Crested wheatgrass (Agropyron cristatum [L.] Gaertn.) is one of the most commonly seeded exotic species in the western United States. Although many degraded Wyoming big sagebrush (Artemisia tridentata ssp. wyomingensis) plant communities have been seeded with crested wheatgrass during rehabilitation efforts, seed banks of these communities have not been characterized. I sought to characterize and explain the variation among 33 seeded communities in the northeastern Great Basin. Hierarchical clustering and principal components analysis identified four possible seed bank categories in these communities. Seed bank categories varied from high to low crested wheatgrass dominance. The crested wheatgrass-dominated category is a particularly attractive setting to develop management strategies that reduce crested wheatgrass dominance and increase native plant diversity. It is also a common practice to seed crested wheatgrass in mixture with native species after a disturbance to increase diversity. Empirical estimates as to how the density of crested wheatgrass seed and seedlings interfere with native species establishment have not been defined. A greenhouse experiment was established using an addition series design to determine the influence of interference between crested wheatgrass and four important native species. The existence of seed bank categories of Wyoming big sagebrush communities seeded with crested wheatgrass agrees with the hypothesis that seed banks closely resemble floristic composition. In addition, these results support the hypothesis that seed bank composition has a strong influence on succession in these communities, and characterizing seed banks is necessary to develop ecologically based management strategies for seeded Wyoming big sagebrush communities. Interference from crested wheatgrass on many native species suggests that further management practices to enhance diversity in crested wheatgrass-dominated communities are necessary to reduce competition from crested wheatgrass in the seed bank as well as the aboveground vegetation. These results also suggest that the practice of simultaneously seeding native species with crested wheatgrass may likely result in poor native species persistence unless combined seed bank density and seeding rate of crested wheatgrass is sufficiently low.

A comparative index to digestible nutrients was established for Russian wildrye, crested wheatgrass, and intermediate wheatgrass seeded on foothill areas of Utah. The nutritive values of these grasses were compared by the lignin ratio method, the chlorophyllan ratio method, and the in vivo rumen bag process. Average digestion coefficients computed by the lignin ratio method for dry matter, ether extract, cellulose, and gross energy were highest for intermediate wheatgrass. Protein was more digestible for Russian wildrye. Digestible energy was adequate in all species to meet requirements for late lactation. Results from the chlorophyllan ratio method were excessively high and were not considered valid. Dry matter loss of Russian Wildrye was greater than losses from either of the wheatgrasses. Losses of dry matter due to 24-hour in vivo rumen fermentation and rinsing only of nylon bags were closely correlated.

I investigated the effect of energy sector development and introduced crested wheatgrass (Agropyron cristatum) on grassland birds on Canadian Forces Base Suffield. I conducted point counts and mapped breeding territories in 2007 and 2008 for Savannah sparrows (Passerculus sandwichensis), chestnut-collared longspurs (Calcarius ornatus), and Spragues pipits (Anthus spragueii). I found that Savannah sparrows favored areas with taller vegetation, human disturbances and crested wheatgrass in both years. Longspurs used shorter vegetation and in were tolerant of disturbance. Crested wheatgrass was avoided by longspurs in both years. Pipit territories contained similar vegetation to longspurs, were sensitive to disturbance, and avoided placing territories in areas containing crested wheatgrass or trails in both years. Well sites, pipelines and junctions were not avoided by the three species. My research suggests that reducing the number of trails and the spread of crested wheatgrass will increase habitat availability for sensitive species of grassland birds. / Ecology

