Despite the recognized role of soil arthropod fauna on nutrient cycling and decomposition processes, many aspects of the effects of sylvicultural methods in forest ecosystems upon their biology remain poorly understood. The long term effects of prescribed fires on soil arthropods in forest ecosystems in the Pacific Northwest have never been studied. Soil samples were taken from three sites located in the Willamette National Forest in 1992: paired sites that were either clear-cut without burning and clear-cut with burning 40 years ago. One hundred and eight samples were processed; the arthropods were separated, identified and counted. To study the biology and behavior of some arthropods, eight species of oribatid mites were reared in laboratory conditions. Their life cycle, feeding behavior and reproduction were studied. Results indicated that there were no statistical significant treatment differences either in terms of total numbers of organisms or biomass. However, the majority of the commonest taxa did show offsetting treatment responses. A total of 204 taxa were found in the three sites. The most important groups included Collembola, mites, and insects. Other groups also represented, but in smaller numbers, were spiders, symphylans, pseudoscorpions, and centipedes. Of all these groups, oribatid mites was the best represented and appears to be a useful indicator of disturbances. / Graduation date: 1995

Monitoring wildlife habitats has become important to forest ecosystem management because it provides valuable information about the response of forests and their species to harvest practices, impacts from recreational use, conservation efforts, and natural and human-caused disturbances. Monitoring is a complex task that requires a variety of abiotic and biotic measurements and decisions about what should be measured, and when and where measurements should be taken. Riparian habitats contain unusually high diversity and are important to land managers. Wildlife assessments of riparian areas have focused on vertebrate species such as amphibians, birds, and mammals, but have largely ignored the arthropod components of the habitats. Arthropods constitute over 85% of all species and posses characteristics that make them valuable for tracking environmental changes. The purpose of this study was to gather site-specific data about epigaeic, riparian beetle community composition of the H.J. Andrews Experimental Forest (HJA). The patterns of beetle distribution, abundance, and diversity were analyzed and the results were used to characterize and compare the riparian and adjacent upslope beetle communities. Almost 8,000 beetle specimens representing about 250 species were collected from 141 pitfall traps placed along 10 transects in 3 different channel morphologies along Lookout Creek in the HJA. Traps were opened during six 30-day sampling periods over 2 years. Riparian and adjacent upslope beetle communities had high diversity measurements. The average difference of the calculated Simpson's Diversity Index between the two communities was 0.0116 and represented about 1% of the average riparian diversity. Analysis of species-curves indicated that the riparian habitats contained a higher total number of species. Multivariate Principal Coordinate Analysis indicated that the two habitats had distinctly different beetle communities. Multigroup Discriminant Analysis correctly classified 89.7% of the sampling units as the habitat group into which they were assigned a priori. Detailed recommendations for monitoring riparian habitats were discussed. / Graduation date: 2000

This study describes the composition of forest landscapes surrounding northern spotted owl (Strix occidentalis caurina) nests in the central Cascade Mountains of Oregon. I compared forest composition around 126 owl nests in 70 pair territories with forest composition around 119 points drawn randomly from all terrestrial cover-types, and around 104 points drawn randomly from the old-forest (closed canopy, > 80 yrs) cover type. All nest sites and random points were drawn from U.S. Forest Service lands and were not drawn from privately owned lands or Wilderness Areas. Forest cover was classified on a Landsat Thematic Mapper image. I quantified the percentage of old-forest within 200 concentric circular plots (0.04-5.0-km radii), centered on each analyzed point, using a geographic information system. I used logistic regression to make spatially-explicit inferences. Owl nests were surrounded by more old-forest when compared to points drawn randomly from all terrestrial cover types: there was significantly (P<0.05) more old-forest around the owl nests in plots as large as 1.79 km in radius. When compared to points drawn randomly from the old-forest cover type, owl nests were surrounded by significantly (P<0.05) more old-forest in plots with 0.17-0.80-km radii. Exploratory analyses suggest that the landscape scales most pertinent to northern spotted owl nest site positioning in this study area appear to be (in descending order): the surrounding 10-15 ha (~200-m radius), the surrounding 25-30 ha (~300-m radius), the surrounding 200 ha (800-m radius), and possibly the surrounding 700 ha (1,500-m radius). This study supports the assertion that northern spotted owls are strongly associated with older forests. The results also indicate that owl nests are most associated with higher proportions of old-forest near the nest implying that the arrangement of habitat is important for nest-site selection/positioning Since spotted owls in the central Cascade Mountains of Oregon are known to have home-ranges that average 1,769 ha, it is important to recognize that these results apply to nest-site selection/positioning on the landscape and not to the amount of habitat necessary for pair persistence or successful reproduction. / Graduation date: 1998

