The objective of this study was to develop a 3-D model for The Pontchartrain Estuary that was capable of long-term mass conservative simulation of salinities. This was accomplished in a multi-stage approach involving: a physical model of salinity exchange through a pass; a 3-D FVCOM model of the physical experiment; the development and testing of an FVCOM model for an idealized Pontchartrain Basin; and for the entire estuary. The data from the physical model tests were used to validate the performance of the FVCOM model with density-driven flows. These results showed that hydrostatic FVCOM captured the primary internal wave movement. The idealized basin simulations were used to evaluate several issues related to salinity transport, namely the relative importance of baroclinic forcing, tidal forcing and hydrology. The idealized domain also permitted the testing of sigma-gradients, spatial distribution of friction coefficients, wind stress and various boundary treatments. The results showed that the density-driven exchange of saltwater at the open boundary required a baroclinic boundary condition for salinity as well as a lateral filter at the boundary on each sigma layer. A new radiative baroclinic open boundary condition was developed for FVCOM. When tides and hydrology were included, the FVCOM model was shown to reproduce the seasonal salinity that has been observed for long-term periods. It was also found that the simulation of tides and salinity in FVCOM is very sensitive to the spatial distribution of the friction coefficient; relatively low friction was required in the open water regions and high friction was needed in the passes and waterways to reproduce the tides and salinity distribution. A variable friction coefficient option was coded on FVCOM. The findings from the idealized model were utilized to setup two models for the actual estuary. Both models extend from Lake Maurepas, one to the Chandeleurs Islands and the other to Mobile Bay. The baroclinic open boundary and variable friction were implemented in these models. They were calibrated for tides and salinity. The 2008 Bonnet Carré Spillway Opening was applied to the first model. A tidal pumping effect in Lake Pontchartrain was observed and captured by the model.

This research investigated patterns in elevation change in newly built wetlands in the Atchafalaya delta, and newly restored wetlands in the Sacramento-San Joaquin River delta. RSETs were used to measure small changes in elevation, and soil cores were processed to examine mineral and organic contributions. Elevation change was highly variable, responding to influences from water level, river discharge, storms and vegetation. Mineral matter consistently added more to the marsh soil through volumetric and gravimetric contributions. Organic contributions were not significantly different across sites, suggesting the type of emergent vegetation at a site may not be the most important factor. Sites with the lowest elevations had the highest rates of positive elevation change. Higher elevation sites were more exposed and had negative rates of elevation change. The findings suggest ideal sites for marsh building are in areas that receive sediment input, are protected from high-energy events, and can support emergent vegetation.

The addition of renewable energy in a previously diesel-powered off-grid micro-grid results in what is known in the field as a Stand-Alone Hybrid Renewable Energy System (HRES). Such initiative is a near-term target of both federal and provincial governments in Canada. Not only does it reduce environmental hazards like leaks, spills and air pollution, but the combination of renewable energy and fossil fuel generators can increase stability and lower the cost of electricity. It is deemed a crucial step towards a clean energy future, but also a necessity in the reconciliation process with Indigenous Peoples of Canada - many of who inhabit off-grid communities. The addition of renewable energy can greatly increase the independence of a community by reducing reliability on external diesel suppliers and creating job opportunities. To be successful, HRES need to be carefully planned; the variable and uncertain behaviour of natural resources add a level of complexity to the preliminary design stage. Energy systems are therefore simulated and optimized to estimate the lifecycle cost by determining the nature and capacity of their components and their operational strategy. Chapter 2 goes over the preliminary design stage of two HRES in British Columbian communities. Many modelling tools are available, ranging from full-factorial and linear optimization techniques that can solve single-objective problems, to meta-heuristic algorithms. One of the distinctions between different HRES modelling tools is the foresight horizon being used. Linear programming tools commonly have a perfect foresight over the typical year analysed, for both demand and natural resources. This can lead to an overly optimistic prediction of the lifecycle cost of a system when the reality of implementations comes with uncertainties. On