A timber frame is a structural building system composed of heavy timber members connected using carpentry-style joinery that may include metal fasteners. A common variant of mortise-and-tenon joints are keyed (or wedged) through-tenon joints. No research on the behavior of wedged joints in timber frames is available. This research provides design knowledge of keyed through-tenon joints from experimental observations and comparisons between mathematical models and experimental measurement. Evaluation of through-tenon keyed mortise and tenon joints was performed by measuring tensile load and stiffness of white oak (Quercus alba) and Douglas-fir (Pseudotsuga menziesii) joints with four- and 11-inch tenons with one and two keys and comparing these results to mathematical models developed from the National Design Specification of Wood Construction (NDS), General Dowel Equations for Calculating Lateral Connection Values (TR-12), and engineering mechanics. Variables included joint species (white oak or Douglas-fir), protruding tenon length (four or 11 inches), and number of keys (one or two). Joints were tested to ultimate load, then model input specimens were cut from tested joints and additional key stock to generate inputs for joint load predictions that were compared to experimental joint load results for validation. Forty joints were tested with white oak keys and six of these joints were retested with ipe (Tabebuia) keys. Joints with four-inch tenons behaved in a brittle manner with tenon failures. Most joints with 11-inch tenons behaved in a ductile manner with key bending and crushing failures. Joint load and stiffness was similar between white oak and Douglas-fir joints. Joints with 11-inch tenons had greater load and stiffness than with four-inch tenons. Joints with two keys had greater load and stiffness than joints with one key, after normalizing joint load and stiffness responses on key width. Joints retested with ipe keys had greater load than joints originally tested with white oak keys. Tenon relish (row tear-out) failure was predicted for all joints with four-inch tenons. Horizontal key shearing was predicted for all joints with 11-inch tenons. Ratios of predicted ultimate joint load divided by experimental ultimate joints load (calculated/tested) or C/T ratios were used to validate the models chosen for load prediction. C/T ratios showed that ultimate load model predictions over predicted joint load which was due to occurrence of unpredicted tenon failures and simultaneously occurring key failures where models predicted key failures independently. Design safety factors (DSFs) were developed by dividing experimental ultimate joint load by governing allowable (design) load predictions. C/T ratios and DSFs were most similar between white oak and Douglas-fir joints and most different between joints with one and two keys. Alternative design values (ADVs) were developed for comparison to design load predictions. Comparisons between ADVs and DSFs showed that model predictions were most conservative for joints fastened with denser keys than joint members. / Master of Science

Douglas-fir (Pseudotsuga menziesii) and lodgepole pine (Pinus contorta) in southwestern Alberta were studied to determine the water sources used and the effect of changing soil moisture on tree ecophysiological function. The hydrogen stable isotope ratios of water from local groundwater and precipitation were compared to tree stem water to determine the amount of stem water coming from those two sources. There were no significant differences between species in the portion of summer precipitation taken up. However, Douglas-fir shifted towards using more groundwater as shallow soil moisture declined. In addition, Douglas-fir showed large changes in shoot water potential, but maintained relatively constant rates of oxygen evolution, whereas lodgepole pine exhibited smaller changes in shoot water potential and had severely reduced rates of oxygen evolution during mid-summer drought. Lower leaf-area to sap-wood area and higher leaf δ13C (carbon isotope composition) suggested a less efficient hydraulic system in Douglas-fir compared to lodgepole pine. / x, 91 leaves : ill. ; 29 cm

