Thesis (MSc)--Stellenbosch University, 2001. / ENGLISH ABSTRACT: The burgeoning problem of alien plant invasions in South Africa necessitates effective decision-making based on an understanding of the complex processes that govern these invasions. Due to the spatial nature of the problem, this study explored the use of Geographic Information Systems and spatial models for predicting the spread of alien vegetation and assessing the effectiveness of clearing strategies. The Spatially Explicit Individual Based Simulation (SEIBS) model was identified as a potentially useful tool for alien plant management. This thesis documents the further investigation and development that was necessary before recommendations could be made regarding the future use of the model. The landscape version of the SEIBS model was adapted to allow for the convenient input and output of spatial data, making it possible to simulate invasions in different areas. An ArcView extension was developed in order to facilitate the pre- and post-processing of the spatial data required and created by the model. Changes were also made to the fire routine of the model. The new version of the model was called Clear. A series of model tests for Pinus pinaster were conducted to assess the sensitivity of the Clear model to spatial resolution, initial spatial fragmentation and heterogeneity. These tests revealed that the model was sensitive to changes in resolution and needed to be reparameterised when using different resolutions. The initial level of fragmentation was shown to have a major influence on the invasion rate. Although greater levels of spatial heterogeneity with respect to vegetation age did not significantly affect the spread rate, it did increase the effectiveness of clearing strategies based on clearing juvenile or sparse vegetation. Based on these tests, it is concluded that the model can be readily applied to different areas, provided the influence of spatial characteristics is understood and accommodated. The Clear model was shown to be a useful tool for evaluating clearing strategies and for investigating invasion rates. It is recommended that the model be introduced to a wider audience, in order to obtain user feedback and further improve the accessibility of the model. / AFRIKAANSE OPSOMMING: Die toenemende probleem van uitheemse indringerplante in Suid Afrika, noodsaak effektiewe besluitneming wat gebaseer is op 'n begrip van die ingewikkelde prosesse wat indringing beheer. As gevolg van die ruimtelike geaardheid van die probleem, is die gebruik van Geografiese Inligtingstelsels en ruimtelike modelle vir die voorspelling van die verspreiding van indringerplante en die evaluasie van die effektiwiteit van opruimingstrategieë in hierdie studie ondersoek. Die Spatially Explicit Individual Based Simulation (SEIBS) model is as 'n moontlike geskikte hulpmiddel vir die bestuur van uitheemse indringerplante geïdentifiseer, alhoewel verdere ondersoeke en ontwikkeling nodig was voordat aanbevelings vir die gebruik van die model gemaak kon word. Vir hierdie studie is die landskapweergawe van die SEIBS model aangepas om die maklike toevoer en afvoer van ruimtelike data te fasiliteer. 'n ArcView uitbreiding is ontwikkel om met die voor- en naprosessering van ruimtelike data, wat deur die model gebruik en geskep is, te fasiliteer. Veranderinge is ook aan die vuur sub-roetine van die module gemaak. Die nuwe weergawe van die model word Clear genoem. 'n Reeks toetse is vir Pinus pinaster gedoen om die sensitiwiteit van die Clear model te toets teenoor ruimtelike resolusie, aanvanklike vlak van versnippering en vlak van heterogeniteit. Vanuit die toetse het dit geblyk dat die model sensitief was ten opsigte van verandering in resolusie en dat die model se parameters verstel moes word wanneer verskillende resolusies gebruik word. Daar is ook gewys dat die die vlak van aanvanklike versnippering 'n groot impak op die verspreidingstempo het. Alhoewel hoër vlakke van ruimtelike heterogeniteit teenoor plantegroei nie 'n merkbare impak op die verspreidingstempo gehad het nie, het dit wel die effektiwiteit van opruiming-strategieë, gebaseer op die opruiming van jong of yl verspreide plante, verbeter. Die gevolgtrekking wat uit die toetse gemaak kan word is dat die model geredelik op verskillende areas toegepas kan word, op die voorwaarde dat die invloed van ruimtelike eienskappe in ag geneem word en in berekening gebring word. Dit word aanbeveel dat die model wyer bekendgestel word om sodoende gebruikersterugvoer te bekom.