This thesis examines factors limiting understory herb presence and flowering in young second-growth Douglas-fir (Pseudotsuga menziesii) forests on the west side of the Cascade Mountains, Oregon, USA. I studied the belowground effects of canopy trees on understory herbs and shrubs in old-growth forests using trenched plots from which tree roots were excluded. Effects of tree density and stand age were tested by comparing the understory community composition of old-growth stands and pre-commercially thinned and unthinned young second-growth stands. I also examined the effect of conifer basal area on understory herb presence and flowering within one young second-growth watershed. In young stands, I focused on three groups of understory herb species: disturbance-responsive (release), forest generalist and old-growth associated. The effects of root trenching on vegetation and soil moisture were tested in closed-canopy and gap locations in two old-growth Douglas-fir (Pseudotsuga menziesii) forests. Ten years after installation, trenched plots averaged 92% total understory cover while untrenched plots averaged 47% cover. Trenched plots under closed canopies were moister than control plots throughout the growing season; the trenching effect on soil moisture became apparent in the generally wetter gaps only at the end of the growing season. Vegetation responses to trenching were concomitantly larger under closed canopies than in gaps. Stands that had been pre-commercially thinned 20 years earlier exhibited understory composition more similar to old growth than did unthinned stands. Thinned stands exhibited higher frequencies, abundances and density of flowering of old-growth associated herbs than did unthinned stands, but lower than did old-growth stands. Forest generalist and release species showed mixed responses to thinning. I used both general linear models and classification and regression tree models to explore the association of herb species presence and flowering with conifer basal area and abiotic variables. Both modeling approaches yielded similar biological insights. Flowering was more sensitive than presence to current stand basal area. Flowering of old-growth associated and release species was negatively correlated with conifer basal area. Linear models allowed clearer hypothesis tests, while tree-based models had greater explanatory power and provided information about interactions between variables. / Graduation date: 2004

Forest stands were studied to determine if old-growth forest structure could be mimicked in younger stands via overstory manipulation. Cover and species composition of understory plants were systematically sampled in sixteen thinned second-growth stands and sixteen adjacent unthinned second-growth Douglas fir (Pseudotsuga menziesii (Mirabel) Franco.) stands. The stands were thinned twenty-four to thirteen years ago. These were compared to seven nearby old-growth stands. Thinned and unthinned stands had matching elevations, aspect, and soils, yet differed primarily in management treatment. Leaf area indices were determined for these stands. Thinned stands differed from the old-growth and unthinned stands in having significantly higher cover values and species numbers, apparently resulting from increased light to the forest floor and a greater variety of microhabitats created by thinning. Young unthinned and old-growth stands were comparable in terms of cover and richness, but differed in species composition. Diversity indices showed no difference in species diversity between the three types of stands. Ordination of the species/sample data using Detrended Correspondence Analysis showed that understory species composition of the young unthinned and thinned stands was nearly identical. Species composition of old-growth stands differed from thinned and unthinned stands. The ordination indicated that age of the stands, structure of the canopy layers and climate were major determining factors in the species composition of the understory plant communities. Management manipulation of the second growth stands did not yield stands with understory vegetation communities that mimicked those of old-growth stands. The conclusions of this study were: 1) Shrub cover increased with thinning as compared to unthinned and old-growth stands. 2) Thinning increased the species richness of the stands, without increasing the number of exotics. 3) Diversity was not altered by thinning. Old-growth, thinned and unthinned stands did not differ in diversity values. 4) Patterns of community composition in thinned stands were more similar to unthinned equivalent stands than to nearby old-growth. / Graduation date: 1996