The western spruce budworm (Choristoneura occidentalis Freeman) is the most widespread and destructive defoliator of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) forests in British Columbia. Over the past two decades, western spruce budworm outbreaks have been sustained and widespread in the interior of British Columbia, leaving the forest industry and many forest-dependent communities increasingly vulnerable to the economic consequences of these outbreaks. While a great deal is known about the impact of western spruce budworm outbreaks on tree growth and form, substantial knowledge gaps remain as to the historic variability of western spruce budworm outbreaks and the consequences of defoliation on fundamental characteristics such as wood structure. This research focused on describing historic and contemporary western spruce budworm outbreaks across multiple spatial and temporal scales in south-central British Columbia using dendrochronology and wood anatomy techniques. Outbreak histories over the past 435 years were reconstructed using a network of tree-ring chronologies from central British Columbia, revealing that 12 western spruce budworm outbreaks have occurred since the early 1600s, with a mean return interval of 30 years. Further, the research illustrates that outbreaks observed over the last 40 years are not unprecedented, which does not support the perception that western spruce budworm is moving northward into central British Columbia. To evaluate the effects of a single western spruce budworm outbreak on the anatomical characteristics of Douglas-fir stemwood, tree ring data was collected from permanent sample plots that sustained both periodic and chronic western spruce budworm feeding. In mature even-aged stands of Douglas-fir, a documented outbreak occurred from 1976 to 1980 in the coastal transition zone of southern British Columbia. Based on microscopic wood anatomical measurements it was shown that the tree rings formed during this outbreak had significantly lower percentages of latewood, reduced mean cell wall thickness and smaller radial cell diameters relative to wood formed during periods without budworm feeding. Western spruce budworm defoliation temporarily modified cellular characteristics, which has implications for wood quality. In uneven-aged stands of mature Douglas-fir, located in the xeric southern interior of British Columbia, there has been a sustained western spruce budworm outbreak since 1997. Tree rings formed during this outbreak had progressively larger earlywood lumen area and radial cell diameter, reduced latewood cell wall thickness, latewood radial cell diameters, and lower percent latewood. Mixed-effects models revealed that climatic variables, defoliation severity, defoliation duration, and in limited cases canopy class were the best predictors of xylem features. The severity and duration of western spruce budworm defoliation, as well as site factors that influence moisture conditions effect the degree and direction of anatomical changes in the stemwood of Douglas-fir. This research fills a number of knowledge gaps by providing insights into the temporal and spatial dynamics of western spruce budworm outbreaks in central British Columbia over multiple centuries, and the plasticity of anatomical features in the stemwood of Douglas-fir during discrete western spruce budworm outbreaks. These research findings suggest that Douglas-fir forests are resilient to western spruce budworm outbreaks over space and time. / Graduate

Tree-ring studies have demonstrated that conifer latewood measurements contain information on long-term North American monsoon (NAM) variability, a hydroclimatic feature of great importance to plants, animals, and human society in the US Southwest. This paper explores data-treatment options for developing latewood chronologies aimed at NAM reconstruction. Archived wood samples for five Douglas-fir (Pseudotsuga menziesii, Mirb. Franco) sites in southeastern Arizona are augmented with new collections. The combined dataset is analyzed along with time series of regionally averaged observed precipitation to quantify the strength of regional precipitation signal in latewood time series and to identify ways of increasing the signal strength. Analysis addresses the signal strength influences of including or excluding ‘‘false’’ latewood bands in the nominal ‘‘latewood’’ portion of the ring, the necessary adjustment of latewood width for statistical dependence on antecedent earlywood width, and tree age. Results suggest that adjusted latewood width chronologies from individual sites can explain around 30% of the variance of regional summer (July–August) precipitation—increasing to more than 50% with use of multiple chronologies. This assessment is fairly insensitive to the treatment of false latewood bands (in intra-annual width and 𝛿¹³C variables), and to whether latewood-width is adjusted for dependence on earlywood-width at the core or site level. Considerations for operational chronology development in future studies are (1) large tree-to-tree differences in moisture signal, (2) occasional nonlinearity in EW-LW dependence, and (3) extremely narrow and invariant latewood width in outer portions of some cores. A protocol for chronology development addressing these considerations is suggested.

The fixed bed drying of western hemlock and Douglas-fir biomass particles at temperatures ranging from 50��C to 200��C and air velocities from 0.3 to 0.9 m/s was investigated. The objectives were to describe the drying characteristics of the particles, fit a model for thin-layer drying, and develop and test a deep bed drying model based on the thin-layer model. The effects of temperature and air velocity were determined in a bed approximately 1.3 cm in depth and a model for the drying curve was developed. The thin-layer model was then used to predict what would happen in a deeper bed. Model results were compared to drying curves measured in a 23-cm-deep bed. The deep bed model predicted both the experimental drying times and the moisture and temperature profiles in the bed. / Graduation date: 2013

Cloning of trees using somatic embryogenesis could have a major impact on tree breeding and commercial plantation forestry. To initiate somatic embryogenesis in Anatolian black pine (Pinus nigra Arnold. subspecies pallasiana), one-year old cones containing immature seeds were collected from eight trees located in METU campus, Ankara. Embryogenic tissues were derived from immature zygotic embryos excised from the seeds. The zygotic embryos at the time of collection were at the precotyledonary stage of development. For this study, Douglas-fir cotyledon revised medium (DCR) supplemented with 13.6&micro / M 2,4-D, 2.2&micro / M BAP, 0.5 g/L casein hydrolysate, 0.25 g/L L-glutamine and 3% sucrose was used. The media was solidified with 0.2% gelrite. Embryogenic tissue initiation was calculated for each genotype and collection date. Overall initiation frequencies were recorded as 0.92% for 2004 and 1.96% for 2005. Highest initiation frequency was calculated for 5-July 2005 sampling time (4.06). ANOVA revealed significant differences between trees and collection date for initiation frequencies. Also, ECL (Established cell lines) recorded after five subcultures. Overall, 0.38% and 0.62% of the initial explants were converted into ECLs for 2004 and 2005 respectively.