M.Tech. (Fine Art) / Unemployment and the crime levels which escalate as a result of it, are very realistic social problems in South Africa and training for job skills and entrepreneurship has become a priority. This study identifies the use of invasive plant fibres as a viable resource for the hand papermaking industry and the development of a new cultural industry. The research' funded by the National Research Foundation (NRF) and the Technikon Research Committee (TRC) has been implemented in an established poverty relief programme called Phumani Paper', which creates jobs especially in rural or impoverished sectors through technology transfer and training. A curriculum is currently being developed for current and future participants of this programme to obtain qualifications in hand papermaking. An investigation has been undertaken to define suitable invasive plant species for the purpose of this research. The investigation included the extent of invasive encroachment, plant identification, methods of control and government legislation. Initially water hyacinth (eichhomia crassipes) was identified as a suitable fibre, but because Phumani Paper supports 21 small paper businesses in seven provinces, research had to be undertaken on several different species found in each of the regions. Species such as moth catcher (Araujia sericifera), milkweed (Asclepias fruticosa), pampas grass (Cortadeira selloana), giant reed (Arundo donax) and bugweed (Solanum mauritianum) have been identified as viable fibres for the hand papermaking industry, due to their fibrous characteristics...

The objective of the study was to assess the farmers’ knowledge and perceptions of P. incana invasion and impacts on rangeland, livestock production and livelihood in four communal grazing areas. Twenty households per village that own livestock were randomly selected to conduct an interview using an open and closed ended questionnaire. In each household, an old and knowledgeable respondent about P. incana invasion was used as a selection criterion. For scientific assessment of range condition, the experiment was conducted in a 270 m x 100 m trial plots that were established in 2014. The area was divided into 3 open and 3 fenced subplots each having an area of 45 m x 100 m established laid down a slope gradient. Each plot was divided into four equal subplots to apply the control treatments. Therefore, the trial layout was a stratified block experiment of four treatments replicated three times. The four control treatments applied to P. incana invasion were chopping and resting (CR), resting without chopping (RWOC), grazing and chopping (GC), grazing without chopping (GWOC). A step point method was applied in each transect to monitor species composition and P. incana cover. On the fenced plots two exclosures of 1mx1m size were randomly distributed to make sure that resting is maintained. Within exclosures 0.25 m2quadrates in each were put randomly to measure biomass. Density of P. incana was determined from 5 m x 5 m quadrates that were placed randomly within each sub-plot. The density and height of individual shrub plant was measured. All rooted live woody plants were recorded and counted in each quadrate. The results revealed that species composition was not affected (P>0.05) by the treatments applied. The herbage height and basal cover had greater values (P<0.05) under rested plus chopping (CR) treatment followed by grazed plus chopping (GC). Grass biomass production and essential nutrients showed a greater concentration (P<0.05) under CR treatment than any treatments applied. The results on perceptions revealed that goats had the highest mean household holding number than any other livestock species across the selected villages in all the villages. Respondents indicated that both goats and cattle had equal or more importance than sheep. Three out of four villages raised cattle primarily for income generation and secondarily for cultural purposes like slaughtering during weddings ceremonies, amazila and also during woman circumcisions (Ntonjane). Similarly, goats and sheep were primarily raised for cash income generation. The results revealed that livestock population decreased over ten to twenty years (one-two decades) and poor rangeland condition is assumed to be the cause of the situation as feed is the key factor for production. The area was dominated by the grasses rather than by browse woody species so the above results are possible as the P. incana is more competitive than grasses. It was concluded that P. incana is detrimental to both rangeland and animal production. Forage and animal performance as well as farmers’ economic status (livelihoods) were negatively affected by P. incana invasion and the scientific assessment reported it to be detrimental to agriculture. Thus chopping and resting (CR) treatment resulted in a greater improvement of the basal cover, biomass production, soil nutrients as well as grass height than the other treatments applied. Therefore it can be concluded that chopping and resting the veld invaded with P. incana can improve the range and consequently animal production.