the other hand, tools that use myopic foresight, or no knowledge of future parameters, can lead to pessimistic lifecycle cost estimates since the demand and power output of certain renewable energy technologies, like solar panels, can be known within a few hours. The purpose of Chapter 3 of this thesis is to guide readers towards the right tool in the context of energy system modelling for the preliminary design of HRES. It was found that the degree of importance of choosing the appropriate foresight approach is a function of renewable energy penetration, autocorrelation, and storage capacity. A system with a high renewable energy share, a low short-term (few hours) autocorrelation, and an optimal storage size will result in the highest NPC difference between the two methods. When planning for long-term HRES design, the choice of the foresight horizon can either be representative of a lower/upper cost boundary (perfect and myopic foresight respectively) or of the real-time predictability of the power output of the chosen renewable energy power source. The use of energy system modelling tools is often reserved for highly qualified personnel and is therefore costly for prospective communities. To improve community readiness with minimal investment, a simple alternative to energy system modelling is proposed in Chapter 4 for the integration of tidal stream turbines in British Columbia. A series of three logical conditions was demonstrated to inform on the viability of a project in terms of cost reduction in comparison to the business as usual scenario. These conditions were found to also be useful for determining the minimum scale, or the economic break-in scale, for a tidal stream turbine given a remote community. In this context, communities are found to be best described by the local price of diesel fuel as an easily accessible metric to represent the current cost of electricity, their electrical load scale, and the local tidal current resource. Ten British Columbian communities were selected to validate the results by comparing the set of conditions to a complete energy system modelling approach and four were found to reach savings of 10 % or more as compared to the business as usual scenario. The long-term objective of this work is to provide remote communities with an integrated, affordable, and turnkey solution for the displacement of diesel in their energy systems. The next steps in achieving this include augmented optimization tools to quantify and capture non-monetary value so that the modelling and multi-criteria decision-making steps of the design process can be bridged together. / Graduate

\"Chovia desde o dia 16, aumentando a intensidade das chuvas a partir das 18 horas do dia 17. As primeiras barreiras começaram a ceder nas primeiras horas da manhã do dia 18 e, às 13 horas, mobilizou-se a totalidade da avalanche\" - (CRUZ, 1974). A descrição do cenário catastrófico no Município de Caraguatatuba, no mês de Março de 1967, que resultou em um dos maiores desastres naturais ocorridos no Brasil (*), fomenta a discussão sobre as probabilidades de ocorrência de eventos pluviométricos e/ou elevações do nível do mar nas Zonas Costeiras. Diante dessa realidade, surgiu essa Dissertação, que abordou aspectos hidrológicos, hidráulicos marítimos e, estatísticos, baseados em dados pretéritos de pluviômetros e marégrafos da região do Litoral Norte do Estado de São Paulo. Foram obtidos resultados práticos para futuros projetos de macro-drenagem, obras fluviais e marítimas, que associam a probabilidade de ocorrência conjugada de determinadas chuvas, com correspondentes elevações dos níveis de maré. Também foram realizados estudos da evolução desses eventos conjugados, ao longo de sessenta anos. - (*) De acordo com informações de Cruz (1974), 436 mortes foram registradas oficialmente, 400 casas destruídas, 3.000 desabrigados numa população total de 15.000 habitantes. Foram formados depósitos de 4 m a 5 m de altura no Rio Santo Antônio com blocos entre 30 t e 100 t. / \"It was raining since March 16th, increasing the intensity of the rainfall at 6:00 pm on March 17th. The first slopes started to sag in the first hours of March 18th, at 1:00 pm, occurred all the debris-flow\" - (translated from CRUZ, 1974). The catastrophic scenario of the city of Caraguatatuba, in March of 1967, previously described, resulted from one of the most serious natural disasters in Brazil. It fosters discussions about probabilities of rainfall events and /or rise in the sea level in coastal areas. Hence, this research is a consequence of this reality, which approached hydrological, hydraulic and statistical areas, based on past data of rainfall stations and tidal stations in the region of the North Coastal Zone of the State of São Paulo (Brazil). Practical results were used for future designs of macro- drainage, fluvial and maritime projects, that associate the probability of occurrence of certain types of rainfall coupled with their corresponding increase in tidal levels. There were also evaluated studies of the evolution of these events combined over sixty years.