Thesis (Ph. D.)--North Carolina State University, 2004. / Title from PDF title page (viewed Mar. 27, 2005). Includes vita. Includes bibliographical references.

Final Report / Contract on 43-8371-4-628 / For: USDA, Forest Service, Southwestern Region / Regional outbreaks of western spruce budworms (Choristoneura occidentalis Freeman) have recurred at least three times in northern New Mexico since the early 1920's when the U. S. Forest Service first began systematic forest-pest surveys and documentation (Lessard 1975, U. S. Forest Service documents). The current outbreak was first noticed in a small area on the Taos Indian Reservation in 1974, and since then the defoliated areas have increased in New Mexico and Arizona to more than 370,000 acres of Federal, Indian, State and private lands (Linnane 1984). Losses in timber values can generally be ascribed to radial growth loss, height growth loss, topkilling, reduced regeneration, and mortality (Carlson et al. 1983, Fellin et al. 1983). A damage assessment project was initiated in 1978 and was aimed at obtaining measurements of some of these losses in budworm infested stands on the Carson National Forest, New Mexico (Holland and Lessard 1979). A large data base has subsequently been developed, including yearly measurements on topkilling, mortality, defoliation, and insect population changes (Stein 1980, 1981, Stein and McDonnell 1982, Rogers 1984). A growth assessment study was undertaken in 1982 to determine the feasibility of using dendrochronological methods to identify the timing of past outbreaks and to quantify radial growth losses associated with budworm defoliation (Swetnam 1984). Results of this work showed that three major outbreaks during the twentieth century were clearly visible in the tree-ring samples obtained from currently infested trees. The radial growth of host trees was corrected for age, climate and other non-budworm environmental effects, and then growth losses were computed as a percentage of expected growth (Swetnam 1984). Additional collections were obtained in 1984 in order to expand the scope of the radial growth study. The objectives included 1) assessment of a larger number of tree -ring samples, 2) comparison of radial growth losses between the two primary host species - Douglas-fir (Pseudotsuga menziesii) and white fir (Abies concolor), 3) comparison of radial growth losses between age classes, and 4) analysis of the relationship between yearly measurements of defoliation, insect populations and radial growth. This report summarizes the findings of the above analyses. Increment core samples from the 1982 collections are included here, therefore this report supersedes the earlier report (Swetnam 1984). Information is also presented on observations derived from the dated tree-ring series on the timing of occurrence of known and inferred spruce budworm outbreaks for the past 284 years (1700- 1983). This is the longest record of spruce budworm occurrence yet developed for western North America.

Western larch (Larix occidentalis Nutt.) is an important component of the montane forest in southeastern British Columbia. It grows in cool-temperate (IDF and ICH zones) and boreal climates (MS zone and, occasionally, ESSF zone). Larch is considered a very shade-intolerant species which can tolerate low light (partial shading) only during the seedling stage (the first 5 growing seasons). Typically, it regenerates after fire in the open on seedbeds exposed by burning. However, to what extent it tolerates low light and how various light environments affect its survival and growth is not known. The objective of our study was to determine the variation in survival and growth of western larch seedlings in relation to light availability and site conditions.

The influence of salal on tree growth has attained considerable attention in coastal British Columbia. Field observations, surveys, and studies in the CWH zone have indicated poor growth performance of crop tree species in salal-dominated plantations and natural immature and old-growth stands. Where sites have been burned and planted, tree growth has improved; similar effects have been observed for naturally regenerated stands. Immature stands that developed after wind disturbance or harvesting feature rapid growth and nearly complete absence of salal. As studies have shown that ericaceous plants negatively impact tree growth, the salal on potential harvest sites has been considered undesirable. This study examined (1) the possible influence of salal on the stand, soil nutrient status and site index, and (2) the relations between site index, salal, plant communities, and site in disturbed, immature, coastal Douglas-fir ecosystems. We compared vegetation and environmental characteristics of 101 ecosystems, and examined differences in foliar and soil nutrient characteristics and site index between stands with high and low salal cover through analysis of variance (ANOVA) and regression analysis.