The common warthog (Phacochoerus africanus) has recently been recognised as an alien invasive species in the Eastern Cape and their population is increasing rapidly within the region. This then raises a concern as it is recorded that invasive species have negative impacts on both economic and ecological aspects of biodiversity in the receiving community. There are few studies that have documented the impacts of this species in the Eastern Cape. This study therefore seeks to determine the diet and population trends of this species in the Addo Elephant National Park (AENP) Main Camp and the results obtained can be correlated with previous studies and used to quantify the impacts of this species in the Eastern Cape. Warthogs are specialized grazers, it was therefore hypothesized that they will have potential impacts on grazing resources in the AENP Main Camp. The diet of common warthogs in the AENP Main Camp was determined through microhistological analysis of faecal material of samples collected seasonally. Population trends of common warthogs were determined in relation to the effects of predation on the population growth and population age structure of these animals within the AENP Main Camp. The results were compared with the findings for common warthog population growth and population age structure of Great Fish River Nature Reserve (GFRNR), a predator free population. It was hypothesized that the population growth rate of common warthogs in the AENP Main Camp before the introduction of lions and hyaenas in the AENP would not differ from the population growth rate of common warthogs in GFRNR. Secondly, the post-lion/hyaena common warthog population in the AENP Main Camp would differ from both the AENP Main Camp pre-lion/hyaena and GFRNR common warthog populations due to the presence of large predators in the AENP Main Camp. For population age structure it was hypothesized that the two populations, AENP Main Camp and GFRNR would differ because of predators increasing mortality. The diet of common warthogs was dominated by grass (87.4 percent), with Cynodon dactylon being the dominant grass species. Common warthogs in the AENP Main Camp should be properly managed because they have potential impact on grasses. The population growth of common warthogs in the AENP Main Camp showed no effect of predation, with population growth not differing from that of GFRNR. Predation had an effect on common warthog population age structure with AENP Main Camp and GFRNR populations differing, particularly in terms of adult structure. Thus, population growth of common warthogs in the AENP Main Camp is not determined by the presence of predators. These findings highlight the status of common warthog as an invasive species in the AENP Main Camp and potentially impacting on grass species and show little effects of top-down population regulation. These results show a need of monitoring common warthog population as well as their potential impacts in the area.

Impact assessment is a requirement for development in many countries across the globe, seeking to inform the decision-maker as to the environmental, social and economic impact of an ongoing or proposed project. Socioeconomic impact assessment (SEIA) is a means of informing decision-makers as to the socioeconomic effects a project could have, or is having, thus contributing to informing adaptive management practices. However, the tendency of socioeconomic impact assessment to highly quantitative economic methods of analysis raises the question of whether the desired results are achieved by the process. The purpose of the research was to determine whether highly quantitative forms of economic analysis are suitable for measurement of impacts in a social context where distributive as well as net impact is important; to critically analyze the method utilized in achieving highly quantitative economic impact assessment results; and lastly to draw conclusions and make recommendations regarding the efficacy of monitoring processes used to inform adaptive management practices. The research was conducted by means of a case study focusing on three SEIAs carried out on the same entity, namely the Addo Elephant National Park. Managed by South African National Parks (SANP), it began expanding its borders in the early 2000s. Funded by the World Bank, SANP was required to carry out a comprehensive Strategic Environmental Assessment (SEA) in 2003 to ensure the expansion did not have negative environmental, social and economic repercussions, and where such consequences were unavoidable, to ensure that mitigation and management thereof was informed by useful monitoring exercises. Given the need for resettlement and issues of economic distributive concern raised in the 2003 SEA, the three socioeconomic impact assessments conducted from 2005 – 2010 as part of the ongoing monitoring exercises formed an ideal framework for answering the primary research questions. The findings indicate that despite consistent terms of reference, different assessors interpret mandates from the commissioning body in different ways, leading to varied applications of the same theory, some methodologically better than others. Economic multiplier analysis was found to be inadequate as a measure of the distributive effects of economic impact. Moreover, a lack of consistency, accountability and transparency in the monitoring process led to three sets of results that were incomparable over time and thus inadequate as a means to inform adaptive management practices. Asymmetries of and between power and expertise in the commissioning body and the assessors led to breakdowns of the assessment process in terms of accountability and integrity and resulted in a failure to properly define the scope of the study and measure the relevant indicators. The following recommendations were made: that the economic multiplier method be complemented by additional methods of analysis when utilized in disparate social contexts where distribution of economic benefit is important; that monitoring practices be systematized at an early stage of the process to ensure comparable results useful in informing ongoing management practices; and that what an assessment measures and how it measures it be clarified with reference to an objective source. Finally, the number of factors for consideration in any impact assessment means that measurement of the full picture suffers resource constraints, emphasizing the need for impact assessment oversight to recognize the deficiencies of the process whilst still acknowledging that ‘some number is better than no number’.

