Made available in DSpace on 2015-03-05T16:20:40Z (GMT). No. of bitstreams: 0 Previous issue date: 27 / Nenhuma / A conversão de florestas naturais em monoculturas arbóreas vem formando mosaicos heterogêneos na paisagem, onde é desconhecida a capacidade de colonização das espécies vegetais do sub-bosque. Essa mudança no uso da terra leva a modificações na composição de espécies de plantas, podendo alterar o seqüestro de carbono em florestas. Este estudo procurou caracterizar diferenças na composição, riqueza de espécies, riqueza funcional e abundância de plantas que colonizam o sub-bosque de florestas e monoculturas arbóreas, investigando também como a diversidade de espécies e diversidade de grupos funcionais podem influenciar no acúmulo de biomassa no sub-bosque. O estudo foi conduzido na Floresta Nacional de São Francisco de Paula, RS, Brasil, em três repetições de cada um dos seguintes ambientes: áreas de monoculturas de Pinus, Eucalyptus e Araucaria angustifolia e Floresta Ombrófila Mista. Em cada uma das 12 áreas de estudo, foram demarcadas 5 parcelas de 8m² e 5 parcelas de 1m³. Foram realizados levantamentos de ri / The conversion of native forests to tree monocultures have been producing an heterogeneus landscape mosaic where the capacity of colonization of understorey plants from adjacent forest is still unknown. These changes in land use may lead to modifications on plant species composition which can alter forest carbon storage. These study investigates differences in species composition, species richness, abundance and functional richness of plants that colonized the understorey of forests and tree monocultures. It also investigates how species diversity and the diversity of functional groups could influence biomass storage in the understorey. The study was conducted at the São Francisco de Paula National Forest, RS, Brazil, using three replicates of each of the following habitas monocultures of Pinus, Eucalyptus and Araucaria angustifolia and Araucaria forest. In each of the 1 study sites five 8m² plots and five 1m³ plots were defined. Richness, abundance and biomass storage of plant species were surveyed and sepa

As pastagens nativas possuem grande importância na produção de produto animal e nos serviços ecossistêmicos. Muitos estudos realizados sobre esse recurso natural demonstram seu potencial produtivo. Este é diretamente ligado ao manejo utilizado, e a oferta de forragem adotada exerce forte influência tanto na produção primária quanto secundária. Em resposta disso também se sabe que a composição da vegetação responde de maneira positiva ou negativa ao regime de desfolha. Entretanto, uma abordagem mais holística relativa à essa resposta e ao uso do diferimento ainda era uma deficiência desta área do conhecimento. No sentido de supri-la, foi desenvolvido o presente trabalho resultando em dois artigos. Ambos se valem do uso de grupos de espécies para caracterização dos ambientes. O primeiro aborda o padrões de diversidade e o comportamento da vegetação após 28 anos de regimes de desfolhas contrastantes mediada pelas ofertas de forragem 4 kg de MS por 100 kg de PV e 8-12 (8 primavera e 12 no restante do ano) kg de MS por 100 kg de PV. Não houve diferença na riqueza e diversidade entre os ambientes, mas verificou-se resposta negativa na composição de espécies na área com sobrepastejo: espécies que são menos eficientes como forrageiras e na proteção do solo, as ruderais, sucederam as espécies conservadoras de recurso que proporcionam maior estabilidade na disponibilidade de forragem e são mais eficientes na proteção do solo. Já quando a intensidade de pastejo foi moderado houve predominância de espécies conservadoras. O segundo artigo testa duas épocas de diferimento nos dois ambientes citados anteriormente objetivando avaliar a vegetação perante esses tratamentos com a hipótese que o diferimento aumenta a contribuição de espécies hibernais e as demais espécies de interesse forrageiro no curto e médio prazo. Esse segundo artigo conclui que o diferimento no curto prazo não foi suficiente para aumentar a contribuição de espécies hibernais, porém aumentou o número de inflorescências dessas espécies que reforçam o banco de sementes do solo, podendo-se inferir que tais espécies aumentem sua contribuição no longo prazo. No curto (seis meses) e médio prazo (dois anos) o redirecionamento da composição florística por efeito de diferimento é dado pelo aumento de espécies cespitosas e conservadoras de recursos no ambiente degradado por longo tempo de sobrepastejo. O uso de grupos de espécies mostra-se como indicadores adequados para caracterizar o grau de herbivoria exercido em pastagens complexas. / Rangelands have great importance in the animal production and ecosystem services. Many studies on this natural resource demonstrate their productive potential, which is a direct consequence of management. The adopted herbage allowance exerts strong influence on primary and secondary production through changes in botanical composition. However, a more holistic approach on interactions and responses of vegetation to grazing and the use of deferment still a presents shortcoming of this area of knowledge. The present work was developed to contribute in filling these gaps, resulting in two papers. Both avail themselves