No description available.

Pollen and packrat midden data from the Great Basin indicate that the Pleistocene vegetation of the region was a combination of subalpine conifers on coarse substrates and steppe plants on finer valley bottom soils. While some of the modern dominants of the woodland zone were apparently absent from this region, other woodland and montane plants apparently persisted through the late Pleistocene. Some features of the Pleistocene environment, such as the large pluvial lakes, apparently disappeared by 12,000 yr B.P., while subalpine plants remained well below their modern elevational limits after 11,000 yr B.P. Limber pine and Rocky Mountain juniper apparently did not retreat from the lower mountain slopes until after 6500 yr B.P. Montane and woodland conifers, rare or absent in the region during the Wisconsin, dispersed across the region in the Middle Holocene. Other plants apparently did not reach their modern geographic limits until after 3000 yr B.P. The low elevational occurrences of subalpine species suggest that the late Pleistocene climate of the Great Basin was characterized by summer temperatures that were as much as 10°C cooler than those of today, and that there was some augmentation in the level of mean annual precipitation. The vegetational records provide no evidence of greater than modern levels of summer precipitation. While the persistence of montane plants at relatively low elevations implies cool or moist conditions through the Early Holocene, evidence from lacustrine systems suggests that there was a trend toward increasingly dry conditions during this period. The main period of migrations of woodland plants seems to slightly postdate the warmest and/or driest part of the Holocene, and these migrations may have been related to relatively high levels in summer temperatures, summer precipitation, and/or winter temperatures. Pollen data from a high elevation site, in conjunction with changes in water level in lower elevational lakes, suggest a return to cooler and/or moister conditions after 4000 yr B.P.

No description available.

