Knowledge of ecological characteristics of trees, sites and tree growth on different sites is fundamental for silvicultural decision-making and planning. With the biogeoclimatic ecosystem classification in place, silvicultural management in British Columbia has been given an ecological foundation; however, relationships between growth and site have not yet been fully investigated. The purpose of this study was to determine how height growth of Douglas-fir within the drier portion of the CWH zone varies with site. We adopted site index (m @ 50 yr bh) as a species-specific measure of forest productivity, recognizing that it indicates height growth performance at a selected point in time. If forest productivity is correlated with ecological measures of site quality, what site factors should be used to quantify the relationships? Because of compensating effects, the numerous site factors can be reduced to four primary (synoptic) factors that directly affect plant establishment and growth: climate (light and temperature), soil moisture, soil nutrients, and soil aeration (not used in this study).

The Mountain Hemlock (MH) zone includes all subalpine forests along British Columbia’s coast. It occurs at elevations where most precipitation falls as snow and the growing season is less than 4 months long. The zone includes the continuous forest of the forested subzones and the tree islands of the parkland subzones (Figure 1). Old-growth stands are populated by mountain hemlock, Pacific silver fir, and Alaska yellow-cedar, and are among the least-disturbed ecosystems in the world. Canopy trees grow slowly and are commonly older than 600 years, while some Alaska yellow-cedars may be up to 2000 years old. Understanding regeneration patterns in the MH zone has become increasingly important as logging continues towards higher elevations of the zone where snowpacks are deeper.

Pacific silver fir (Abies amabilis (Dougl. ex Loud.) Forbes) is an important timber crop species in coastal forests of B.C. Its range extends from sea-level to almost timberline, and from the hypermaritime region on the west coast of Vancouver Island to the subcontinental region on the leeward side of the Coast Mountains. With this relatively wide climatic amplitude, a large variability in the height growth pattern of Pacific silver fir can be expected, since climate is considered to be the most influential determinant of the trajectory of height over age of forest trees. This variability, however, is not reflected in the height growth curves and site index tables used to estimate Pacific silver fir site index, since the curves and tables were developed from low-elevation stands on Vancouver Island. Consequently, when these curves and tables are applied to high-elevation or submaritime stands, we get biased estimates of site index. Accurate estimates of site index are necessary for accurate yield predictions. Furthermore, they are essential for making rational decisions about whether to cut the forest in situations where potential tree growth is marginal, such as in high-elevation forests.

In BC, it is required that harvested areas be regenerated with a mixture of tree species whenever appropriate to the site. This policy is based upon the assumption that increases in stand productivity, reliability, and/or biodiversity can be achieved in mixed-species stands. However, the knowledge justifying this policy is at best incomplete. Differences in forest productivity of mixed-species stands have been attributed mostly to competition. However, an increasing number of studies are providing evidence to support alternate theories, in which positive plant interactions play a major role. Positive plant interactions are divided into two components: (i) competitive reduction through structural and physiological differences in above and below ground structures, and (ii) facilitation through any positive effect on the growing environment of one plant species by another. These theories have yet to be tested in forest ecosystems. The objectives of this study, with respect to naturally established, unmanaged, second-growth stands of western hemlock (Hw) (Tsuga heterophylla (Raf.) Sarg.), western redcedar (Cw) (Thuja plicata Donn ex D. Don in Lamb.), and their mixtures, were: (1) to review the mechanisms of positive plant interactions and their potential to occur in these mixtures, and (2) to compare the productivity of these three stand types, using relative and absolute yield.

The three major components in the site classification of the biogeoclimatic ecosystem classification system are: climatic regimes, soil moisture regimes (SMRs) and soil nutrient regimes (SNRs). Both SMRs and SNRs can be identified in the field using soil characteristics and indicator plants. In the case of SMRs a quantitative classification was also developed that allow comparison of SMRs in different subzones. However, similar quantitative classification has not yet been developed for SNRs. This pamphlet summarizes and compares the results of several regional studies conducted in different biogeclimatic zones. Each of theses studies aimes to develop a quantitative SNR classification (Table 1). The comparison will examine: (1) how well the field-based classification matches quantitative classification, and (2) which direct measures distinguish best between field-identified SNRs.

