGS and CS and LCS. PRS and HGS Sections are dominantly composed by shallow infaunal deposit-feeding bivalves (Ennucula frigida, Enncula musculosa Yoldia peninsularis), preserved preferentially with closed articulated valves, however out of life position, indicating remobilized fauna; the absence of signs of spalling, abrasion and incrustation indicates that the bioclasts were not affected by intense chemical, physical or biological bioestratinomic processes. The occurrence of pyritized bioclasts in HGS is interpreted as the result of decomposition of the entombed organisms, still alive, in an anoxic environment. The assemblages of CS and LCS Sections are dominantly constituted by deep infaunal suspension-feeding bivalves (Neilo (N.) rongelii), preserved preferentially with disarticulated valves, indicating that the physical biostratinomic processes were more operative; however there are few signs of spalling or other taphonomic signatures such as incrustation and bioerosion. Disarticulated valves are preferentially preserved convex down indicating that bioclasts were placed in suspension and redeposited in a position other than that of greatest hydrodynamic stability; the orientation of these bioclasts in plan view shows a preferential direction, indicating weak currents and a longer time of exposition at the water/sediment interface; the occurrence of rare remains of deep infaunal suspension-feeding bivalves (Panopea (P.) cf. P. regularis) and crabs (Antarctidromia inflata) preserved in life position is indicative that this community was entombed in situ by sediments in suspension that brought the thanatocenosis of disarticulated valves. The results obtained in the present study reiterate the importance of taphonomic and paleoecological studies for the understanding of the depositional dynamics of the past and contribute a great number of data useful in the characterization of glacial and periglacial environments.

Trabalhos objetivando a tafonomia de concentrações fossilíferas geradas em ambiente glacial ou periglacial são raros. Neste contexto, a presente dissertação realizou a caracterização tafonômica das concentrações fossilíferas da Formação Cape Melville, Grupo Moby Dick (Mioceno Inferior), Ilha Rei George, Antártica, tendo em vista a elucidação da gênese destes depósitos. Para tanto, dados relativos a um total de 534 espécimes foram obtidos nas camadas ricamente fossilíferas das quatro seções, denominadas Pingüineira (PRS), Hard Ground (HGS), Chaminé (CS) e Lava Crag (LCS), levantadas no topo da Península Melville entre Janeiro e Fevereiro de 2003. Destas, apenas a Seção PRS apresenta duas camadas fossilíferas distintas (PRS-C1 e PRS-C2). A fauna estudada é composta predominantemente por restos de moluscos bivalves. Restos menos abundantes de corais, caranguejos, gastrópodes e braquiópodes, além de icnofósseis, também estão presentes. A análise da composição taxonômica, ecológica e tafonômica permitiu identificar semelhanças entre as Seções PRS e HGS e entre as Seções CS e LCS. As seções PRS e HGS são compostas dominantemente por bivalves depositívoros da infauna rasa (Ennucula frigida, Enncula musculosa e Yoldia peninsularis), preservados preferencialmente com as valvas articuladas fechadas, porém fora da posição de vida, indicando remobilização da fauna pré-soterramento; a ausência de sinais de fragmentação, abrasão e incrustação indica que os bioclastos não foram afetados por processos bioestratinômicos químicos, físicos ou biológicos intensos. A ocorrência de bioclastos piritizados na Seção HGS é interpretada como resultante da decomposição dos organismos soterrados, ainda vivos, em ambiente anóxico. As assembléias das seções CS e LCS são constituídas dominantemente por bivalves suspensívoros da infauna profunda (Neilo (N.) rongelii), preservados preferencialmente com as valvas desarticuladas, indicando que os processos bioestratinômicos físicos foram mais atuantes, sendo, no entanto, raros os sinais de fragmentação e de outras assinaturas tafonômicas como incrustação e bioerosão. As valvas desarticuladas estão preservadas preferencialmente com a convexidade voltada para baixo indicando que os bioclastos foram colocados em suspensão e redepositados fora da posição de maior estabilidade hidrodinâmica; a orientação em planta destes bioclastos exibe direção preferencial, indicando atuação de correntes fracas e um maior tempo de exposição na interface água/sedimento; a ocorrência de raros restos de bivalves suspensívoros escavadores da infauna profunda (Panopea (P.) cf. P. regularis) e de caranguejos (Antarctidromia inflata) preservados em posição de vida é indicativa de que esta comunidade foi soterrada in situ por sedimentos em suspensão que trouxeram a tanatocenose de valvas desarticuladas. Os resultados obtidos no presente estudo reiteram a importância dos estudos tafonômicos e paleoecológicos no entendimento da dinâmica deposicional do passado, contribuindo com um grande conjunto de dados úteis na caracterização de ambientes glaciais e periglaciais. / Works focusing on the taphonomy of fossil concentrations generated in glacial or periglacial environment are rare. In this context, the present dissertation carried out the taphonomic characterization of the fossil concentrations of the Cape Melville Formation, Moby Dick Group (Lower Miocene), King George Island, Antarctica, in order to elucidate the genesis of these deposits. To this end, data relative to a total of 534 specimens were obtained in the richly fossil layers of the four sections, called Pingüineira (PRS), Hard Ground (HGS), Chaminé (CS) and Lava Crag (LCS), investigated at the top of the Melville Peninsula between January and February of 2003. Of these, only the PRS Section presents two distinct fossil layers (PRS-C1 and PRS-C2). The studied fauna is mainly composed of remains of bivalve clams. Less abundant remains of corals, crabs, gastropods and brachiopods, as well as trace fossils, also are present. Taxonomic, ecological and taphonomic analyses allowed similarities to be identified between PRS and HGS and CS and LCS. PRS and HGS Sections are dominantly composed by shallow infaunal deposit-feeding bivalves (Ennucula frigida, Enncula musculosa Yoldia peninsularis), preserved preferentially with closed articulated valves, however out of life position, indicating remobilized fauna; the absence of signs of spalling, abrasion and incrustation indicates that the bioclasts were not affected by intense chemical, physical or biological bioestratinomic processes. The occurrence of pyritized bioclasts in HGS is interpreted as the result of decomposition of the entombed organisms, still alive, in an anoxic environment. The assemblages of CS and LCS Sections are dominantly constituted by deep infaunal suspension-feeding bivalves (Neilo (N.) rongelii), preserved preferentially with disarticulated valves, indicating that the physical biostratinomic processes were more operative; however there are few signs of spalling or other taphonomic signatures such as incrustation and bioerosion. Disarticulated valves are preferentially preserved convex down indicating that bioclasts were placed in suspension and redeposited in a position other than that of greatest hydrodynamic stability; the orientation of these bioclasts in plan view shows a preferential direction, indicating weak currents and a longer time of exposition at the water/sediment interface; the occurrence of rare remains of deep infaunal suspension-feeding bivalves (Panopea (P.) cf. P. regularis) and crabs (Antarctidromia inflata) preserved in life position is indicative that this community was entombed in situ by sediments in suspension that brought the thanatocenosis of disarticulated valves. The results obtained in the present study reiterate the importance of taphonomic and paleoecological studies for the understanding of the depositional dynamics of the past and contribute a great number of data useful in the characterization of glacial and periglacial environments.