Thesis (PhD)--Stellenbosch University, 2013. / ENGLISH ABSTRACT: Historical vicariant processes due to glaciations, resulting from the large-scale environmental changes during the Pleistocene (0.012-2.6 million years ago, Mya), have had significant impacts on the geographic distribution of species, especially also in marine systems. The motivation for this study was to provide novel information that would enhance ongoing efforts to understand the patterns of biodiversity on the South African coast and to infer the abiotic processes that played a role in shaping the evolution of taxa confined to this region. The principal objective of this study was to explore the effect of Pleistocene climate changes on South Africa′s marine biodiversity using five intertidal gastropods (comprising four rocky shore species Turbo sarmaticus, Oxystele sinensis, Oxystele tigrina, Oxystele variegata, and one sandy shore species Bullia rhodostoma) as indicator species. Sequence data obtained from partial segments of the mitochondrial cytochrome oxidase c subunit 1 (COI), and the nuclear ribosomal DNA (encompassing part of 5.8S, second Internal Transcribed Spacer and part of 28S, hereinafter called ITS2; or comprising part of the first Internal Transcribed Spacer, 5.8S, second Internal Transcribed Spacer and part of 28S, hereinafter called ITS), were used as genetic markers to construct phylogeographic patterns and to investigate demographic histories of the taxa. Population structure was investigated using haplotype network analyses, pairwise ΦST statistics, analysis of molecular variance (AMOVA), isolation by distance analyses, Bayesian analysis of population structure (BAPS) and coalescent analysis of gene flow. Demographic history was analysed through Fu′s Fs tests, mismatch distributions, and Bayesian skyline plots. Demographic analyses suggest that all five intertidal gastropods studied experienced demographic expansions dating to the late Pleistocene. The sandy shore direct developer B. rhodostoma began expansion after the LGM (c. 15 kya) whereas for the four rocky shore broadcast spawners (T. sarmaticus, O. sinensis, O. tigrina, and O. variegata) the onset of expansion coincided with or preceded the LGM (c. 25, 60, 50, 40 kya, respectively). Consistent with recent range expansions and gene flow patterns, the population genetic structure in all species was characterised by shallow or a lack of population differentiation. Oxystele variegata was an exception as it showed a deep disjunction, of late Pleistocene origin, between individuals in the west coast Namaqua Bioregion and those in the south coast Agulhas Bioregion. These results provide strong evidence of the vital role that Pleistocene climatic changes and current regimes played in shaping the nature and distribution of biodiversity on the South African coast. In addition, gene flow in all species, except O. tigrina, was remarkably asymmetrical with the regions around Cape Infanta and Port Elizabeth acting as source populations. Considering the generally weak population genetic structure and gene flow patterns detected for most gastropod species studied here, it is recommended that T. sarmaticus, O. sinensis, O. tigrina and B. rhodostoma be managed as panmictic populations, and that the region encompassing Cape Infanta, and Port Elizabeth should be prioritised for conservation as it appears to harbour source populations. Oxystele variegata was the only species showing distinct population structure and in this instance, species specific conservation efforts should recognize this divergence by treating the two genetic assemblages as distinct management units. / AFRIKAANSE OPSOMMING: Historiese vikariante prosesse kan toegeskryf word aan glasiasie, en het tot gevolg gehad dat grootskaalse veranderinge in die omgewing plaasgevind het tydens die Pleistoseen (,012 - 2.6 miljoen jaar gelede, Mjg). Dit het 'n beduidende impak gehad op die geografiese verspreiding van spesies, veral ook in die mariene stelsels. Die motivering vir hierdie studie was om nuwe data te voorsien wat sal bydrae tot die voortgesette pogings om die patrone van biodiversiteit langs die Suid-Afrikaanse kus te verstaan. Dit sou ook help om die abiotiese prosesse af te lei wat 'n rol gespeel het in die evolusie van taksa wat in hierdie streek voorkom. Die hoofdoel van hierdie studie was om die effek van die Pleistoseen klimaatsveranderinge op Suid-Afrika se mariene biodiversiteit te bepaal deur gebruik te maak van vyf intergety slak spesies as indikatore (vier wat in rotsagtige gebiede voorkom: Turbo sarmaticus, Oxystele sinensis, Oxystele tigrina, Oxystele variegata en 'n sanderige strand spesies: Bullia rhodostoma). Volgorde data verkry vanaf gedeeltelike segmente van die mitochondriale sitochroom oksidase c subeenheid 1 (COI), en die kern ribosomale DNA (bestaande uit 'n deel van 5.8S, tweede interne getranskribeerde spasieërders en 'n deel van 28S), hierna genoem ITS2 is gebruik as genetiese merkers om filogeografiese patrone te dokumenteer en ook om die demografiese geskiedenis van die spesies te ondersoek. Bevolking struktuur is ondersoek deur gebruik te maak van haplotipe netwerk analise, paarsgewyse ΦST statistiek, analise van molekulêre variansie (AMOVA), isolasie deur afstand analise, Bayesiaanse analise van die bevolking struktuur (BAPS) en analise van gene vloei. Demografiese geskiedenis is ontleed deur Fu se Fs toetse, misparing verdelings, en Bayesiaanse luglyn kurwes. Demografiese ontleding dui daarop dat al vyf die intergety slakke wat ondersoek is demografiese uitbreidings ervaar het wat terugdateer tot die einde van die Pleistoseen. Die sanderige strand direkte ontwikkelaar, B. rhodostoma, het die uitbreiding begin na die LGM (c. 15 Kya), terwyl vir die vier rotsagtige kusbewoners wat eiers oor 'n uitgebreide gebiede versprei (T. sarmaticus, O. sinensis, O. tigrina, en O. variegata) het die aanvang van die bevolkings uitbreiding saamgeval met die laaste galsiasie of dit voorafgegaan (c. 25, 60, 50, 40 Kya, onderskeidelik). In ooreenstemming met die onlangse reeks bevolkings uitbreidings, is die bevolking genetiese struktuur in alle spesies gekenmerk deur weinig differensiasie. Oxystele variegata was 'n uitsondering en het 'n ontwrigting van laat Pleistoceen oorsprong getoon tussen individue langs die weskus Namaqua Biostreek en dié in die suid kus Agulhas biostreek. Hierdie resultate voorsien sterk bewyse van die belangrike rol wat die Pleistoseen klimaatsveranderinge gespeel het in die vorming en verspreiding van biodiversiteit langs die Suid-Afrikaanse kus. Daarbenewens, geen vloei in alle spesies, behalwe O. tigrina, was merkwaardig asimmetries. Kaap Infanta en Port Elizabeth verteenwoordig moontlik die bron bevolkings. Met inagneming van die geringe bevolking genetiese struktuur en geenvloei patrone wat waargeneem is vir die meeste slak spesies wat bestudeer is, word dit aanbeveel dat T. sarmaticus, O. sinensis, O. tigrina en B. rhodostoma bestuur word as 'n panmiktiese bevolking, en dat die streek wat Kaap Infanta en Port Elizabeth insluit geprioritiseer moet word vir bewaring. Oxystele variegata was die enigste spesie wat duidelike bevolking struktuur getoon het en in hierdie geval, moet spesie spesifieke bewaringspogings aangewend word.