The structure and composition of mixed-conifer forest (MCF) in central Oregon has been altered by fire exclusion and logging. The resulting increased density, spatial contagion, and loss of fire resistant trees decrease the resiliency of this ecosystem to fire, drought, and insects. The historical and current composition and structure of MCF are characterized by steep environmental gradients and a complex mixed-severity fire regime. This inherent variation makes it difficult to determine the magnitude of anthropogenic effects and set objectives for restoration and management. As a result, there is a lack of consensus regarding how MCF should be managed and restored across the landscape. My primary research objectives were to: (1) Characterize the current structure and composition of MCF and how these vary with environmental setting; and (2) Characterize establishment and tree growth patterns in MCF in different environmental settings. To address these objectives, I collected field data on structure and composition and increment cores across a range of environmental conditions in MCF of the eastern Cascades and Ochoco Mountains. I used cluster analysis to identify four stand types based on structure and composition in the eastern Cascades study area and four analogous types in the Ochoco Mountains study area. Variation in understory composition and the presence of large diameter shade tolerant species distinguish each type. Stand types occupied distinct environmental settings along a climatic gradient of increasing precipitation and elevation. At relatively dry PIPO sites understories were dominated by ponderosa pine. At wetter PIPO/PSME and PIPO ABGC sites understories were dominated by shade tolerant species, but ponderosa pine was dominant in the overstory. At the coolest and wettest PIPO/PSME/ABGC sites understories were dominated by grand fir and shade tolerant species were common in the overstory. In the eastern Cascades current density of all live trees and snags was 432, 461, 570, 372 trees per hectare (TPH) for the four stand types identified. Stand types in the drier Ochoco Mountains were currently less dense at 279, 304, 212, and 307 TPH. Current MCF densities in both areas are 2-3 times higher than densities estimated for the late 19th and early 20th centuries from other studies in those two areas. Reconstruction of cuts in each stand type indicates that the density of large diameter ponderosa pine has been reduced by approximately 50% in all stand types in both study regions. Age histograms demonstrate that current density and composition of MCF stand types is a product of abrupt increases in tree establishment following fire exclusion in the late 19th century. The number of trees established increased after 1900 in all stand types, but the timing and composition of changes in establishment varied with climate. At dry PIPO sites increases in establishment were delayed until the 1920s and 1930s and were composed of ponderosa pine. At PIPO/PSME and PIPO/ABGC sites with intermediate precipitation, establishment was dominated by ponderosa pine prior to 1900, but after 1900 establishment was dominated by a large pulse of Douglas-fir and grand fir. At the wettest PIPO/PSME/ABGC there was less evidence of changes in structure and composition over time. My results indicate that compared to dry pine and dry-mixed conifer sites, relatively productive moist mixed-conifer sites were characterized by large changes in structure and composition. Such sites could be considered more ecologically altered by lack of fire than drier forest types that had high fire frequencies but slower rates of stand development and less plant community change. Radial growth patterns of cored ponderosa pines differed between the eastern Cascades and Ochoco Mountains. In the eastern Cascades mean growth rates and variance decreased during favorable climatic periods after 1900. This is likely related to increased competition, and provides evidence that current stand density lacks a temporal analog in the 18th and 19th centuries. Sensitivity of growth to climate and harvest suggest competition for water in the denser forest of the eastern Cascades, and indicates thinning will increase the diameter growth rate of large old pines. In the Ochoco Mountains, ponderosa pine tree growth was less responsive to climate prior to fire exclusion in the late 1800s, and growth did not respond to fire events. This suggests competition among trees was historically low in this region. After fire exclusion growth became more responsive to wet and dry climatic cycles, which may indicate that increased density and competition made trees more responsive to climate variability. Patterns of slow and fast growth appeared to differ between study regions and likely differ at the sub-regional scale. Further analysis of the relationship between growth and climate in different environmental settings is needed to distinguish where stand development has been modified by disruption of fire regimes. / Graduation date: 2013