O presente trabalho dedica-se a uma modelagem da interação entre corpos celestes, em regime Newtoniano, levando-se em consideração as influências que suas deformações e viscosidades internas exercem sobre seus movimentos orbitais e suas velocidades angulares. A abordagem adotada é uma variação do conhecido problema do corpo pseudo-rígido, a qual simplifica drasticamente a determinação dos equilíbrios relativos e torna a questão da dinâmica matematicamente acessível. Com este tratamento, podemos relacionar ou comparar os resultados com aqueles estabelecidos na literatura, dentre eles: formato de equilíbrio de um fluido isolado em rotação, deformação de maré causada pela interação gravitacional e o torque de maré induzido no mesmo. Pela simplicidade do modelo pode-se ainda fazer uma análise qualitativa da dinâmica do sistema e obter estimativas sobre a velocidade com que se aproxima dos equilíbrios. / The present work is devoted to model the interaction among celestial bodies, in a Newtonian regime, but considering the role played by the internal deformation and viscosity on the orbital motion and angular velocities of the components of the system. The work is mainly developed with an alternative approach to the pseudo-rigid body model, which simplifies the determination of the relative equilibria and allows precise conclusions about the dynamics. So, we are able to compare the results of this theory with those established in the literature, namely: the equilibrium shape of an isolated fluid in rotation, the tidal elongation induced by gravitational interaction and the tidal torque. Due to its simplicity, we can further perform a qualitative analysis of the dynamics of the system and estimate the velocity of attraction of the equilibrium states.

Looking over a tidal wetland, the tidal network characterised by its intricate system of bifurcating, blind-ended tidal courses clearly stands out from the overall landscape. This tidal landform exerts a fundamental control on the morphology and ecology within the tidal environment. With today’s recognition of the ecological, economical and societal values provided by tidal wetlands, which has been notably reflected in the development of restoration management strategies across Europe and USA, there is a need to fully understand the nature and development of tidal networks as well as their relationships with associated landforms and biotic components (e.g. vegetation), to eventually guarantee the success of current and future restoration practices. Accordingly, this research aims to bring further insights into the bio-physical controls on the geomorphology of tidal networks. To this end, a combination of remote sensing, modelling and field activities was employed. A geo-spatial analysis was performed at Queen Mary, University of London (UK), to address the variability of tidal network patterns. A series of network scale morphometric variables was extracted using airborne LiDAR data among selected tidal networks across the UK depicting different planview morphologies, and supplemented with the collection of corresponding marsh scale environmental variables from published sources. Multivariate statistics were then performed to characterise the variability of tidal network patterns and identify the inherent environmental controls. The analysis has revealed that every network type can be characterised based upon measures of network size and complexity, with each network pattern depicting proper morphometric aspects. Particularly, the stream Strahler order and the median depth of the network main channel have the highest discriminating weight on the patterns investigated. High correlation between the latter variable and network main channel width has revealed that linear, linear-dendritic and dendritic networks followed a transitional gradient in their aspect ratio approximated by a power law and thus are seen to depict similar erosional processes. To the contrary, meandering networks clearly depart from this relationship, and show particular segregation in their aspect ratios with respect to dendritic networks. Globally, differentiation on network morphometric properties has been linked to environmental conditions specific to the marsh physiographic setting within which a tidal network develops. Conceptually, tidal networks seem to adapt to marsh environmental conditions by adopting suitable morphologies to drain their tidal basin effectively. An eco-geomorphic modelling framework was developed at University of Trento (Italy), to address tidal network morphological development. In line with current theories as well as modelling advances and challenges in the field of tidal network ontogeny, emphasis was thus placed on the investigation of tidal channel formation and evolution in progressive marsh accretional context. Under these environmental conditions, tidal network development can be ascribed to the combination of two channel-forming processes: channel initiation results from bottom incisions in regions where topographic depressions occur; channel elaboration results from differential deposition, contributing to the deepening of the tidal channels relative to the adjacent marsh platform. Further evolutionary stages including channel reduction proceed from the horizontal progradation of the marsh platform which may lead eventually to channel infilling. Moreover, both qualitative and quantitative results allude to an acceleration of the morphological development of the synthetic tidal networks with increasing sediment supply. These different observations thus emphasise the prevalence of depositional processes in shaping tidal channels. In a second stage, the investigation was extended to the role of the initial tidal flat morphology as an inherent control on tidal network development, by considering different scenarios of topographic perturbations, which has revealed its legacy on tidal network morphological features. Modelling experiments have also acknowledged salt marsh macrophytes as a potential control on network evolution depending on their biomass distribution within the tidal frame. However, tidal channel morphodynamcis appears to be sensitive to the way biomass growth is mathematically parameterised in the model. In view of the current challenges in transcribing mathematically such a dynamic process and the relevance of bio-physical interactions in driving salt marsh and tidal network evolution, a field survey was conducted in a temperate salt marsh in the Netherlands, as part of the mobility to UNESCO-IHE (Netherlands) in partnership with University of Antwerp (Belgium), to assess vegetation distribution and productivity in the tidal frame. Particularly, emphasis was placed on extending investigations on the possible presence of relationships involving vegetation properties in different climatic and ecological conditions from those characterising these previously documented relationships. Regression analysis has revealed that biomass growth can be expressed as a linear function of marsh relative elevation, providing therefore direct empirical validation for corresponding assumptions reported in the literature and used in the present modelling framework; surprisingly, that increase did not correlate with an increase in species richness and diversity. Analysis of likely associations between vegetation morphometrics and total standing biomass yielded only a single linear relationship linking the latter variable to stem height. In truth, these observations may bear reconsiderations on the global validity of the assumptions used in the formulation of some eco-geomorphic processes which are applied in the study and prediction of wetland resiliency facing climate change.