Shallowly-buried channel systems have been imaged on the New Jersey Shelf with high-resolution seismic imaging. These channels formed as riverine systems that occupied the exposed shelf during the Last Glacial Maximum, ~18 ka. Subsequent sea level rise ~15-12 ka modified the valleys, forming estuaries and filling the channels with estuarine sediments. The infill sediments within the channel provide evidence for the response of the shelf to the Late Pleistocene glacio-eustatic sea level rise, but little work has been done on samples from these strata. This study aids in the ground-truthing of previous stratigraphic results by analyzing the cores collected within the infill sediments. The seismic stratigraphy of fill sediments from the mid-shelf and outer-shelf channels are structurally dissimilar. The mid-shelf channel fill stratigraphy is dominated by finely-laminated U-shaped reflectors throughout the section, with cut and fill geometries. In contrast, the outer shelf channel fill stratigraphy is a well-ordered sequence of 4 identifiable, primarily flat-lying seismic units. We collected five cores in mid-shelf channels (~30 m water depth), one in an outer shelf channel (~80 m of water depth) and one core in the trangressive ravinement surface. Cores were logged for density and seismic velocity. Grain size analysis was conducted by settling column and laser particle size analyzer. Radiocarbon analysis of the stratigraphy was conducted with the shell fragments and organic mud within the samples. The foraminiferal assemblages aided in determining the depositional environment. Using these data I investigated the differences in depositional environment of the mid- and outer-shelf channels systems, and consider these results in the context of sedimentary models for estuarine processes. The radiocarbon dates and foraminiferal are consistent with channel infill in an estuarine environment. Grain size and density log data indicate that the mid-shelf channel fills are sandier than the outer-shelf channel fills, which leads me to infer that the sediment from the mid-shelf channels was deposited in a higher energy environment than that of the sediment in the outer shelf channels. The stratigraphic differences and locations of the channel systems are similar to the Zaitlin (1994) model of incised valley infill, but infill of the mid-shelf channel system could possibly be the result of a catastrophic meltwater flood event occurring ~14 ka as glacial lakes to the north broke their dams and flooded the mid-shelf. / text