Potential uses of dry weight information for simulation, and sources of data, for simulation of forest fuels were discussed. Tree and crown component dry weights of 23 Douglas fir, 18 western hemlock, and 23 western red cedar trees expanded to 314 "model" trees were investigated. Data were collected on the University Research Forest near Haney, British Columbia. Multiple regression techniques were used for the analysis. The best four independent variables are listed and the possibility of retaining a single independent variable, with its implications, are discussed. All predicting equations were conditioned to zero intercept and statistically tested for significance by the F-test. Of the independent variables tested, the combined variable of breast height diameter squared times total tree height was most closely related to tree component dry weights. The product of diameter and crown width squared was found to be the best single variable to describe crown component dry weights for Douglas fir. The dry weights of western hemlock and western red cedar crown components were most closely associated with breast height diameter times crown length. Reliable estimates of tree component dry weights were obtained. Crown components were highly variable and widely dispersed about the mean which resulted in less accurate estimates. / Forestry, Faculty of / Graduate

The structure of a forest stand is characterized by: (a) species composition, (b) age, (c) size (diameter and height), and (d) spatial (horizontal and vertical) arrangement of the trees. Depending on the species, site, and disturbance history, the stand structure varies with time, thus providing a snapshot of a particular development stage. Research on growth and stand structure has shown that the spatial distribution of trees is one of the key determinants of stand productivity. Forest inventories and ecological surveys carried out in British Columbia (BC) have shown that the structure of naturally established, unmanaged stands varies from simple (single-species, single-storied, and even-aged) to complex (multi-species, multi-storied, and uneven-aged). Only a few studies have quantitatively characterized this range of structural complexity, with nearly all studies focusing on old-growth stands. BC forest policy requires that harvested areas be regenerated with a mixture of tree species whenever a mixture is suited to the site. This policy is based upon the assumption that under appropriate conditions, increases in stand productivity, reliability, and/or biodiversity can be attained in mixed-species stands. This assumption has not yet been tested for forest ecosystems. One mechanism by which different tree species can reduce crown competition for light is through vertical separation (the development of multiple canopy strata). Canopy stratification is not easily recognized in mixed-species stands, particularly when species have similar shade tolerance and height growth patterns, and no quantitative methods have been developed to detect stratification. The diameter frequency distribution of two-storied stands have been characterized by inverted J-shaped as well as modal curves. Although it would be more appropriate to characterize stand structure by height frequency distributions, these distributions have not been developed. We suggest that (i) a stand is stratified if there are distinct, quantitatifiable modes in the size distribution; either diameter, height, or crown height, and (ii) height or crown height distributions will be the most sensitive measures. To characterize the structure of western hemlock (Tsuga heterophylla (Raf.) Sarg.) (Hw) and western redcedar (Thuja plicata Donn ex D. Don in Lamb.) (Cw) second-growth stands, and to investigate its influence on tree growth, we (1) described and compared size (diameter, height, and crown height) frequency distributions in single- and mixed-species stands, (2) determined whether mixed-species stands develop a stratified canopy, and (3) examined whether interactions between hemlock and redcedar affect tree growth.

The influence of tree species on forest soils has been the subject of study for at least a century. Of particular interest have been western hemlock (Tsuga heterophylla (Raf.) Sarg.) and western redcedar (Thuja plicata Donn ex D. Don) – two of the most common tree species in coastal and southern British Columbia, but each with a different nutrient amplitude. It has generally been found that acid, mycogeneous Mor humus forms develop in hemlock stands, while less acid and more zoogenous Mormoder, Moder, or even Mull humus forms develop in redcedar stands. The objective of this study was to determine the influence of hemlock and redcedar, growing separately and together, on forest floor nutrient properties. The questions addressed were: (1) does each stand type have unique forest floor nutrient properties? and (2) can any forest floor nutrient property discriminate between stand types?

Trees compete for various resources such as sunlight, water, and nutrients, which can be expressed as numerical terms, called competition indices (CI). Competition between individual trees is correlated with their growth and mortality. Therefore, CIs are used as independent variables to develop, improve and modify growth and yield models. This study was conducted to test the effect of neighborhood competition on tree diameter growth among Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), western hemlock (Tsuga heterophylla (Raf.) Sarg) and red alder (Alnus rubra Bong.), in the Pacific Northwest Coast Range, USA. After testing seven distance-independent CIs and three distance-dependent CIs, only the distance-independent CIs were found to significantly affect the diameter growth model. Among them, CIs with basal area and diameter information were the most impactful. As a result, a simple CI was very effective in a model that accounts for the basal area information of different tree species.

This thesis is a case study using paleoethnobotanical analysis of Kilgii Gwaay, a 10,700-year-old wet site in southern Haida Gwaii to explore the use of plants by ancestral Haida. The research investigated questions of early Holocene wood artifact technologies and other plant use before the large-scale arrival of western redcedar (Thuja plicata), a cultural keystone species for Haida in more recent times. The project relied on small-scale excavations and sampling from two main areas of the site: a hearth complex and an activity area at the edge of a paleopond. The archaeobotanical assemblage from these two areas yielded 23 plant taxa representing 14 families in the form of wood, charcoal, seeds, and additional plant macrofossils. A salmonberry and elderberry processing area suggests a seasonal summer occupation. Hemlock wedges and split spruce wood and roots show evidence for wood-splitting technology. The assemblage demonstrates potential for site interpretation based on archaeobotanical remains for the Northwest Coast of North America and highlights the importance of these otherwise relatively unknown plant resources from this early time period. / Graduate / cohenjenny2@gmail.com