A series of experiments were conducted during 2005 and 2006 to evaluate five perennial grass species for forage yield and quality, steer performance and grazing capacity, animal intake, plant energy reserves and economic return under grazed conditions. In 1999, two 0.8 ha replicates each of Paddock meadow bromegrass (<i>Bromus riparius</i>Rehm.), Carlton smooth bromegrass (<i>Bromus inermis</i>Leyss.) and AC Knowles hybrid bromegrass (<i>B. riparius</i> x <i>B. inermis</i>) were seeded. In 2003, two 0.8 ha replicates each of AC Goliath crested wheatgrass (<i>Agropyron cristatum </i> (L.) Gaertn.), hybrid bromegrass, and Courtenay tall fescue (<i>Festuca arundinacea </i> Schreb.) were seeded. A long established stand of crested wheatgrass acted as the control pasture. For 2003 established pastures, AC Goliath crested wheatgrass (7515 kg ha-1) had greater (P<0.05) cumulative dry matter yield than hybrid bromegrass (3136 kg ha-1) during the 2005 grazing season. Average (2005-2006) crude protein (CP) was greatest (P<0.05) for hybrid and smooth bromegrass for 1999 established pastures at start and middle of period one. Control pastures had the greatest (P<0.05) neutral detergent fiber (NDF) mid-grazing period. Over 2 years, smooth bromegrass had greater acid detergent fiber (ADF) (P<0.05) than control pastures at the end of the grazing period one. Average (2005-2006) in vitro organic matter digestibility (IVOMD) was greatest for hybrid and meadow bromegrass (P<0.05) at the start of grazing period one. Control pastures (129 g kg-1) had lower CP levels at the start of the 2005-2006 (average) grazing period 1 (P<0.05) compared to species seeded in 2003. Control and hybrid bromegrass pastures had the greatest NDF and ADF levels at the start of grazing period 1 (2005-2006 average) while tall fescue pastures had the lowest (P<0.05) NDF and ADF levels. Over 2 years, control pastures had the lowest IVOMD at start of grazing (P<0.05). In 2006, hybrid and smooth bromegrass had greater etiolated re-growth than control pastures (P<0.05). In 2006, grazed plants seeded in 1999 had greater (P<0.05) etiolated re-growth than ungrazed plants. For 2003 seeded grasses, crested wheatgrass produced greater (P<0.05) etiolated re-growth than tall fescue and control pastures. Average daily gain was similar (P>0.05) for all 1999 and 2003 seeded grasses. Overall, bromegrasses seeded in 1999 produced greater animal grazing days (AGD) than control pastures (P<0.05). Total beef production (TBP) was greater (P<0.05) for hybrid and meadow bromegrass compared to the control. All species seeded in 2003 produced greater AGD (P<0.05) compared to the control. Crested wheatgrass produced greater (P<0.05) TBP than the control over both years of the study. The C33:C32 alkane ratio estimated greater DMI (P<0.05) for hybrid bromegrass (9.9 kg d-1) and control pastures (9.6 kg d-1) compared to crested wheatgrass (6.8 kg d-1) or tall fescue (6.8 kg d-1) during period 1 in 2006. Over 2 years, net return to labor, equity and personal draw was greater (P<0.05) for hybrid bromegrass ($91.24 ha-1) compared to the control (-$54.32 ha-1). For 2003 seeded pastures, all pastures generated positive returns over 2 years. Crested wheatgrass ($92.49 ha-1) had greater net return than control pastures (-$54.32 ha-1) (P<0.05). Finally, the results of this grazing study indicate beef producers can manage these grasses during the summer grazing season and maintain high levels of animal performance and pasture production. This study has demonstrated that bromegrasses, crested wheatgrass and tall fescue could work well in a complementary grazing system.

A series of experiments were conducted during 2005 and 2006 to evaluate five perennial grass species for forage yield and quality, steer performance and grazing capacity, animal intake, plant energy reserves and economic return under grazed conditions. In 1999, two 0.8 ha replicates each of Paddock meadow bromegrass (<i>Bromus riparius</i>Rehm.), Carlton smooth bromegrass (<i>Bromus inermis</i>Leyss.) and AC Knowles hybrid bromegrass (<i>B. riparius</i> x <i>B. inermis</i>) were seeded. In 2003, two 0.8 ha replicates each of AC Goliath crested wheatgrass (<i>Agropyron cristatum </i> (L.) Gaertn.), hybrid bromegrass, and Courtenay tall fescue (<i>Festuca arundinacea </i> Schreb.) were seeded. A long established stand of crested wheatgrass acted as the control pasture. For 2003 established pastures, AC Goliath crested wheatgrass (7515 kg ha-1) had greater (P<0.05) cumulative dry matter yield than hybrid bromegrass (3136 kg ha-1) during the 2005 grazing season. Average (2005-2006) crude protein (CP) was greatest (P<0.05) for hybrid and smooth bromegrass for 1999 established pastures at start and middle of period one. Control pastures had the greatest (P<0.05) neutral detergent fiber (NDF) mid-grazing period. Over 2 years, smooth bromegrass had greater acid detergent fiber (ADF) (P<0.05) than control pastures at the end of the grazing period one. Average (2005-2006) in vitro organic matter digestibility (IVOMD) was greatest for hybrid and meadow bromegrass (P<0.05) at the start of grazing period one. Control pastures (129 g kg-1) had lower CP levels at the start of the 2005-2006 (average) grazing period 1 (P<0.05) compared to species seeded in 2003. Control and hybrid bromegrass pastures had the greatest NDF and ADF levels at the start of grazing period 1 (2005-2006 average) while tall fescue pastures had the lowest (P<0.05) NDF and ADF levels. Over 2 years, control pastures had the lowest IVOMD at start of grazing (P<0.05). In 2006, hybrid and smooth bromegrass had greater etiolated re-growth than control pastures (P<0.05). In 2006, grazed plants seeded in 1999 had greater (P<0.05) etiolated re-growth than ungrazed plants. For 2003 seeded grasses, crested wheatgrass produced greater (P<0.05) etiolated re-growth than tall fescue and control pastures. Average daily gain was similar (P>0.05) for all 1999 and 2003 seeded grasses. Overall, bromegrasses seeded in 1999 produced greater animal grazing days (AGD) than control pastures (P<0.05). Total beef production (TBP) was greater (P<0.05) for hybrid and meadow bromegrass compared to the control. All species seeded in 2003 produced greater AGD (P<0.05) compared to the control. Crested wheatgrass produced greater (P<0.05) TBP than the control over both years of the study. The C33:C32 alkane ratio estimated greater DMI (P<0.05) for hybrid bromegrass (9.9 kg d-1) and control pastures (9.6 kg d-1) compared to crested wheatgrass (6.8 kg d-1) or tall fescue (6.8 kg d-1) during period 1 in 2006. Over 2 years, net return to labor, equity and personal draw was greater (P<0.05) for hybrid bromegrass ($91.24 ha-1) compared to the control (-$54.32 ha-1). For 2003 seeded pastures, all pastures generated positive returns over 2 years. Crested wheatgrass ($92.49 ha-1) had greater net return than control pastures (-$54.32 ha-1) (P<0.05). Finally, the results of this grazing study indicate beef producers can manage these grasses during the summer grazing season and maintain high levels of animal performance and pasture production. This study has demonstrated that bromegrasses, crested wheatgrass and tall fescue could work well in a complementary grazing system.