Topography growth and sediment fluxes in active subduction margin settings are poorly understood. Geological record is often scarce or hardly accessible as a result of intensive deformation. The Hawke Bay forearc basin of the Hikurangi margin in New Zealand is well suited for studying morphstructural evolution. It is well preserved, partly emerged and affected by active tectonic deformation during Pleistocene stage for which we have well dated series and well-known climate and eustasy. The multidisciplinary approach, integrating offshore and onshore seismic interpretations, well and core data, geological mapping and sedimentological sections, results in the establishment of a detailed stratigraphic scheme for the last 1.1 Ma forearc basin fill. The stratigraphy shows a complex stack of 11 eustasy-driven depositional sequences of 20, 40 and 100 ka periodicity. These sequences are preserved in sub-basins that are bounded by active thrust structures. Each sequence is characterized by important changes of the paleoenvironment that evolves between the two extremes of the glacial maximum and the interglacial optimum. Thus, the Hawke Bay forearc domain shows segmentation in sub-basins separated by tectonic ridges during sea level lows that become submerged during sea level highs. Over 100 ka timescale, deformation along active structures together with isostasy are responsible of a progressive migration of sequence depocenters towards the arc within the sub-basins. Calculation of sediment volumes preserved for each of the 11 sequences allows the estimation of the sediment fluxes that transit throughout the forearc domain during the last 1.1 Ma. Fluxes vary from c. 3 to c. 6 Mt.a⁻¹. These long-term variations with 100 ka to 1 Ma timescale ranges are attributed to changes in the forearc domain tectonic configuration (strain rates and active structure distribution). They reflect the ability of sub-basin to retain sediments. Short-term variations of fluxes (<100 ka) observed within the last 150 ka are correlated to drastic Pleistocene climate changes that modified erosion rates in the drainage area. This implies a high sensitiveness and reactivity of the upstream area to environmental changes in terms of erosion and sediment transport. Such behaviour of the drainage basin is also illustrated by the important increase of sediment fluxes since the European settlement during the 18th century and the following deforestation.

Microbial methane from subsurface organic-rich units such as coals and shale support approximately 5% of the United States and Canada's energy needs. In the deep subsurface, microbial methane is formed by the metabolism of primarily CO2, H2, and acetate by methanogens. These metabolites are the by-products of multi-step biodegradation of complex organic matter by microbial consortia. This study investigates microbial methane in the Illinois Basin, which is present in organic-rich shallow glacial sediments (surficial), Pennsylvanian coals (up to 600 m depth), and the Upper Devonian New Albany Shale (up to 900 m depth). Findings from the study show that hydrogeochemical conditions are favorable for methanogenesis in each reservoir, with a decrease in groundwater flushing rates corresponding to a decrease in average reservoir depth and an increase in carbon isotopic fractionation. The deeper reservoirs (coals and shale) were paleopasteurized, necessitating re-inoculation by methanogens. The microbes were likely advectively transported from shallow sediments into the coals and shale, where areas of microbial methanogenesis correlate with freshwater recharge. The recharge in the shale was primarily sourced from paleoprecipitation with minor contributions from glacial meltwater during the Pleistocene (4He ages). All areas sampled in the shale were affected by Pleistocene recharge, however groundwater ages in areas of microbial methanogenesis are younger (average 0.33 Ma) than areas with thermogenic methane (average 1.0 Ma). Estimates of in-situ microbial methane production rates for the shale (10-1000 TCF/Ma) are 104-106 times slower than laboratory rates. Only limited biodegradation is observed in the shale. In-situ stimulation of methane production may be most effective if aimed at increasing production of the supporting microbial consortia as well as methanogens. Trace metal concentrations in the shale are below known levels of inhibition or enhancement, with the exception of Fe, suggesting that microbial methanogenesis is not repressed by any of the measured trace metals and may be improved with the addition of Ag, Co, Cr, Ni, and Zn.

A stratigraphic radiocarbon study at the Wilicox Playa and vicinity in Cochise County, southeastern Arizona has revealed a sedimentary sequence reflecting the lake level chronology of ancient Lake Cochise. The lower green clay was deposited in a lake at least 30 miles long from before 30,000 years ago until about 13,000 years ago. A marl formation which began depositing about 25,000 years ago and continued until the lake diminishe to about its present state 13,000 years ago, indicates a warm, moist climate at that time. The period from 13,000 B.P. (before present) and 11,000 B.P. was one of alluvial deposition north of the playa and channel cutting east of the playa. An upper green clay, 6 to 8 inches thick, represented the final phase of Lake Cochise, lasting from 11,500 B.P. until 10,500 B.P. A thin marl layer was deposited during this phase, indicating warm moist conditions again. The lake rapidly receded, some playa sediments deflated from the surface, and dunes formed north of the playa. Preliminary studies of two playas near Lordsburg, New Mexico and one at the San Augustin Plains, New Mexico, indicate these fluctuations were responding to a general climate change rather than isolated tectonic disturbances. The climate chronology concluded from this study is consistent with known climatic variations in the world.