Este estudo visa à descrição pontual do comportamento hidrodinâmico do estuário do Rio Mearim, analisando os padrões de corrente de maré, do registro de marés e das possíveis relações entre eles. Durante novembro e dezembro de 2012 um ADCP foi fundeado no canal do estuário, onde coletou dados por 31 dias. Os dados foram qualificados e processados para futura análise e inter-relações. Foi observado que a coluna de água estuarina no ponto de medição move-se como uma coluna de água homogênea com fluxo bidirecional no eixo NE-SW. As medições ocorreram no período de baixo índice de pluviosidade. O espectro de energia de velocidade de correntes indicou contribuições parciais dos harmônicos M2, M4, M6 e M8 no padrão de correntes encontrados. Defasagens no tempo entre os picos de máxima velocidade de corrente durante enchente e vazante mostraram a dominância do processo de maré vazante sobre a enchente no estuário. A análise integrada apresentada permitiu uma melhor compreensão no entendimento do comportamento das correntes forçadas pela maré, as chamadas correntes de maré, no estuário do Rio Mearim. / This study aim to describe the punctual hydrodynamic behavior of Mearim River estuary, analyzing the pattern of the tidal current, the tidal data and possible relationship between them. During 2012 november and december an ADCP was deployed in the estuary channel, where it collected data for 31 days. The data were qualified and processed for further analysis and inter-comparisons. It was observed the estuarine water column in the point of deployment moved as a homogeneous water column with bidirectional flux in axis NE-SW. The deployment was made in a period of low pluviosity index. The energy spectrum of current velocity indicated partial contribution of harmonics M2, M4, M6 e M8 in the current pattern found. Delays between the maximum velocity during flood and ebb conditions, showed the dominance of the edd process over the flood in the estuary. An analysis integrated presented on this paper allowed a better understanding of the behavior of the current forced by tides, called the tidal current in the Mearim River estuary.

Salt marshes provide a range of ecosystem services and yet are subjected to anthropogenic impacts that alter the biogeochemical processes underlying these services. In particular, human activities may modify salt marsh greenhouse gas (carbon dioxide, methane, nitrous oxide) emissions by changing plant and microbial communities, hydrological regime, and sediment chemistry. Quantifying the effects of human impacts on greenhouse gas emissions is important for complete carbon budgets, and for effective management of salt marshes and the ecosystem services they provide. In Chapters 1 and 2, I investigate the effects of hydrology and plant invasion on greenhouse gas emissions. First, I show how the restriction and restoration history of four salt marshes influence methane flux in unpredictable ways. Despite comparable salinity, methane emissions from one partially restored marsh were 25 times higher than unimpacted reference sites 13+ years after restoration, but emissions from other restored sites were equal or lower. Next, I show that greenhouse gas emissions associated with invasive Phragmites australis are not different from those associated with native Spartina alterniflora. These Chapters demonstrate the de-coupling of greenhouse gas emissions, and carbon sequestration more generally, from ecosystem degradation and restoration. In Chapters 3 and 4, I quantify greenhouse gas fluxes and microbial community structure under precipitation changes that may occur with global climate change. In a field experiment, doubled rainfall and drought had significant transient impacts on porewater salinity following storms, and on the community structure of plants (doubled rainfall) or microbes (drought), yet greenhouse gas fluxes and other biogeochemical processes were not affected. The absence of biogeochemical change indicates functional redundancy and resistance or resilience exist in the microbial community, suggesting marshes may continue providing services as precipitation changes. In a lab experiment, rewetting intact cores to simulate tidal inundation or rainstorms produced a nitrous oxide pulse 10-20x the baseline flux rates, without changing the microbial community. A model of rewetting event frequency suggests that pulsed emissions may be responsible for the majority of marsh nitrous oxide emission. Precipitation change may increase coastal nitrous oxide emission if it causes more or stronger storms, and thus more rewetting events.