Populations of small prey species (< 92 kg) can be regulated by competition, habitat-mediated responses, climate change, disease and predation. Populations of small prey species have declined in the Addo Elephant National Park Main Camp Section since the reintroduction of apex predators (lion and spotted hyaena). One possible explanation is predation by lion and spotted hyaena. However, small prey species are not generally considered preferred prey items of these predators, hence their contribution to predator diets needs to be explored. Global Positioning System (GPS) cluster analysis has emerged as a viable approach to locate lion kills for lion diet estimation in AENP, however this technique is known to bias diet estimates to large prey species. Understanding the impact of lion predation on populations of small prey in Main Camp therefore requires better estimates of lion diets. This study investigated the potential underestimation of small prey species in lion diet estimates by combining the data from scat analysis and carcasses located at GPS clusters. The adjusted lion diets were then used to investigate if the impact of lion predation could explain declines in small prey, by relating lion prey preferences to prey population trends in Nyathi. Using the results from Nyathi, it could then be inferred whether lions could have been responsible for the declines observed in Main Camp. Six GPS-collared lions provided location data used to construct individual movement paths, along which spatio-temporal clusters of GPS locations (n = 4896) were identified for ground investigation. Carcasses (n = 643) were recorded at the GPS clusters and scats (n = 256) were collected at GPS clusters with or without carcasses. Data from the located carcasses and scat collections were then used to create a history of feeding events for each collared lion in Main Camp-Colchester and Nyathi from a) carcasses alone or b) a combination of scats and carcasses, based on an estimated short and long lion throughout period. Relative prey availability was estimated based on camera trapping in Nyathi in high lion, low lion and no lion use areas, as lions may encounter some prey species more than other prey species. Jacobs’ index prey preference was used to investigate the importance of prey species to lion diets at three landscape scales: 1) Nyathi, 2) lion UD and 3) core lion UD. Lastly, prey population trends were investigated in Nyathi to provide an indication of possible changes in prey populations. Large prey species dominated the adjusted diets of lions in Main Camp-Colchester and Nyathi. However, when comparing the GPS carcass-derived lion diet estimate to the combined lion diet estimate, small prey were underestimated more often than large prey in the GPS carcass-derived lion diet estimate. Missed feeding events were therefore a function of prey body size. In addition, missed feeding events were also representative of the lion diets based on carcasses, with the tendency for species more often consumed to be missed more often in the diet record. However, even though small prey species were underestimated, they represented the minority of species consumed by lions and were predominantly avoided by lions in Nyathi. Lion prey preferences thus did not correlate to the declines in small prey species in Naythi, suggesting other potential drivers are important for small population trends in Nyathi. Lion predation cannot be ruled out as a driver of the decline in small prey species in Main Camp, due to different ecological conditions (e.g. vegetation density) in Main Camp compared to Nyathi, during the observed small prey population declines. Lion prey preferences in Nyathi revealed fine-scale variations in species specific preferences by lions, as well as site-specific differences in prey preferences when compared to published lion prey preference meta-analyses. These findings suggest that metaanalysis studies should be used as a guideline from which hypotheses about local predatorprey interactions can be developed rather than as an absolute measure of prey preference.

