This thesis explores potential seeding techiniques to limit harmful effects of preemergent herbicide on a seeded species while simultaneously reducing abudance of annual invasive grasses. The first chapter examines the use of activated carbon seed coatings and furrows to limit herbicide effect on seeds of a perrenial bunchgrass. We found that both carbon coatings and furrows mitigated some of the herbicide effects, but that only when the two techniques were combined did we observe unaffected seedling emergence, plant density, and aboveground growth. Therefore, we suggest to management that use of carbon coatings and furrows after herbicide application can likely be used to reduce invasive annual grasses while simultaneously establishing a native bunchgrass. In chapter 2, we examine the effects of a novel preemergent herbicide indaziflam, on native seeds and compare it against a common preemergent herbicide, imazapic. We found that indaziflam provides superior long-term control of annual invasive grasses than imazapic, but that it is also more detrimental to native seeds. Our results suggest that indaziflam is best suited for control purposes only, and is hard to incorporate in restoration seeding efforts due to its strong effects on native seed.

Current revegetation practices in the lntermountain West include the use of Nordan (Agropyron desertorum) and Hycrest (A. cristatum x desertorum) crested wheatgrass on rangeland susceptible to cheatgrass (Bromus tectorum) invasion, although little quantitative data exists that compares their competitive abilities. We evaluated both the competitive ability of Hycrest and Nordan in two-species mixtures with cheatgrass, and evaluated seedling establishment characteristics for all three species in a greenhouse study. Linear and nonlinear models were developed for a range of densities for each species to predict median above-ground biomass and tiller numbers. In both experiments, increasing Hycrest and Nordan densities reduced their own biomass and tiller production while cheatgrass biomass and tiller production was not influenced. However, increasing cheatgrass densities reduced both Hycrest and Nordan biomass and tiller production, as well as its own biomass and tiller production. Examination of trends in competition indices, such as relative resources totals, substitution rates and perceived densities indicated that as seedlings, Hycrest was a better competitor with cheatgrass than Nordan at lower crested wheatgrass densities (130 plants/m2 ). Results from this experiment indicate that Hycrest is a better competitor than Nordan with cheatgrass and suggest that seeding Hycrest at lower densities than currently recommended may optimize its seedling growth when competing with cheatgrass. Future research concerning competition in these species using similar designs should focus on competition in successive years after establishment and on field experiments to verify these results.

Wyoming big sagebrush (Artemisia tridentata Nutt.) is an important native shrub in the Great Basin because of its wide distribution where it dominates over 60 million ha and provides essential habitat and forage for many varied species. The hand collection of sagebrush seed often results in seed scarcity and the available seed quantities are at times inadequate to revegetate large areas that have been disturbed, resulting in a demand for sagebrush seed. Study locations were selected near Scipio and Sahara sand dunes of Utah, and treatments were 1-) control, no treatment applied area left undisturbed 2-) general chemical strip thinning 3-) general chemical thinning of entire stand, 4-) general mechanical strip thinning, and 5-) general mechanical thinning of the entire stand. Significant differences among treatments in seed yields were collected in 2011 at Scipio but not at Sahara. At Scipio, the mechanical strip of competing sagebrush in 3m strips was the most effective of all treatment and produced 2.47kg/ha compared to 4.624kg/ha in the control, but the mechanical land area was only utilizing half the compared control area. The chemical treatments produced 1.819kg/ha and 1.31kg/ha. The percent of sagebrush mortality by each treatment determined the level of competition killed in treatment areas. All treatments at both locations killed at least 57% of the sagebrush. Chemical treatments had a consistent kill rate at both locations, although lower than anticipated, but mechanical kill was the highest at 93% in Scipio. Both mechanical and chemical treated plots had increased cover levels of cheatgrass when compared to the control plots.

Thesis (M.S.)--Oregon State University, 2006. / Printout. Includes bibliographical references (leaves 27-31). Also available on the World Wide Web.

The conversion of sagebrush-steppe communities of the Great Basin into cheatgrass-dominated communities is one of the most dramatic ongoing land conversions in North America. Although restoration of these communities is a high priority to landowners and land management agencies, restoration of native vegetation is difficult. Several restoration treatments intended to increase the success of aerially‐seeded perennial grasses in cheatgrass-invaded sagebrush ecosystems were assessed to determine their effects on perennial seedling emergence and soil seed bank density and composition. Assessed restoration treatments were: 1) vegetation manipulation (sagebrush thinning and prescribed burning); 2) imazapic herbicide application; 3) seedbed amendments (aerial seeding with activated carbon addition, aerial seeding with sucrose addition); and 4) seeding frequency. The effects of these treatments were evaluated in two distinct sagebrush shrubland ecosystems in northern Utah. One is characterized as a remnant sagebrush stand with a cheatgrass-dominated understory and the other as a cheatgrass near-monoculture, completely lacking a sagebrush component. In the seed bank study, responses were assessed immediately and 1 year following treatment while in the seedling emergence study, they were assessed 2 and 3 years following treatment. Main effects of vegetation manipulation, herbicide application, and seedbed addition treatments and their interactions on perennial seedling emergence are described in Chapter 2. The effects of seeding frequency on perennial seedling emergence are also described in Chapter 2. Herbicide demonstrated potential for increasing native perennial grass emergence, although this response was delayed and not seen until 3 years post-application. Burning showed potential for increasing the emergence of perennial grasses 2 years post-burn. Results also suggest that potential exists to increase native perennial grass emergence through an increase in seeding frequency. In Chapter 3 I evaluated the effects of vegetation manipulation, herbicide application, and seedbed addition on seed pool dynamics. These results suggest that herbicide and sucrose may be useful tools for reducing exotic species richness in cheatgrass-invaded systems. Herbicide also showed potential for reducing cheatgrass seed bank densities. Additionally, results demonstrated that the reductions in cheatgrass seed bank densities observed immediately after fire are still observed 1 year post-burn.