on the use of species groups for characterization of environments. The first addresses the diversity patterns and behavior of vegetation after 28 years of contrasted defoliation regimes mediated by herbage allowance of 4 kg DM per 100 kg BW and 8-12 (8 spring and 12 in the rest of the year) kg DM per 100 kg BW. There was no difference in the richness and diversity between environments but there was a negative response in species composition in the overgrazed area: species that are less efficient as forage and soil protection, the ruderals, succeeded conservative resource species that provide greater stability in the availability of forage and are more efficient in protecting the soil. When grazing intensity was moderate conservative species predominated. The second article seeks to verify effects of two periods (spring or fall) of deferment in both environments that already were studied in the first chapter, with the hypothesis that deferment increases the contribution of cool season species and other species of interest to the livestock system in the short and medium term. This second article concludes that the short-term deferment was not enough to increase the contribution of cool season species, but increased the number of inflorescences of these species, which may contribute to strengthening presence of these species in the soil seed bank, and we may infer that these species increase their contribution in the long term. In the short (six months) and medium (two years) term redirection of the floristic composition, the effect of deferment can be seen by the increase of tussock species and resources conservation species on the environment degraded by overgrazing. Grouping species by common attributes shown to be suitable to describe community changes in a complex pasture.

For several decades, Moose population (Alces alces) in Sweden has been growing considerably. This increase has resulted in a greater impact and damage in trees, especially in Scots pine (Pinus sylvestris) which is the basis of Swedish forestry and, therefore the economical losses are considerable. For this reason, for several years, to know Moose feeding behaviour and to look for measures to reduce it is impact on forestry have marked the guidelines of research. The Moose damages the pine mainly in winter, when there are no herbaceous plants and deciduous trees to eat. This project has focused on reducing the moose damage from a biodiversity point of view, following a line similar to the "Optimal forage theory". According to this theory, a moose selects the food according to the composition and the richness of the species. Results have conducted to relation plant-richness with Moose Browsing Damage on Scots pine and with more abundance of trees around a Scots pine, Moose Browsing Damage decreases considerably. In this project I also study the association between Moose activity and the plant-richness. Both (Moose Browsing damage on Scots pine and Moose activity) were tested with Shannon´s and Simpson´s index, two biodiversity indexes. The amount of different plants species nearby a Scots pine were measured and studied, as well. Also, I did the Pearson´s coefficient as an initial relationship between the variables (and observed if it is negative or positive) and ANOVA analysis. Although the conclusions weren´t as clear as I expected, in my opinion this study offered another tool to reduce the before mentioned impact.

As pastagens nativas possuem grande importância na produção de produto animal e nos serviços ecossistêmicos. Muitos estudos realizados sobre esse recurso natural demonstram seu potencial produtivo. Este é diretamente ligado ao manejo utilizado, e a oferta de forragem adotada exerce forte influência tanto na produção primária quanto secundária. Em resposta disso também se sabe que a composição da vegetação responde de maneira positiva ou negativa ao regime de desfolha. Entretanto, uma abordagem mais holística relativa à essa resposta e ao uso do diferimento ainda era uma deficiência desta área do conhecimento. No sentido de supri-la, foi desenvolvido o presente trabalho resultando em dois artigos. Ambos se valem do uso de grupos de espécies para caracterização dos ambientes. O primeiro aborda o padrões de diversidade e o comportamento da vegetação após 28 anos de regimes de desfolhas contrastantes mediada pelas ofertas de forragem 4 kg de MS por 100 kg de PV e 8-12 (8 primavera e 12 no restante do ano) kg de MS por 100 kg de PV. Não houve diferença na riqueza e diversidade entre os ambientes, mas verificou-se resposta negativa na composição de espécies na área com sobrepastejo: espécies que são menos eficientes como forrageiras e na proteção do solo, as ruderais, sucederam as espécies conservadoras de recurso que proporcionam maior estabilidade na disponibilidade de forragem e são mais eficientes na proteção do solo. Já quando a intensidade de pastejo foi moderado houve predominância de espécies conservadoras. O segundo artigo testa duas épocas de diferimento nos dois ambientes citados anteriormente objetivando avaliar a vegetação perante esses tratamentos com a hipótese que o diferimento aumenta a contribuição de espécies hibernais e as demais espécies de interesse forrageiro no curto e médio prazo. Esse segundo artigo conclui que