The challenge of tackling global climate change and our increasing reliance on power means that new and diverse renewable energy generation technologies are a necessity for the future. From a number of technologies reviewed at the outset, the cross-flow tidal turbine was chosen as the focus of the research. The numerical investigation begins by choosing to model flow around a circular cylinder as a challenging benchmarking and evaluation case to compare two potential solvers for the ongoing research, ANSYS CFX and OpenFOAM. A number of meshing strategies and solver limitations are extracted, forming a detailed guide on the topic of cylinder lift, drag and Strouhal frequency prediction in its own right. An introduction to cross-flow turbines follows, setting out turbine performance coefficients and a strategy to develop a robust numerical modelling environment with which to capture and evaluate hydrodynamic phenomena. The validation of a numerical model is undertaken by comparison with an experimentally tested lab scale turbine. The resultant numerical model is used to explore turbine performance with varying Reynolds number, concluding with a recommended minimum value for development purposes of Re = 350 × 103 to avoid scalability errors. Based on this limit a large scale numerical simulation of the turbine isconducted and evaluated in detail, in particular, a local flow sampling method is proposed and presented. The method captures flow conditions ahead of the turbine blade at all positions of motion allowing local velocities and angles of attack to be interrogated. The sampled flow conditions are used in the final chapter to construct a novel blade pitching strategy. The result is a highly effective optimisation method which increases peak turbine power coefficient by 20% for only two further case iterations of the numerical solution.

Submitted by Jean Medeiros (jeanletras@uepb.edu.br) on 2016-08-12T12:17:58Z No. of bitstreams: 1 PDF - Renato Pereira Dantas.pdf: 2391944 bytes, checksum: 12056a11ac436bdd4e65fb9e44ddf75b (MD5) / Made available in DSpace on 2016-08-12T12:17:59Z (GMT). No. of bitstreams: 1 PDF - Renato Pereira Dantas.pdf: 2391944 bytes, checksum: 12056a11ac436bdd4e65fb9e44ddf75b (MD5) Previous issue date: 2016-02-03 / Coordenação de Aperfeiçoamento de Pessoal de Nível Superior - CAPES / The incorporation of a geographic perspective on ecology, has promoted insights were landscape models provide frameworks for understanding the hydrological and geological variability in estuarine regions. In this context, the geomorphological arrangements have explained the Nekton structure at different scales. These models are based on two hypothesis : i) the landscape determines the hydrodynamics and the configuration of the fiscal chemist variables ii) the geomorphology may affect the distribution and habitat use by species since certain geometries can support or attract higher densities. These factors have been observed in drainage systems that composed estuaries, the tidal creeks. In these sites, processes such as energy dissipation tidal stratification, tide and circulation of nutrients, are decisive factors in the structuring of the assemblages. In this context estuarine fish, they tend to have a significant energy expenditure adapting to the conditions of salinity and temperature. In this way, creeks can offer tidal and thermal refuge on the structures that form their landscape, playing a role in the persistence of the species during the tidal cycles. The narrowing of the channels geometry causes the dissipation of the tide, especially due to the braking caused by the friction of the banks. Mediated the channel narrows it had observed physical and chemical gradient associated with geomorphology. Thus assemblies may differ over different portions of the channels. For example, the hydraulic dynamics and hyaline circulation can be decisive in the presence of certain species. Soon varations in geomorphological measures, such as length and width, cause changes in the composition of estuarine fish assemblages. Then channels with different morphometric must differ in their population structure. Therefore, it had assumed therefore that the geomorphology inherent to drainage networks are directly connected to their function in the estuarine environment, so physical and chemical changes associated to landscape can attract fish species seeking for refuge, food and reproduction sites. For this reason the present work aimed to verify the way that the geomorphological features (length, width, split, connection to the main channel and meanders), in two second order creeks, affects the composition, distribution and abundance of fish assemblages over a salt gradient on a seasonal basis. Therefore, we made six excursions (3 in the dry and 3 in the rainy season). In each channel, were set four sample points, two of then represented the lower and other two the upper part. At each point were performed three hauls using a beach seine. Environmental factors were measured using a multiparameter meter. The geomorphometrie was obtained through satellite images A total of 6901 individuals included in 76 species of fish were caught. In this study, we found differentiate between areas next the connection with the main channel that show a greater richness and fish density. We attributed this pattern to the proximity of these zones with the main channel, the creek moth size and the width. The lower portions presented more stable in terms of salinity and dissolved oxygen, thus the upper zone was characterized by dominance of small gobiidae. / A incorporação de uma perspectiva geográfica a ecologia, tem promovido insights através dos quais modelos de paisagem fornecem quadros para o entendimento das variabilidades hidrológicas e geológicas nas regiões estuarinas. Nesse contexto, os arranjos geomorfológicos têm explicado a estrutura do nécton em diferentes escalas. Tais modelos fundamentam-se em duas hipóteses: i) a paisagem determina a configuração das variáveis fisioquímicas, e hidrodinâmicas, ii) a geomorfologia pode afetar a distribuição e uso do hábitat pelas espécies uma vez que determinadas geometrias podem suportar ou atrair maiores densidades. Estes fatores são observados nas redes de drenagem que compõem os estuários, os canais de maré. Nestes locais, processos como a dissipação da energia tidal estratificação, maré e circulação de nutrientes, são fatores decisivos na estruturação das assembleias. Em tal contexto peixes estuarinos, tendem a ter um gasto de energia significativo adequando-se as condições de salinidade e temperatura. Desta forma, canais podem ofertar refúgio tidal e térmico, nas estruturas que formam sua paisagem, desempenhando um papel na persistência das espécies durante os ciclos de maré. Os estreitamentos na geometria dos canais provocam a dissipação da maré, sobretudo devido principalmente a frenagem ocasionada pela fricção das margens. A mediada que o canal se estreita observa -se a formação de um gradiente físico e químico associado a geomorfologia. Assim, assembleias podem diferir ao longo de porções distintas dos canais. Por exemplo a dinâmica hidráulica e a circulação hialina pode ser determinante na presença de certas espécies. comprimento, acarretam mudanças na composição das assembleias de peixes estuarinos. Então canais com morfometrias distintas devem diferir quanto a sua estrutura populacional. Assume-se, portanto que a geomorfologia inerente as redes de drenagem estão diretamente ligadas a função das mesmas no ambiente estuarino, ou seja, variações físicas e químicas ligadas a paisagem atraem espécies de peixes em busca refúgio, alimento e reprodução. Por esta razão, o presente trabalho buscou verificar a maneira que as características geomorfológicas (comprimento, largura, divisões, conexão com o canal principal e número de meandros) em dois canais de maré de segunda ordem, afetam a composição, distribuição e abundância das assembleias de peixes ao longo de um gradiente salino em um regime sazonal. Para tanto foram realizadas seis excursões (3 no período seco e 3 no chuvoso). Em cada canal foram estabelecidos quatro pontos amostrais, dois dos quais representavam a zona inferior e dois a porção superior. Em cada ponto foram realizados três arrastos utilizando uma rede beach seine. Os fatores ambientais foram aferidos através de uma soda Logo variações nas medidas geomorfológicas, tais como largura e multiparamétrica. A geomorfometría foi obtida através de imagens de satélite. Um total de 6.901 indivíduos, pertencentes a 76 espécies, foram capturados. Nesse estudo constatou-se diferenciação entre as zonas, onde as regiões próximas a conexão com o canal principal paresentaram maior riqueza e densidade de peixes. Atribuiu-se tal padrão a proximidade destas zonas com o canal principal, tamanho da abertura da boca do canal bem como a largura do mesmo. As zonas inferiores apresentaram-se mais instáveis em termos de salinidade e oxigênio dissolvido, desta forma as zonas superiores foram caracterizadas pela dominância de pequenos gobídeos.