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The western spruce budworm (Choristoneura occidentalis Freeman) is the most widespread and destructive defoliator of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) forests in British Columbia. Over the past two decades, western spruce budworm outbreaks have been sustained and widespread in the interior of British Columbia, leaving the forest industry and many forest-dependent communities increasingly vulnerable to the economic consequences of these outbreaks. While a great deal is known about the impact of western spruce budworm outbreaks on tree growth and form, substantial knowledge gaps remain as to the historic variability of western spruce budworm outbreaks and the consequences of defoliation on fundamental characteristics such as wood structure. This research focused on describing historic and contemporary western spruce budworm outbreaks across multiple spatial and temporal scales in south-central British Columbia using dendrochronology and wood anatomy techniques. Outbreak histories over the past 435 years were reconstructed using a network of tree-ring chronologies from central British Columbia, revealing that 12 western spruce budworm outbreaks have occurred since the early 1600s, with a mean return interval of 30 years. Further, the research illustrates that outbreaks observed over the last 40 years are not unprecedented, which does not support the perception that western spruce budworm is moving northward into central British Columbia. To evaluate the effects of a single western spruce budworm outbreak on the anatomical characteristics of Douglas-fir stemwood, tree ring data was collected from permanent sample plots that sustained both periodic and chronic western spruce budworm feeding. In mature even-aged stands of Douglas-fir, a documented outbreak occurred from 1976 to 1980 in the coastal transition zone of southern British Columbia. Based on microscopic wood anatomical measurements it was shown that the tree rings formed during this outbreak had significantly lower percentages of latewood, reduced mean cell wall thickness and smaller radial cell diameters relative to wood formed during periods without budworm feeding. Western spruce budworm defoliation temporarily modified cellular characteristics, which has implications for wood quality. In uneven-aged stands of mature Douglas-fir, located in the xeric southern interior of British Columbia, there has been a sustained western spruce budworm outbreak since 1997. Tree rings formed during this outbreak had progressively larger earlywood lumen area and radial cell diameter, reduced latewood cell wall thickness, latewood radial cell diameters, and lower percent latewood. Mixed-effects models revealed that climatic variables, defoliation severity, defoliation duration, and in limited cases canopy class were the best predictors of xylem features. The severity and duration of western spruce budworm defoliation, as well as site factors that influence moisture conditions effect the degree and direction of anatomical changes in the stemwood of Douglas-fir. This research fills a number of knowledge gaps by providing insights into the temporal and spatial dynamics of western spruce budworm outbreaks in central British Columbia over multiple centuries, and the plasticity of anatomical features in the stemwood of Douglas-fir during discrete western spruce budworm outbreaks. These research findings suggest that Douglas-fir forests are resilient to western spruce budworm outbreaks over space and time. / Graduate

Plant macrofossils from packrat middens in two southeastern Utah caves outline development of modern plant zonation from the late Wisconsin. Allen Canyon Cave (2195 m) and Fishmouth Cave (1585 m) are located along a continuous gradient of outcropping Navajo Sandstone that extends from the Abajo Mountains south to the San Juan River. By holding the site constant, changes in the floral composition for a plot of less than one hectare can be observed, even if sporadically, over tens of millennia. At Allen Canyon Cave, engelmann spruce-alpine fir forest was replaced by the present vegetation consisting of pinyon-juniper woodland on exposed ridgetops and cliffside stands of Douglas fir, ponderosa pine, and aspen. Xerophytic woodland plants such as pinyon, Plains prickly pear, and narrowleaf yucca arrived sometime in the middle Holocene between 7200 and 3400 B.P. At Fishmouth Cave, Utah juniper in Holocene middens replaced blue spruce, limber pine, Douglas fir, and dwarf and Rocky Mountain junipers in late Wisconsin samples. Disharmonious associations for the late Wisconsin occur only at the lower site with the xerophytes Mormon tea, Plains prickly pear, and narrowleaf yucca growing alongside subalpine conifers. One possible explanation involves the late Wisconsin absence of ponderosa and pinyon pines from the Colorado Plateaus. Released from competition at their lower limits, subalpine conifers were able to expand into lower elevations and mix with xerophytic plants found today in understories of pinyon-juniper and ponderosa pine woodlands. Quantitative climatic estimates are derived for the late Wisconsin by applying vertical lapse rates for temperature and precipitation to the amount of vegetation depression. The Fishmouth Cave sequence indicates a minimum lowering of 850 m for blue spruce, limber pine, and dwarf juniper. A depression of at least 700 m for engelmann spruce and alpine fir is suggested for the Allen Canyon locality. Use of conservatively low lapse rates for stations below 2080 m yields a 3-4°C cooling from present mean annual temperature and 35 to 60 percent more rainfall than today. Steeper lapse rates associated with more mountainous terrain suggest a 5°C lowering in temperature and up to 120 percent increase over modern precipitation.