o diferimento no curto prazo não foi suficiente para aumentar a contribuição de espécies hibernais, porém aumentou o número de inflorescências dessas espécies que reforçam o banco de sementes do solo, podendo-se inferir que tais espécies aumentem sua contribuição no longo prazo. No curto (seis meses) e médio prazo (dois anos) o redirecionamento da composição florística por efeito de diferimento é dado pelo aumento de espécies cespitosas e conservadoras de recursos no ambiente degradado por longo tempo de sobrepastejo. O uso de grupos de espécies mostra-se como indicadores adequados para caracterizar o grau de herbivoria exercido em pastagens complexas. / Rangelands have great importance in the animal production and ecosystem services. Many studies on this natural resource demonstrate their productive potential, which is a direct consequence of management. The adopted herbage allowance exerts strong influence on primary and secondary production through changes in botanical composition. However, a more holistic approach on interactions and responses of vegetation to grazing and the use of deferment still a presents shortcoming of this area of knowledge. The present work was developed to contribute in filling these gaps, resulting in two papers. Both avail themselves on the use of species groups for characterization of environments. The first addresses the diversity patterns and behavior of vegetation after 28 years of contrasted defoliation regimes mediated by herbage allowance of 4 kg DM per 100 kg BW and 8-12 (8 spring and 12 in the rest of the year) kg DM per 100 kg BW. There was no difference in the richness and diversity between environments but there was a negative response in species composition in the overgrazed area: species that are less efficient as forage and soil protection, the ruderals, succeeded conservative resource species that provide greater stability in the availability of forage and are more efficient in protecting the soil. When grazing intensity was moderate conservative species predominated. The second article seeks to verify effects of two periods (spring or fall) of deferment in both environments that already were studied in the first chapter, with the hypothesis that deferment increases the contribution of cool season species and other species of interest to the livestock system in the short and medium term. This second article concludes that the short-term deferment was not enough to increase the contribution of cool season species, but increased the number of inflorescences of these species, which may contribute to strengthening presence of these species in the soil seed bank, and we may infer that these species increase their contribution in the long term. In the short (six months) and medium (two years) term redirection of the floristic composition, the effect of deferment can be seen by the increase of tussock species and resources conservation species on the environment degraded by overgrazing. Grouping species by common attributes shown to be suitable to describe community changes in a complex pasture.

As pastagens nativas possuem grande importância na produção de produto animal e nos serviços ecossistêmicos. Muitos estudos realizados sobre esse recurso natural demonstram seu potencial produtivo. Este é diretamente ligado ao manejo utilizado, e a oferta de forragem adotada exerce forte influência tanto na produção primária quanto secundária. Em resposta disso também se sabe que a composição da vegetação responde de maneira positiva ou negativa ao regime de desfolha. Entretanto, uma abordagem mais holística relativa à essa resposta e ao uso do diferimento ainda era uma deficiência desta área do conhecimento. No sentido de supri-la, foi desenvolvido o presente trabalho resultando em dois artigos. Ambos se valem do uso de grupos de espécies para caracterização dos ambientes. O primeiro aborda o padrões de diversidade e o comportamento da vegetação após 28 anos de regimes de desfolhas contrastantes mediada pelas ofertas de forragem 4 kg de MS por 100 kg de PV e 8-12 (8 primavera e 12 no restante do ano) kg de MS por 100 kg de PV. Não houve diferença na riqueza e diversidade entre os ambientes, mas verificou-se resposta negativa na composição de espécies na área com sobrepastejo: espécies que são menos eficientes como forrageiras e na proteção do solo, as ruderais, sucederam as espécies conservadoras de recurso que proporcionam maior estabilidade na disponibilidade de forragem e são mais eficientes na proteção do solo. Já quando a intensidade de pastejo foi moderado houve predominância de espécies conservadoras. O segundo artigo testa duas épocas de diferimento nos dois ambientes citados anteriormente objetivando avaliar a vegetação perante esses tratamentos com a hipótese que o diferimento aumenta a contribuição de espécies hibernais e as demais espécies de interesse forrageiro no curto e médio prazo. Esse segundo artigo conclui que o diferimento no curto prazo não foi suficiente para aumentar a contribuição de espécies hibernais, porém aumentou o número de inflorescências dessas espécies que reforçam o banco de sementes do solo, podendo-se inferir que tais espécies aumentem sua contribuição no longo prazo. No curto (seis meses) e médio prazo (dois anos) o redirecionamento da composição