Invasive tree species pose a huge problem in the Garden Route and are particularly damaging to aquatic ecosystems, including wetlands, riparian zones, lakes and estuaries. Therefore, this study aimed to determine priority areas for invasive tree species management, with a focus on aquatic ecosystems. This was achieved by using existing literature to identify priority species, based on their impact on aquatic ecosystems and their associated ecosystem services, and then testing the suitability of SPOT-6 and WorldView-3 multispectral data at detecting these focal species. The priority species identified were: Acacia cyclops (rooikrans), Acacia longifolia (long-leaved wattle), Acacia mearnsii (black wattle), Acacia melanoxylon (blackwood), Acacia saligna (Port Jackson willow), Eucalyptus camaldulensis (red river gum), Pinus pinaster (cluster pine) and Pinus radiata (radiata pine). The Random Forest classifier on SPOT-6 data achieved an overall accuracy of 62.5% and this method was consequently deemed ineffective at separating invasive tree species from other tree species in the Garden Route. The overall accuracy of the WorldView-3 classifier was higher (78.9%) but the cost of the data limited the use of more images for the detection of the focal species throughout the Garden Route. Therefore, to identify priority areas for invasive tree management, criteria derived from existing literature were input into spatial conservation planning software. The analysis identified the: Saasveld section of the Garden Route National Park, the Wilderness Lakes, Knysna Forest, Knysna Estuary, Tsitsikamma Forest around Stormsriver and a disturbed area of fynbos southeast of Kareedouw as management priorities. Currently spatial conservation planning software proved to be cost-affordable and useful tool and is recommended for invasive tree management in the Garden Route.

Magister Scientiae - MSc (Biodiversity and Conservation Biology) / Arundo donax is an invasive species that mostly impacts on sensitive riparian ecosystems. Arundo is also invasive in South Africa, though less is known about its ecology, biology, and impacts. Since California and the Western Cape of South AFrica have similar Mediterranean-type climates, we could assume that the impacts of Arundo on ecosystems in California are likely to be similar in the Western Cape, and that control methods used could be extrapolated for use in South Africa. This thesis attempted to determine what impact Arundo has on biodiversity at the community-ecosystem level. / South Africa

The alien invader plant Parthenium hysterophorus is a Category 1 weed in South Africa, where it poses a serious threat to indigenous vegetation in particular, and to biodiversity in general. In addition to its competitive ability, it is hypothesized that the successful invasiveness of P. hysterophorus is linked to the allelopathic potential of the plant. One compound in particular, parthenin, is alleged to play a major role in this allelopathic potential. Interference between P. hysterophorus and three indigenous grass species (Eragrostis curvula, Panicum maximum, Digitaria eriantha) was investigated on a site with a natural parthenium infestation at Skukuza, Kruger National Park. The trial was conducted over two growing seasons on exclosure plots which eliminated mammal herbivory. P. maximum displayed best overall performance and was eventually able to completely overwhelm P. hysterophorus. Eragrostis curvula and D. eriantha grew more favourably in the second season after becoming better established but were clearly not well adapted to the trial conditions. Although P. maximum was the supreme interferer, all grasses were able to significantly interfere with P. hysterophorus growth in the second season. The ability of P. maximum to interfere with P. hysterophorus growth so efficiently that it caused mortalities of the latter species, indicates that P. maximum exhibits high potential for use as an antagonistic species in an integrated control programme. An investigation on the production dynamics of parthenin in the leaves of P. hysterophorus indicated that high levels of this compound are produced and maintained in the plant up until senescence. The high resource allocation priority of the plant towards this secondary metabolite even in the final growth stages may indicate the use of residual allelopathy to inhibit or impede the recruitment of other species. Studies on the persistence of parthenin in soil revealed that parthenin is readily degraded in soil and that microbial degradation appears to play a predominant role. Significant differences between parthenin disappearance-time half-life (DT50) values were observed in soils incubated at different temperatures and in soils with different textures. Exposure of the three grass species to pure parthenin showed that, in terms of their early development, the order of sensitivity of the grasses was: Panicum maximum>Digitaria eriantha>Eragrostis curvula. It may therefore prove challenging to establish P. maximum from seed in P. hysterophorus stands during the execution of an integrated control programme due to the sensitivity of this grass species to parthenin. From the research findings it appears possible that P. hysterophorus can inhibit or impede the recruitment of indigenous vegetation under natural conditions. At least one mechanism through which this alien species can exert its negative influence on other plant species is the production and release of parthenin. / Dissertation (MSc (Agric) Agronomy)--University of Pretoria, 2007. / Plant Production and Soil Science / unrestricted