Cheatgrass is an invasive weed that has come to dominate large areas of the western United States. Once an ecosystem has been converted to a cheatgrass monoculture, it is extremely difficult to restore native vegetation. Cheatgrass negatively impacts wildlife and increases wildfire frequency and intensity. Understanding how cheatgrass so effectively invades western ecosystems is essential to turning the tide of invasion. One possible key to cheatgrass’ success is alteration of soil nutrient cycling. The goal of this study is to explore how nitrogen (N) may accumulate in cheatgrass soils via redistribution of N within soil N pools. To accomplish this we investigated soil N cycling in soils underneath cheatgrass and crested wheatgrass. We used a 15N isotope tracer to determine the contribution of root exudates to soil N pools. During the 1-week 15N tracer experiment, cheatgrass roots exuded more than twice as much N (0.11 mg N kg-1 soil d-1) as crested wheatgrass roots (0.05 mg N kg-1 soil d-1). We propose that exudation of high N content root exudates leads to the changes in soil N pool size and transformation rates commonly observed in soils under cheatgrass. This research uses a simple and relatively inexpensive isotope tracer to shed light on mechanisms by which invasive plants may alter soil processes. By understanding these mechanisms we may be able to develop strategies for better managing cheatgrass invasion.

Graduation date: 2004

The conversion of sagebrush-steppe communities of the Great Basin into cheatgrass-dominated communities is one of the most dramatic ongoing land conversions in North America. Although restoration of these communities is a high priority to landowners and land management agencies, restoration of native vegetation is difficult. Several restoration treatments intended to increase the success of aerially‐seeded perennial grasses in cheatgrass-invaded sagebrush ecosystems were assessed to determine their effects on perennial seedling emergence and soil seed bank density and composition. Assessed restoration treatments were: 1) vegetation manipulation (sagebrush thinning and prescribed burning); 2) imazapic herbicide application; 3) seedbed amendments (aerial seeding with activated carbon addition, aerial seeding with sucrose addition); and 4) seeding frequency. The effects of these treatments were evaluated in two distinct sagebrush shrubland ecosystems in northern Utah. One is characterized as a remnant sagebrush stand with a cheatgrass-dominated understory and the other as a cheatgrass near-monoculture, completely lacking a sagebrush component. In the seed bank study, responses were assessed immediately and 1 year following treatment while in the seedling emergence study, they were assessed 2 and 3 years following treatment. Main effects of vegetation manipulation, herbicide application, and seedbed addition treatments and their interactions on perennial seedling emergence are described in Chapter 2. The effects of seeding frequency on perennial seedling emergence are also described in Chapter 2. Herbicide demonstrated potential for increasing native perennial grass emergence, although this response was delayed and not seen until 3 years post-application. Burning showed potential for increasing the emergence of perennial grasses 2 years post-burn. Results also suggest that potential exists to increase native perennial grass emergence through an increase in seeding frequency. In Chapter 3 I evaluated the effects of vegetation manipulation, herbicide application, and seedbed addition on seed pool dynamics. These results suggest that herbicide and sucrose may be useful tools for reducing exotic species richness in cheatgrass-invaded systems. Herbicide also showed potential for reducing cheatgrass seed bank densities. Additionally, results demonstrated that the reductions in cheatgrass seed bank densities observed immediately after fire are still observed 1 year post-burn.

Glasshouse and growth-chamber experiments were conducted to evaluate primisulfuron phytotoxicity and the influence of adjuvants on downy brome and Kentucky bluegrass. GR₅₀ (50% growth reduction) values were 0.97 ± 0.57 and 8.07 ± 1.85 g/ha for downy brome and Kentucky bluegrass, respectively. Primisulfuron was applied to downy brome and Kentucky bluegrass at 3 placement sites; foliar, soil, and foliar plus soil. Foliar or foliar plus soil applications were more effective at reducing downy brome dry weights than the soil application of primisulfuron, while Kentucky bluegrass was injured more from the soil or foliar plus soil applications than from the foliar application of primisulfuron. Primisulfuron at 5 g/ha applied alone reduced downy brome dry weights by 5%, whereas, when an adjuvant was added, dry weights were reduced by 52 to 83%. Primisulfuron was more phytotoxic to downy brome at alternating temperatures of 8-16 C and 16-24 C than at 0-8 C. Phytotoxicity of primisulfuron was less when downy brome plants were stressed for soil moisture after herbicide treatments than when the plants were not stressed or only stressed before treatment. / Graduation date: 1999

Thesis (Ph. D.)--Oregon State University, 2011. / Printout. Includes bibliographical references (leaves 123-135). Also available on the World Wide Web.