florística por efeito de diferimento é dado pelo aumento de espécies cespitosas e conservadoras de recursos no ambiente degradado por longo tempo de sobrepastejo. O uso de grupos de espécies mostra-se como indicadores adequados para caracterizar o grau de herbivoria exercido em pastagens complexas. / Rangelands have great importance in the animal production and ecosystem services. Many studies on this natural resource demonstrate their productive potential, which is a direct consequence of management. The adopted herbage allowance exerts strong influence on primary and secondary production through changes in botanical composition. However, a more holistic approach on interactions and responses of vegetation to grazing and the use of deferment still a presents shortcoming of this area of knowledge. The present work was developed to contribute in filling these gaps, resulting in two papers. Both avail themselves on the use of species groups for characterization of environments. The first addresses the diversity patterns and behavior of vegetation after 28 years of contrasted defoliation regimes mediated by herbage allowance of 4 kg DM per 100 kg BW and 8-12 (8 spring and 12 in the rest of the year) kg DM per 100 kg BW. There was no difference in the richness and diversity between environments but there was a negative response in species composition in the overgrazed area: species that are less efficient as forage and soil protection, the ruderals, succeeded conservative resource species that provide greater stability in the availability of forage and are more efficient in protecting the soil. When grazing intensity was moderate conservative species predominated. The second article seeks to verify effects of two periods (spring or fall) of deferment in both environments that already were studied in the first chapter, with the hypothesis that deferment increases the contribution of cool season species and other species of interest to the livestock system in the short and medium term. This second article concludes that the short-term deferment was not enough to increase the contribution of cool season species, but increased the number of inflorescences of these species, which may contribute to strengthening presence of these species in the soil seed bank, and we may infer that these species increase their contribution in the long term. In the short (six months) and medium (two years) term redirection of the floristic composition, the effect of deferment can be seen by the increase of tussock species and resources conservation species on the environment degraded by overgrazing. Grouping species by common attributes shown to be suitable to describe community changes in a complex pasture.

L’ajout de diversité végétale est présenté comme un moyen d’améliorer la durabilité des agrosystèmes. Cependant, il y a encore des manques importants de connaissances sur l’effet de la diversité végétale sur les processus à la base de l’élaboration des rendements. Les systèmes agroforestiers tropicaux couvrent une large gamme de diversité végétale ; ce sont donc de bons modèles pour étudier la relation entre diversité et productivité. Les systèmes agroforestiers de la région de Talamanca au Costa Rica sont particulièrement intéressants car au sein de communautés végétales complexes, ils comprennent des bananiers et des cacaoyers qui sont des cultures de rente importantes et dont la production peut facilement être quantifiée et analysée. Une autre spécificité de ces systèmes est qu’ils présentent des organisations spatiales particulièrement diverses. Analyser comment la diversité des plantes et son organisation influe sur les performances de ces systèmes est particulièrement complexe et nécessite le développement de nouvelles approches. Les objectifs de cette thèse étaient d’étudier : i) quels facteurs affectent la relation entre diversité végétale et productivité ? ii) Comment la diversité végétale influence la productivité globale des systèmes agroforestiers ? et iii) Comment la structure spatiales des communautés de plantes cultivées influence leurs rendements ?Tout d’abord, une méta-analyse a été menée afin d’étudier la relation entre diversité végétale et production dans une très large gamme de systèmes naturels et cultivés. Cette analyse a notamment traité du rôle de la latitude, du climat et de la structure de la canopée sur cette relation. Elle a montré que le gain lié à la diversité végétal tend à diminuer avec la magnitude de cette diversité. Nos résultats montrent également que la réponse de la productivité à la richesse spécifique en plantes dépend énormément du type de communauté considéré, notamment si la communauté comprend des arbres.Ensuite, un réseau de 180 placettes situées dans 20 parcelles d’agriculteurs a été étudié dans la région de Talamanca. Pendant un an, la production de chaque plante a été évaluée. Cette production a ensuite été convertie en revenus en accord avec les prix du marché local. Alors qu’un effet positif de la diversité végétale cultivée a été observé sur le revenu globale (de chaque placette), cet effet était très contrasté si on le considérait séparément pour les différents groupes fonctionnels. Cet effet était positif pour les plantes des groupes appartenant aux strates hautes et négatif pour les