It is well known that invasive alien plants (IAPs) pose a significant threat to natural biodiversity and human well-being. Through various pressures exerted on natural ecosystems, IAPs decrease and alter natural processes that provide important ecosystems services and livelihoods to human communities. These plants also displace and out-compete natural plant communities in the areas in which they invade. Management of IAPs is a complex issue with social, environmental and financial challenges. The long-term sustainable management of IAPs requires a management approach that monitors and measures the outcomes of current management practices, and considers the lessons learnt in future decision making. Adaptive management is such an approach and is characterised by monitoring, assessment, reflection and adaptation. In this dissertation I explore IAP management within the South African National Parks (SANParks) organisation. I focus on the quality of the available IAP clearing data for monitoring and environmental decision-making and explore how strategic adaptive management (SAM) is being applied to IAP management. Strategic adaptive management is a form of adaptive management that SANParks adopted to deal with the inherent unpredictability of the social ecological systems they manage, and is primarily a learning-by-doing approach. In chapter one I describe the various impacts that IAPs have on natural environments. I also discuss management of IAPs, the national Working for Water (WfW) programme and how SANParks control IAPs in partnership with WfW. Clearing of IAPs within the borders of SANParks is funded by the Department of Environmental Affairs‘ (DEA) Natural Resource Management programmes and carried out through the Working for Water programme, under SANParks‘ Biodiversity Social Projects programme and is done on a contract basis. Contract data are stored in the Working for Water Information Management System (WIMS) database. Although IAP clearing is well established, the efficiency of overall clearing operations within the parks, and quality and reliability of the data available in the WIMS, needs to be examined. In Chapter two, I (i) assess the status of IAP clearing and the direct clearing costs in each of the five national parks (costs are inflated to 2012 equivalents using the consumer price index), (ii) assess the accuracy of workload estimations for WfW IAP clearing, and (iii) discuss the utility of WIMS as a system to collate and store accurate data for monitoring purposes, focusing on the integrity and reliability of the data, the accuracy of density estimations and whether it can inform sound decision making. The planning of future clearing contracts lacks efficiency due to inaccuracies and unreliability of input data, specifically estimates of invasive alien plant density. The accuracy of density estimation and cost of clearing varies greatly between parks and does not appear to reflect the reality on the ground. In Chapter 3, through an interview process with IAP managers, I 1) unpack the steps in the SAM ‗cycle‘ and investigate the effectiveness of implementation of each step according to respondents; 2) assess the barriers that are currently keeping managers from implementing effective IAP management; 3) discuss whether learning and adaptation is taking place, and 4) discuss how the SAM process can be modified to promote its effectiveness.

Thesis (PhD (Conservation Ecology and Entomology))--University of Stellenbosch, 2009. / ENGLISH ABSTRACT: Natural and semi-natural ecosystems and human communities worldwide are under siege from a growing number of destructive invasive alien species. Alien species are those whose presence in an area is due to intentional or accidental introduction as a result of human activities. Some alien species become invasive, and some cause tremendous destruction to the ecosystem and their stability, but we do not yet understand fully the many factors that determine the levels of invasiveness in alien species. However, management of alien plants requires a detailed understanding of the factors that make them invasive in their new habitat. The aim of this study was to explore in detail the processes and potential for invasion of Schinus molle (Peruvian pepper tree) into semiarid savanna in South Africa and to examine the potential for this species to invade further in these ecosystems, and in other South African biomes. In this thesis I explored the patterns and processes of invasion of S. molle in semiarid savanna using small-scale experiments to investigate physical and ecological barriers to invasion that prevent or accelerate the invasion of this species. I examined factors such as