This study presents an investigation into the population dynamics and social structure of a small, closed elephant population. Specifically, it examined population growth rates for evidence of density-dependent regulation. It also quantified the association patterns of female elephants groups, and male elephants groups. Social structure was examined using Hamilton’s kinship theories of inclusive fitness, and age. Male-female patterns of association were also examined for inbreeding avoidance behaviours. The study population was located in Addo Elephant National Park, South Africa. Density-dependence was assessed using a long-term data set. Densities were considerably higher than estimated carrying capacities. Population growth rate was positively correlated with increasing density. No relationship between birth rate, the age of first calving or calf sex ratio and elephant density was detected but there was a positive relationship between birth rate and rainfall during conception year. Mortality rates, particularly for juveniles, were low, and mean inter-calf interval was 3.3 years. There is no evidence of density dependent regulation in this population. These findings indicate that density dependence should not be considered as an option in the control of elephant numbers in this Park, or where elephant resources are not seasonally limited. Examination of association patterns of the adult female component revealed that associations were not random at the population, family or individual scale. This is the second study on African elephants to confirm previous behavioural studies that predicted that preferred associates were close maternal relatives. This supports many studies showing that social species preferentially associate with their kin. The adult males in this population were found to have a well differentiated society with non-random associations. Generally, males were found to have weak associations with most other males and strong associations with only a few males. This association pattern was found to be persistent over the time frame of the study, as indicated by the time lag analysis. Males returned to their natal family, even when maternally related females were in oestrus. Oestrous females directed positive behaviours towards musth males. It appears that behavioural inbreeding avoidance mechanisms in this small, closed population are inhibited: musth status seems to override inbreeding avoidance. General principles from this case study were interpreted in terms of their applicability to other small, closed populations.

Elephants, as megaherbivores and keystone species have major impacts, both positive and negative, on ecosystems such as the Subtropical Thicket of the Eastern Cape. The feeding behaviour of elephants was quantified so as to firstly determine the preferred feeding heights of elephants, and secondly to determine if feeding behaviour and impact varies with size and sex. The preferred feeding height was determined with experimental feeding trials. Feeding height, pluck size, foraging rate and the proportion of plant material discarded were used to test for differences between elephants of different sizes and sexes in the Addo Elephant National Park. The influence of plant growth form on sex-specific feeding was also considered. Elephants preferred to feed at the lowest heights. The preferred feeding height was not related to body size. A wide range of feeding heights was recorded including and extending beyond, both the preferred and maximum feeding height of co-existing indigenous browsers. There was no difference in feeding height between the sizes and sexes. In free ranging conditions, all elephants fed at levels above the preferred foraging height when browsing. There was no difference in feeding behaviour between the different sized elephants, or between males and females. Adult elephants however exhibited larger ranges of feeding heights, pluck sizes, foraging rates and intake rates, including the maximum and minimum values. Growth form influenced the feeding of male and female elephants differently. Elephant feeding behaviour appears to limit opportunities for resource partitioning by way of feeding height segregation. Elephants are also capable of dominating the browse resources available to co-existing browsers by removing large amounts of forage at lower heights. The extent of elephant impact seems to be comparable between sizes and sexes, although adults and especially larger males are often able to use foraging opportunities that other elephants can not effectively utilize. Findings suggest that the feeding heights of elephants are determined by forage availability and reflect the diet in terms of browse or grass. The findings can be used, together with browse inventory methods, to determine browse resources available to elephants, and can also be used to develop monitoring tools to assess the impacts of elephants on plants.

Thesis (M.Sc. (Ichthyology & Fisheries Science)) - Rhodes University, 2009.

The Addo Elephant National Park (AENP) is the only existing national park situated in succulent thicket. This unique veld type is endemic to the Eastern Cape, and forms an important centre of endemism for small succulents and geophytes which comprise the subdominant component of the vegetation. It is regarded as the most threatened vegetation type in the Eastern Cape, as much of it has been severely degraded by farming activities and its rate of regeneration is very low. Thus the AENP represents an important sanctuary for certain species. However, large herbivores, particularly elephants, pose a further threat to the vegetation, and areas from which elephants have been excluded in the park (botanical reserves) have been shown to be more speciesrich than the surrounding vegetation. Most elephant-impact studies have focussed on the large shrub component of the vegetation, and only preliminary studies have been done in AENP to date. As the elephant population continues to grow, there is pressure to utilise the botanical reserves to increase the amount of available grazing. It is thus important that the botanical reserve system be highly effective and efficient in terms of area. Existing botanical reserves were established ad hoc and are therefore not necessarily optimal in this regard. An iterative reserve-selection algorithm was used to maximise plant species conservation in the most efficient area.

