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Recent research has challenged the long-standing hypothesis that forests in the Upper Midwest of the United States developed during wetter periods and retreated during dry periods. We explored this debate by examining patterns of tree establishment on an oak savanna in east-central Minnesota within the context of variable moisture availability and fire suppression. We used superposed epoch analyses (SEA) to evaluate the mean moisture conditions for a 21-year window surrounding tree establishment dates. Before effective fire suppression (1809–1939), 24 of 42 trees with pith dates (62%) grew to 30-cm height during dry years (Palmer Drought Severity Index < -1), versus only 5 of 42 (12%) that established in wet years (PDSI > 1). Significantly more trees established during dry periods (negative PDSI values) than would be expected with the proportion of wet-to-dry years (x²= 10.738, df = 1, p-value = 0.001). Twenty of the complete sample of 74 trees with pith dates (27%) established during drought in the 1930s. We hypothesize that dry conditions limited plant productivity, which in turn decreased competition between grasses and tree seedlings and reduced rates of accumulation of fine fuels, enabling seedlings to grow tall enough to resist subsequent fires. We recommend SEA as a methodological approach to compare historical climate conditions with the timing of regeneration success in other regions of forest expansion.

The nuclear behavior, vegetative incompatibility, and induction of basidiocarp formation and basidiospore germination of Inonotus andersonii was investigated. Safranin O-KOH and Hematoxylin stains were used to determine the number of nuclei in basidiospores, vegetative hyphae, and basidiocarp tissue. Both uni- and binucleate basidiospores were found in the tubes of basidiocarp tissue. The nuclei in the hyphal cells of presumptive heterokaryons typically numbered some multiple of two per cell which suggests that division is conjugate. Subhymenial hyphae and immature basidia were often uninucleate but tramal hyphae were multinucleate. Antagonistic hyphal interactions developed between different vegetative isolates on both 2% MEA and oakwood test blocks. Self-crosses were compatible. Basidiocarps formed in 12 wk on 2% MEA medium containing ground oakwood and wheat. Most attempts to induce spore germination failed. However, basidiospores obtained at the end of this study from actively sporulating basidiocarps germinated on 2% MEA.

Dendrochronologists have used the presence of spiral grain as an indicator of old trees for most of the history of the field, although this relationship has been little studied. We examined cross-sections from dead trees and used a 12-mm Haglof Swedish Increment borer to collect cores from living white oak (Quercus alba L.) trees in an Eastern Deciduous Forest stand in southwestern Illinois. Spiral grain is the alignment of wood fibers to the longitudinal axis of trees and is driven by patterns of initial cambial cell division. In this study, we examine environmental factors that may affect spiral grain severity, the usefulness of non-destructive sampling methods (using the 12-mm increment borer), and the relationship between tree age and spiral grain. We tested Brazier’s method (1965) of averaging the spiral grain angle from two radii taken 180 degrees apart (i.e. one diameter in the tree) to get representative grain angles for the whole circumference of a tree at a certain height. The 12-mm increment borer did not produce consistent results in this study; therefore, the collection of cross-sections is advised for the study of spiral grain in white oaks. Brazier’s method should not be used in white oaks and should not be applied universally to all tree species. The severity of spiral grain is expressed in the xylem and may not be expressed in the bark of the tree. Left spiral grain does generally increase in white oaks with age, although this relationship is not always consistent, so a tree without severe spiral grain is not necessarily young.

This thesis contributes to the understanding of two of the most intriguing questions that forest ecologists have faced over recent decades: 1) how high diversity is maintained in species-rich ecosystems; and 2) what is the role of spatio-temporal environmental variation in structuring forest communities. The aims of the research were to ascertain how species composition varies both spatially and temporally and how changes in the vegetation can be understood in the context of species coexistence theories (niche versus neutral). A group of 38 sympatric species, including 9 species of Quercus, on which little ecological research has been undertaken, were used in this study. The data used in this project include eleven years of periodic remeasurements of permanent plots established in high-altitude oak forests in Mexico. Adult, sapling and seedling trees were studied as well as their environmental surrounding. Spatial and temporal variations in forest composition were analysed using multivariate statistical approaches. The results show that there are discrete communities in these mixed oak forests that correspond to specific environments. At a broad scale the study area can be classified into two floristic zones, a mesic zone characterised by associations that include Quercus candicans, Q. laurina and Q. castanea and; a xeric zone dominated by Q. crassipes. However of a finer scale of analysis important variation in composition was associated with different life stages of the trees, with adult trees showing much stronger environmental associations than seedlings and saplings. Successional pathways and rates vary at relatively fine scales. This may be as a result of dominance alternation between dominant canopy species. Micro-niche zonation processes caused by a high degree of environmental heterogeneity combined with individual species traits explain the coexistence of phylogenetically similar sympatric Quercus species. A hierarchy of processes, each acting at a different spatial and temporal scale, determines species diversity and coexistence. The overall findings support the idea that niche differentiation rather than chance events such as dispersal limitation, are more important in permitting species coexistence.

Sixty individual ring-width series of oak (Quercus robur L.) from six sites in the northwestern Iberian Peninsula, ranging from 50 to 120 years, were grouped using hierarchical cluster analysis with different types of distance measures. Euclidean distances as well as other linkage distances based on statistics used to crossdate tree-ring series (Gleichläufigkeit and coefficient of correlation with its corresponding t-value) were compared. In addition, a new distance measure based on a corrected inversion of the Student’s t is proposed in the present paper, which takes into consideration the number of years used for series comparison. The Euclidean distances, commonly used in ecological analyses, inefficiently identified homogeneous units of trees based on their ring-width patterns. Among crossdating statistics, the correlation coefficient was more effective than Gleichläufigkeit, but the most satisfactory results were obtained when 1/t was used as distance measure. Finally, these methods of cluster analysis have been implemented into a computer program for future use of the dendrochronological community.

In this thesis, the behaviour of glued laminated timber combined with hardwood and softwood lamallae is investigated. The influence of hardwood in the tension and compression zone, in terms of strength and stiffness is evaluated.The basis of evaluation consists of determining the behaviour of beams with various combinations of hardwood solely in the tension zone along with beams with hardwood in the tension and compression zone. The influence of different amount of hardwood for both cases is studied my means of experimental and analytical methods.Experimental data attained by performing bending tests are evaluated for different combinations made from spruce and oak. By comparing the experimental and analytical data an increase in the strength and stiffness in various combinations is observed and potrayed which varies based on different wood species.

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The sudden oak death pathogen, Phytophthora ramorum, is present in southwestern Oregon, and while an eradication effort is underway, the potential impact of the polyphagous pathogen on surrounding vegetation is unknown. Plant communities in the area are substantially different from those affected in California, although tanoak (Lithocarpus densiflorus), evergreen huckleberry (Vaccinium ovatum) and Pacific rhododendron (Rhododendron macrophyllum) are hosts found in both areas. Other species are likely to be susceptible to the pathogen. Detached leaf and whole plant assays were used to test species commonly associated with tanoak, as well as three rare or endemic ericaceous species of the western Siskiyou Mountains and their associated taxa. Leaves and plants were challenged with zoospore suspensions that were capable of generating symptoms in the known hosts tanoak and evergreen huckleberry. Most (78%) of the previously unchallenged species developed necrotic lesions in detached leaf assays although severity (% leaf area necrotic) was variable. All three of the ericaceous species of conservation concern: Arctostaphylos hispidula, Kalmiopsis leachiana, and Leucothoe davisiae were susceptible in detached leaf assays. Factors important in determining whether or not these species will become infected in their native habitat are discussed. / Graduation date: 2004