pollination; seed production; seed dispersal; seed predation and viability, all known to contribute to invasiveness. I highlighted the critical role of microsite conditions (temperature, humidity, water availability) in facilitating S. molle seedling establishment in semi-arid savanna and demonstrated that microsite type characteristics need to be considered for management and monitoring of the species in South Africa. I demonstrated the ability of S. molle to out-compete indigenous woody plants for light and other resources and also showed that disturbance of natural ecosystems was not a prerequisite for invasion, although human activities such as tree planting have played a major role in disseminating this species in South Africa. Predicting the future distribution of invasive species is very important for the management and conservation of natural ecosystems, and for the development of policy. For this reason, I also assessed the present and potential future spatial distribution of S. molle in South Africa by using bioclimatic models and a simulation-based spread model. I produced accurate profiles of environmental conditions (both biophysical and those related to human activities) that characterize the planted and naturalized ranges of this species in South Africa, by linking species determinants, potential habitat suitability and likely spread dynamics under different scenarios of management and climate change. All those components provided insights on the dynamics of invasions by fleshy-fruited woody alien plants in general, and on S. molle invasions in South Africa in particular. I developed a conceptual model that described S. molle population dynamics leading to an understanding of the processes leading to the invasive spread of this species in South Africa. This work also emphasized the need for policy review concerning the invasive status of S. molle in South Africa, and recommendations are made for future research. / AFRIKAANSE OPSOMMING: Die natuurlike and half-natuurlike ekosisteme sowel as menslike gemeenskappe wereldwyd word bedreig deur ‘n groeiende hoeveelheid indringerplantspesies. Indringerplantspesies (daardie spesies wie se teenwoordigheid toegeskryf kan word aan opsetlike of toevallige inbringing deur menslike toedoen) is ‘n bedreiging nie net vanweë die massiewe verwoesting van die ekosisteme en ekosisteemstabilitiet nie, maar ook omdat ons nog nie ten volle verstaan hoe hulle van skaars in hul natuurlike omgewing tot dominant in hul nuwe habitat gaan nie. Bestuur van indringer plante vereis ’n begrip van biologise en ekologiese faktore wat lei tot hulle indringing in die nuwe habitat. Die primêre doel van hierdie studie was om in detail uit te vind wat die prosessesse en potensiaal is vir die indringing van S. molle (die Peruviaanse peper boom) in droë savanna en om indringingspatrone in droë savanna met huidige en potensieële toekomstige patrone in ander Suid-Afrikaanse biome te vergelyk. In hierdie tesis ondersoek ek die patrone en prosesse wat verband hou met die indringing van S. molle in droë savanna deur gebruik te maak van kleinskaalse eksperimente om fisiese en ekologise hindenisse te ondersoek wat indringing van S. molle in Suid-Afrika voorkom of versnel. Dit sluit faktore in wat bekend is om by te dra tot indringing van plant spesies, soos bestuiwing, saadproduksie, saadpredasie en kiemkragtigheid. Ek het die kritieke rol beklemtoon van mikroomgewingskondisies (temperatuur, humiditeit, waterbeskikbaarheid) in die fasilitering van S. molle saailingvestiging in droë savanna en het gedemonstreer dat die tipe mikroomgewingskarakteristieke in ag geneem moet word by betuur en monitering van die verspreiding van S. molle in Suid-Afrika. Ek het die vermoë van S. molle om inheemse plante te uitkompeteer gedemonstreer, en het gewys dat versteuring van natuurlike ekosisteme nie ’n voorvereiste vir S. molle indringing was nie, hoewel menslike aktiwiteite soos boomaanplantings ’n groot rol speel deur by te dra tot indringing van hierdie spesie in Suid-Afrika. Voorspelling van toekomstige verspreiding van indringerspesies is baie belangrik vir die bestuur en bewaring van natuurlike ekosisteme, sowel as vir ontwikkeling van wetgewing. Daarom is die huidige en potensiele toekomstige ruimtelike verspreiding van S. molle in Suid-Afrika bereken deur inkorporering van bioklimaatsmodelle en simulering gebasseer op ’n verspreidingsmodel. Ek het derhalwe akkurate profiele van omgewingstoestande (beide fisiese en daardie wat verband hou met menslike aktiwiteite) wat die aangeplante en natuurlike omvang van die spesie in Suid-Afrika kenmerk geproduseer deur spesiedeterminante, potensieële geskiktheid van habitatte en moontlike verspeidingsdinamika onder verskillende bestuursscenarios en kimaatsverandering te koppel. Al hierdie komponente verskaf insig in die dinamika van die indringing van houtagtige plante met vlesige vrugte oor die algemeen en S. molle in besonder in Suid- Afrika. Hierdie werk beklemtoon ook die behoefte vir hersiening van beleidsrigtings wat betrekking het op die indringerstatus van S. molle in Suid-Afrika en maak aanbevelings vir toekomstige navorsing.