The tourism industry is becoming increasingly more competitive, making it essential for tourism destinations to understand why tourists travel to specific destinations. Travel motives are seen as important factors affecting tourist behaviour. Understanding the travel motives of tourists is essential information for the development of a tourism product and its marketing strategy. Tourists to different destinations have different internal reasons for doing so, and wish to fulfil certain needs. These needs motivate tourists to choose a certain destination and, therefore, identifying the travel motives of tourists to tourism destinations will help the destination to better target their marketing strategies to tourists with common motives. The aim of this study is to determine the travel motives of tourists to selected national parks in South Africa as well as to determine whether there are differences and/or similarities between these motives. The parks concerned are the Addo Elephant-, Augrabies Falls- and Golden Gate Highlands National Parks. This will assist SANParks in their future marketing efforts and help to better understand why the tourists choose to visit the respective parks. A literature review regarding tourist behaviour was carried out to achieve Objective 1, which was to conduct a theoretical analysis of tourist behaviour. The primary reasons for understanding tourist behaviour are to be able to retain your customer for life, to fulfil the tourists’ needs and to develop better target marketing strategies. To understand tourists’ travel motives better, studying tourist behaviour becomes important. The tourist’s decision-making process (that forms part of the tourism behaviour model) is divided into six stages. These are, awareness of a need; seeking information; evaluation of information; decision making; action or buying and postpurchase evaluation. The tourist behaviour model consists of a number of factors that influence this decision-making process, including internal (for example perception, learning and motivation) and external (for example culture, social class and family) influences as well as personal (race, gender and age) and market-related (climate, economy and government) influencers. The most important influence relevant to this study is motivation. To achieve Objective 2, which was to conduct a theoretical analysis of travel motives of tourists to nature based products, a literature review concerning the travel motives of tourists was carried out. This revealed that a number of different travel motives to tourism destinations have been identified. Internal needs motivate tourists to travel to specific destinations, in the hope of fulfilling these needs. Research on travel motives is based on a number of theories, and some of the important motives that arose were: nature, wildlife, escape, relaxation, education, novelty, socialisation, family recreation and destination attractions. In conclusion, there are similarities as well as differences in the travel motives of tourists to different natural areas. Objectives 3 and 4 included determining the profiles as well as the travel motives of tourists to the selected national parks. Secondary data was used to determine the profiles and travel motives of the tourists. The first section focused on the profiles of the tourists, which proved to be quite similar in all three parks, with minor differences regarding province of residence and the languages spoken. The second section focused on the travel motives of these tourists, and a factor analysis was used to extract six factors from the data. These factors were: relax and socialise, park attractions, nature and wildlife, knowledge-seeking, photography and park activities. The factors of tourists to each park were compared by means of a practical significance test (ANOVA) to determine the differences in travel motives to each park. The results revealed that certain travel motives were more important to certain of the three selected national parks, for example: knowledge-seeking is more important to tourists visiting the Addo Elephant National Park, relax and socialise is more important to tourists visiting the Augrabies Falls National Park, and park activities are more important to tourists visiting the Golden Gate Highlands National Park. By determining the differences and similarities in travel motives to the three selected national parks, the aim of this study was achieved. The results are in agreement with other researchers’ findings that there are a number of travel motives motivating tourists to visit certain destinations, and this study determined that these motives differ from destination to destination. / Thesis (M.Com. (Tourism))--North-West University, Potchefstroom Campus, 2012.