Oviparous reproduction in Carditamera floridana was studied in St. Andrew's Sound, near Mexico Beach, Florida, between May 1992 and February 1994. The oviparous mode of development of Carditamera floridana is unique in the number of juveniles produced, the extent of parental care involved, the subtropical range of the species, and the lack of a distinct vitellogenic mechanism. / Typescript. / "1995." / "Submitted to the Department of Biological Science in partial fulfillment of the requirements for the degree of Master of Science." / Advisor: William H. Heard, Professor Directing Thesis. / Includes bibliographical references.

No description available.

Westphalian non-marine bivalves in Nova Scotia inhabited fresh to brackish temporary lakes. A series of lacustrine facies are identified on the basis of the faunas. Myalinid genera including the probably pseudoplanktonic Curvirimula, and Naiadites dominate the middle Westphalian A to early Westphalian B. Carbonicola occurs rarely in the late Namurian to early Westphalian A and is the only reported Anthracosiid. Anthraconauta dominates the upper Westphalian C to early Stephanian occurring in a wide range of sediments. The elongate Anthraconaia arenacea group range through the Westphalian D and occur in coarser lithologies. Small-shell Anthraconaia inhabited ephemeral lake margins. Cluster and principal components analysis are used to classify the faunas. The method requires the measurement of a small number of linear and relatively uncorrelated morphological variables and allows the construction of reproducable pictographs. Three new morphospecies are described. Principal components analysis is used to study morphological variation with respect to palaeoenvironment. Consistent morphological trends occur in Naiadites and Anthraconauta and are considered to have been the product of spat selection and phenotypic changes. With increased turbulence and sedimentation rates, the shell's centre of gravity is shifted towards the anter-umbonal region and the bivalves adopt a semi-infaunal mode of life thus, the morphological changes offer increased stability. Evolution was inhibited by the absence of marine incursions and progressive environmental change. Such palaeoenvironmental changes that did occur were local. Competition may have been of evolutionary importance in younger communities of Anthraconaia. The bivalves are of limited stratigraphical use as compared to the British faunas. In conjuction with micro- and macrofloral work, the Namurian/Westphalian A and Westphalian C/D boundaries are lowered and a possible Cantabrian stage recognised. A correlation with the British Chronozones is attempted.

No description available.