No description available.

Fairway crested wheatgrass, which is identified taxonomically as Agropyron cristatum (L . ) Gaertn. (45 ), A. cristatiforme (38) , or A. pectiniforme Roem. and Schult (22), is an economically important range grass belonging to the "crested wheatgrass complex" (24, 38). The crested wheatgrass complex includes diploid, 2n = 14, tetraploid, 2n = 28, and hexaploid, 2n = 42, forms (1, 11, 22). The variety Fairway and Fairway-like derivatives are the only known diploid members of the species complex (24, 38). Meiotic chromosome behavior of Fairway diploids appears to be typical of other diploid species; however, the number of plants examined cytologically has been relatively small. Although Fairway crested wheatgrass is a good seed producer, interplant variation in fertility is high (13, 22, 25, 42). Irregular chromosome behavior is a common source of sterility and may be contributing to the variable seed set in diploid crested wheatgrass. No information is available concerning the relation of meiotic chromosome behavior to fertility in Fairway crested wheatgrass. Polyploid crested wheatgrasses are generally considered to be of autoploid origin, i.e., they are derived by duplication of the chromosome complement of a diploid prototype. Chromosome pairing in the polyploid species (31), in interspecific hybrids (12), and in polyhaploid plants (11) substantiate the autoploid derivation of polyploid crested wheatgrass. Diploid and tetraploid forms of crested wheatgrass have been hybridized by Knowles (24), and chromosome pairing in the hybrids suggest a close relation between the diploid and tetraploid genomes. Colchicine-induced tetraploids of Fairway crested wheatgrass have been produced by Knowles, 1 and these artificial tetraploids are currently being utilized in his crested wheatgrass breeding program. If the full breeding and cytogenetic potentials of diploid crested wheatgrass are to be realized, the meiotic chromosome behavior and the cytotaxonomic status of this species must be fully understood. The present investigation was designed to provide further information concerning the cytogenetic characteristics of Fairway crested wheatgrass and its autotetraploid derivatives. This investigation was established with the following objectives: 1. To examine meiotic chromosome behavior of Fairway crested wheatgrass. 2. To determine the relation of meiotic chromosome behavior to fertility in Fairway crested wheatgrass. 3. To evaluate the effectiveness of several colchicine treatments in doubling the chromosome complement of Fairway crested wheatgrass. 4. To determine the effect of induced polyploidy on plant morphology in colchicine-induced tetraploids of Fairway crested wheatgrass. 5. To determine the meiotic chromosome behavior and fertility of induced tetraploids of Fairway crested wheatgrass.

Aboveground biomass, litter biomass and root biomass of a crested wheatrgrass (Agropyron desertorum [Fisch.] Schult.) dominated community were inventoried in the fall of 1971, 1972, and 1973. In addition, energy, nitrogen, fats and ash determinations were made on the materials collected in 1972 and 1973. The sampling methods used generated data sufficiently precise to detect significant differences (α = .10) among biomass components among years. The chemical contents of the components were similar in the fall of 1972 and the fall of 1973 despite the large differences in growing season precipitation. A simple linear regression formula was generated from which aboveground biomass was predicted using individual plant volume as the independent variable. Regression techniques were tried in an effort to use aboveground biomass to predict root and litter biomass. This approach proved unsuccessful because of high variability within the data. Changes in the biomass of the components were analyzed with respect to differing precipitation regimes. Aboveground biomass responded positively and linearly to increasing growing season precipitation. Litter biomass decreased as current growing season precipitation increased. However, litter increased as a function of increasing previous-growing-season precipitation. Root biomass decreased with increasing previous-growing-season precipitation. It was found that both litter:shoot and root:shoot ratios decreased as a function of increasing growing season precipitation.