The change in total evaporation through alteration of vegetative cover is a major influence on catchment hydrology. The transformation of grassland and scrub habitats to commercial tree plantations, as well as the uncontrolled spread of invasive alien plants (lAPs) to ecologically sensitive systems, riparian zones in particular, are a threat to biodiversity and integrity of natural systems. Furthermore, critical low flow periods are of particular concern to water managers and local communities, as well as the associated impacts of potentially compromised water resources for rural livelihoods. The Working for Water (WfW) programme was implemented in 1995 by the Department of Water Affairs and Forestry and its main goals are to remove lAPs in order to improve water supply while at the same time providing employment to marginalised communities. In this study, the hydrological response to lAPs in the Upper Blyde River catchment is assessed. This is done by developing a classification structure for lAPs as a land use using detailed mapping available from WfW for use in a hydrological model, and then configuring and running the ACRU hydrological model for the Upper Blyde River catchment in Mpumalanga. In the classification, lAPs are represented as spatially explicit land use units in the ACRU model according to the type of habitat they invade, viz. riparian or non-riparian; as well as by type of plant, i.e. tree or shrub; and their area and density. The results obtained from simulating catchment hydrological responses using the ACRU model indicate that riparian lAPs have a great er impact on streamflow than do landscape invasions alone, specifically during periods of low flow. An increase in streamflow after removing lAPs from riparian and non-riparian habitats is a consistent outcome at both subcatchment and catchment scales. Using a spatially explicit method in order to model the hydrological response of different types of lAPs for different density classes in both riparian and non-riparian habitats is found to be a useful technique in determining the degree to which lAPs influence catchment streamflow. Recommendations for future research include focussing hydrological assessments of lAPs on critical flow periods and their impacts on water quality; investigation into the water use of invasive and indigenous vegetation for more accurate estimates from modelling exercises; and finally, applying the classification system for lAPs with other land use sensitive hydrological models for validation, and their wider application by incorporating methodologies into guidelines for use by WfW at national and provincial level. / Thesis (M.Sc.)-University of Natal, Pietermaritzburg, 2003.

Extensive areas of the Eastern Cape Province have been invaded by Pteronia incana (Blue bush), a non-palatable patchy invader shrub that is associated with soil degradation. This study sought to establish the relationship between the invasion and a range of eco-physical and land use gradients. The impact of the invader on soil moisture flux was investigated by comparing soil moisture variations under grass, bare and P. incana invaded surfaces. Field based and laboratory spectroscopy was used to validate P. incana spectral characteristics identified from multi-temporal High Resolution Imagery (HRI). A belt transect was surveyed to gain an understanding of the occurrence of the invasion across land use, isohyetic, geologic, vegetation, pedologic and altitudinal gradients. Soil moisture sensors were calibrated and installed under the respective surfaces in order to determine soil moisture trends over a period of six months. To classify the surfaces using HRI, the pixel and sub-pixel based Perpendicular Vegetation Index (PVI) and Spectral Mixture Analysis (SMA) respectively were used. There was no clear trend established between the underlying geology and P. incana invasion. Land disturbance in general was strongly associated with the invasion, as the endemic zone for the invasion mainly comprised abandoned cultivated and overgrazed land. Isohyetic gradients emerged as the major limiting factor of the invasion; a distinct zone below 619mm of mean annual rainfall was identified as the apparent boundary for the invasion. Low organic matter content identified under invaded areas was attributed to the patchy nature of the invader, leading to loss of the top soil in the bare inter-patch areas. The area covered by grass had consistently higher moisture values than P. incana and bare surfaces. The difference in post-rainfall moisture retention between grass and P. incana surfaces was significant up to about six days, after which a near parallel trend was noticed towards the ensuing rainfall episode. Whereas a higher amount of moisture was recorded on grass, the surface experienced moisture loss faster than the invaded and bare surfaces after each rainfall episode. ii There was consistency in multi-temporal Digital Number (DN) values for the surfaces investigated. The typically low P. incana reflectance in the Near Infrared band, identified from the multi-temporal HRI was validated by field and laboratory spectroscopy. The PVI showed clear spectral separability between all the land surfaces in the respective multi-temporal HRI. The consistence of the PVI with the unmixed surface image fractions from the SMA illustrates that using HRI, the effectiveness of the PVI is not impeded by the mixed pixel problem. Results of the laboratory spectroscopy that validated HRI analyses showed that P. incana’s typically low reflectance is a function of its leaf canopy, as higher proportions of leaves gave a higher reflectance. Future research directions could focus on comparisons between P. incana and typical green vegetation internal leaf structures as potential causes of spectral differences. Collection of spectra for P incana and other invader vegetation types, some of which have similar characteristics, with a view to assembling a spectral library for delineating invaded environments using imagery, is another research direction.