This study presents an analysis of the zooarchaeological remains from Klissoura Cave 1, a Middle Paleolithic through Mesolithic site in Peloponnese, Greece. Changes in subsistence patterns are evaluated across a long sequence (ca. 80,000-10,000 BP) against a backdrop of environmental change. Results are interpreted using models from evolutionary ecology, specifically prey choice, central place foraging, and patch choice models. Two major trends are apparent in the series. One is a decline in the exploitation of high-ranked ungulate species with an overall increase in lower-ranked small game animals. The second is an increase in low-ranked small, fast-fast moving animals (e.g., hares and partridges) at the expense of higher-ranked small, slow-moving animals (e.g., tortoises). These changes cannot be accounted for by environmental shifts alone, though shifts in ungulate diversity likely track the expansion and contraction of plant communities. The increase in use of low-ranked prey indicates human population growth and demographic pressure in southern Greece during the late Pleistocene and early Holocene. In addition to these overarching trends, there are changes in site use during the sequence. In the Middle Paleolithic, foragers used Klissoura Cave 1 more during the winter and overwhelmingly hunted prime-aged adult animals, maternal herds of fallow deer in particular. In the Upper Paleolithic and later periods, the site continued to be used during the winter, in addition to other times of year, but the mortality profiles reflect a natural fallow deer herd structure. There was an intense period of occupation during the Aurignacian period. This is evidenced by numerous clay-lined hearth features, a possible rock-lined structure, and increases in ornaments, as well as abundant lithic and faunal materials. The ungulate faunas are particularly rich during this period, but there is evidence of resource intensification based on increased bone marrow processing and the transport of marrow-rich elements to the site. After this period there was a gradual decline in site use through the end of the Upper Paleolithic and into the Mesolithic, though the exploitation of low-ranked resources (e.g., small, fast-moving game) indicates that populations were on the rise in there region as a whole.

African climate changed considerably throughout the Pleistocene (2.588 million (Ma) to 12 thousand years ago (ka)). The timing, rate, and magnitude of past climate change across the continent impacted the evolutionary and migratory history of many mammalian species, including hominins. Investigating paleoclimatic variability through time at local and regional scales allows for an assessment of the extent to which climate change affected hominin evolution in Africa. This dissertation presents three approaches for increasing the understanding of past climate change in Africa. One method is to critically synthesize the existing literature of African climate (n=85) and hominid demography (n=64) over a restricted time frame (150 ka to 30 ka) and specific spatial scale (regional). Results from this study are two-fold: 1) climate change in Africa during this period was variable by region, responding to different climate-forcing mechanisms, and 2) changes in population and climate were asynchronous and likely created alternating opportunities for migration into adjacent regions, including hominin migrations out of Africa (~140-80 ka). The second approach is to evaluate modern ecological relationships between species and their environment to better quantify interpretations of paleoecological records. A modern distribution study of 33 ostracode species from 104 sites in the southwest arm of Lake Malawi suggest that depth-dependent variables likely define species niches. Relationships between ostracodes, fish and the green algae Botryococcus, were used to inform the paleoecological interpretations in the third study of this dissertation. Additional results suggest that macrocharcoal is likely delivered to the lake basin via river rather than wind-borne methods. The third approach involves primary analysis of climatic indicators from the sedimentary record to chronicle paleoecological and paleoenvironmental change at the basin scale through time. Results from a 380.7 meter-long sediment core recovered from Lake Malawi indicate a change of state likely caused by local tectonism, which affected ostracode assemblages, but had little effect on lake level history through time. Furthermore, the local hydroclimate of Lake Malawi alternately covaried with global glacial/interglacial cycles and local insolation maxima over the past 1.25 Ma. The magnitude and frequency of hydroclimatic variability in the watershed will be further assessed in future research.