During 1980-81, studies with crested wheatgrass (Agropyron desertorum) and fourwing saltbush (Atriplex canescens) were conducted in controlled environment growth chambers as well as under field conditions to achieve the following objectives: 1. To determine the effect of nitrogen fertilizer on the water use efficiency. 2. To determine the effects of various temperature, water stress and nitrogen treatments on the productivity, nitrogen content and carbohydrate reserves. 3. To determine the effects of N fertilization on fall and spring regrowth. Crested wheatgrass and fourwing saltbush plants were maintained in three growth chambers for 60 days under three temperature regimes (11/7, 19/7 and 27/7 C), two soil moisture stress regimes (-0.3 bars and -15 bars) and three N fertilizer levels (0, 50 and 100 kg of N/ha). During the study, tranpiration and plant biomass data were recorded. During the first week of September, 1980, crested wheatgrass and fourwing saltbush pastures at Nephi, Utah, were subjected to three nitrogen fertilizer levels (0, 50 and 100 kg N/ha). After 60 days the fall regrowth was clipped. In the first week of June 1981 spring regrowth of both species was measured. In the fall of 1981, a second experiment was laid out at Nephi where crested wheatgrass and fourwing saltbush plants were subjected to three soil moisture regimes (dry, medium and wet) and three nitrogen fertilizer levels. At the end of a 60 day study period, dry matter yield, root distribution, water content and soil samples at different incremental soil depths were collected. Under controlled environment conditions, the water use efficiency of both species was six percent more with the application of a moderate amount of nitrogen (50 kg/ha). A high temperature regime (27/7 C) and a high water stress regime (-15 bars) increased the water use efficiency of plants by eight and six percent respectively. Results of the growth chamber experiment revealed that nitrogen fertilization had a significant effect on plant biomass, nitrogen percent and total nonstructual carbohydrate reserves of crested wheatgrass and fourwing saltbush. The data further suggested that nitrogen fertilization can substitute for the adverse effects of low temperature and low soil moisture on plant growth. Nitrogen fertilization during fall increased plant biomass, nitrogen percent and total nonstructural carbohydrate reserves in crested wheatgrass and fourwing saltbush. Fall fertilization did not reduce spring regrowth. It is inferred that under limited soil moisture and low temperature during the fall growing season, a moderate amount of nitrogen fertilizer (50 kg N/ha) may increase the forage availability and water use efficiency of crested wheatgrass and fourwing saltbush to the level of plants maintained at moderate temperature and adequate soil moisture. Nitrogen fertilization (50 kg N/ha) of crested wheatgrass and fourwing saltbush during fall does not reduce plant nitrogen percent or carbohydrate reserves which may limit spring regrowth.

Crested wheatgrass arrived in North America at the turn of the twentieth century through the foreign plant exploration missions sponsored by the United States Department of Agriculture. During the first two decades of the new century, scientists tested the grass at agricultural experiment stations. They determined it was useful for grazing and particularly valuable because it could grow in drought conditions with little or no care and would continue to produce high quality feed even after several years of heavy use. Beginning in the 1930s federally sponsored land utilization and agricultural adjustment programs sponsored the use of crested wheatgrass for soil conservation and weed control. The grass protected the soil on the land that had been entered into the acreage reserves and the conservation reserves programs of the federal soil bank. Also in the late 1930s and through the 1960s, rangeland managers used crested wheatgrass to improve forage productivity on public lands that were used for grazing. By the 1970s somewhere between 12 and 20 million acres of crested wheatgrass grew in North America in eleven western states, and in Saskatchewan and Alberta. By 1980 attitudes about agriculture and wilderness had changed in the United States and land management was focused on multiple uses and on protecting ecosystems and native species. Attitudes about grazing and agricultural landscapes had changed and many preferred nonagricultural landscapes and land uses. As a result, crested wheatgrass went from being considered one of the most valuable plants in North America to being considered an invasive weed, in some quarters. Debates in the last 25 years have tried to determine if, where, and how crested wheatgrass belongs in North America. This thesis explains the discourses, or interest groups, that are participating in the current conversation. One impulse is to use empirical evidence to determine whether or not introduced plants like crested wheatgrass belong, but the main contention of this thesis is that empirical studies alone will always be insufficient measures because belonging is also a subjective and experientially or emotionally derived measure.