plantes des groupes appartenant aux strates basses. Ces résultats suggèrent que la complémentarité entre plantes était plus forte pour les plantes des strates hautes et qu’inversement la compétition était plus forte dans les strates plus basses.La seconde phase de l’analyse des données de ce réseau de parcelles a visé à étudier l’effet du voisinage de chaque bananier ou cacaoyer sur leur production. Une approche d’analyse individu-centrée a été développée afin de déterminer si le nombre de voisins d’un groupe donné dans un rayon donné était un bon prédicteur de la croissance ou du rendement de chaque bananier ou cacaoyer. Les résultats montrent que la distance à laquelle la production d’un bananier ou d’un cacaoyer est affectée par ses voisins dépend de la taille de ces. De manière surprenante, les grands arbres ont eu un effet plus faible que les arbres plus petits. Cela suggère que des densités modérées de grands arbres pourraient être compatibles avec une production de bananiers et de cacaoyer avec un haut niveau de rendement. Ces résultats ont été discutés en termes de complémentarité et compétition pour la lumière. Des pistes d’organisation sont proposées et discutées au regard de la maximisation des rendements des cultures de rente et des autres cultures mais aussi dans la provision de services écosystémiques au sens large. / Adding plant diversity is increasingly presented as a mean to improve the sustainability of agrosystems. However, there is still a lack of knowledge on how plant functional diversity alters processes that support production. Because they cover a broad range of plant diversity, agroforestry systems in the tropics are a good case study to better understand the diversity-production relation. Agroforestry systems in the Talamanca region in Costa Rica are particularly interesting because among the cultivated plants they encompass, banana and cacao are two cash crops of major importance and for which production can easily be quantified and analyzed. Another specificity of these systems is that their vertical and horizontal organization is particularly diverse. Understanding how plant diversity and its organization alter the performances of these complex systems is particularly challenging and requires developing new approaches. The objectives of this thesis were to address the following questions: i) Which factors affect the relationship between plant diversity and productivity? ii) How plant diversity influences the global productivity of agroforestry systems? and iii) How the spatial structure of the plant community affects yields?First, a meta-analysis was carried out to address the diversity-production issue among a very broad range of systems world-wide. This analysis focused on how latitude, climate, and canopy structure modify the effect of plant richness on productivity of agricultural and natural ecosystems. It showed that the gain per unit of diversity added decreased as plant richness increased. Our findings also showed that the response of productivity to plant richness largely depends on the type of plants in the community, especially if the community includes trees.Then, we extensively studied the diversity and the productivity of 180 plots located within 20 fields in the Talamanca region. A global evaluation of the productivity of these systems was possible with the estimation of the production of each plant during 1 year. This production was converted into income according to local market prices. While we observed a global positive effect of plant diversity on global income, this effect was contrasted according to the functional group considered (banana, cacao, other fruits, timber, firewood. When considering the functional group separately, there was a positive effect of plant diversity for higher strata groups and a negative effect for lower strata groups. This suggested that complementarity between plants was stronger than competition for those plants occupying the higher strata of the canopy but that competition was stronger than complementarity for plants occupying the lower strata of the canopy.The second part of the analysis of the Talamanca fields dataset focused on the effect of neighbouring plants on the production of banana and cacao plants. An individual-based analysis was developed to determine whether the number of neighbouring plants of a given functional groups explained the potential yield of each banana or cacao plant. We found that the distance at which other plants alters the yield of banana or cacao plants was greater for larger functional groups (fruit or wood trees) than for smaller ones (cacao trees or banana plants). Interestingly, higher strata trees had a smaller effect than lower strata trees, suggesting that moderate densities of tall trees could be compatible with high banana and cacao production. These findings were discussed in terms of complementary and competition with respect to the availability of light at higher and lower strata of the canopy. On an applied perspective, our results suggest that productivity could be maximized by a reasonably number of plant species, and then we proposed new direction to organize fields in order to maximize the production of cash crops while providing supplementary income for farmers and ecosystem services.