Tourism comprises the world's largest industry, and beaches (marine tourism) are considered as one of the major tourism attractions in the world. Marine tourism includes different aspects such as the ocean, coastal water sports, hotels and restaurants, island and beach resorts, recreation, fishing boat operators, cruise ships and charter yacht companies. One of the main contributors to marine and wildlife tourism in South Africa, is South African National Parks (SANParks), comprising of 21 national parks. Of the 21 National Parks six are marine national parks covering 4004.4 square kilometres namely West Coast, Wilderness, Tsitsikamma, Addo Elephant, Agulhas and Table Mountain National Parks. The two most known Marine National Parks in South Africa are Tsitsikamma and Addo Elephant National Parks: Addo Elephant for the fact that it is the only national park in South Africa hosting the BIG 7, namely elephant, rhino, lion, buffalo, leopard, whales and great white sharks. Tsitsikamma National Park is important as it is the first and oldest Marine National Park in Africa. South African National Parks receive thousands of visitors each year and tourists visiting these parks are a source of revenue for national parks and by determining travel motives, marketing can be undertaken more effectively, and specific factors can be taken into account when the marketing strategies are being planned. The literature review indicated that an understanding of tourist motives is a key aspect in understanding tourist behaviour. If the travel motives of tourists are known to product owners, in the case of South African National Parks, they will assist the product owner with product development as well as its marketing strategy. By understanding tourist behaviour one can understand why tourists buy certain products and why they make certain decisions. Tourist behaviour can be analysed in terms of the purchasing process a tourist goes through when making a decision. Different tourists will have different decision making processes and will have different motives for buying different products and services. To be able to understand how buyers proceed through the decision making process one has to look at the tourist decision–making process which consists of different influences like demographic factors, the marketing mix, internal/ psychological, external/social and situational factors. From the literature review, the following travel motivations theories were identified: Maslow's hierarchy of needs, Murray's classification of needs, Mill and Morrison's needs and wants theory, Push and Pull theories and the Sun lust and Wanderlust theory. It was found in the literature review that there are some travel motives that are found in most of the nature based destinations such as “relaxation and escape”, “learn about nature” and “park attributes” to name but a few. The research, however, also found that travel motives are closely linked to what these destinations offer and it was also found that different marine destinations seem also to have different travel motives. The aim of this study is, therefore, to determine the travel motives of tourists to selected Marine National Parks in South Africa. Secondary data was used to determine the visitor's profile as well as the factor analysis on travel motives. Four hundred and eight questionnaires were administered to the Addo Elephant National Park and 495 to the Tsitsikamma National Park. The data analysis consisted of two parts, the first part was descriptive data and here the profile of the visitors to the two selected marine national parks was discussed. In the second part a factor analysis was conducted. The aim of a factor analysis is to reduce the data and to assist in the interpretation of the data. A factor analysis further describes the variance–covariance relationship among a number of variables in terms of a few underlying but unobservable random quantities, called factors. For this research a principal axis factor analysis with Oblimin with Kaiser Normalisation rotation was performed on the 22 travel motive items. The visitors' profiles for both the parks were quite similar except for the following: The largest percentage of the visitors to Tsitsikamma National Park were Afrikaans speaking and not as in the case of Addo Elephant National Park most tourists were English speaking, The major markets for Addo Elephant National Park, were the Western and Eastern Cape Provinces and for the Tsitsikamma National Park the major markets were Gauteng and the Western Cape Provinces. The factor analysis regarding travel motives revealed the following four factors: Education and knowledge seeking, Relaxation and escape, Park attributes and Family togetherness. A comparison was done regarding the travel motives of tourists to the two national parks to determine if there were any differences. A small to medium effect size was found for “relax and escape” and “education and learning about wildlife”, therefore, a small to medium statistical significant difference were found. This research confirmed that different marine destinations have different travel motives. Therefore, marketers can use this information and focus on individual marketing for each park based on the products they offer. / Thesis (M.Com. (Tourism))--North-West University, Potchefstroom Campus, 2012.

The tourism industry is becoming increasingly more competitive, making it essential for tourism destinations to understand why tourists travel to specific destinations. Travel motives are seen as important factors affecting tourist behaviour. Understanding the travel motives of tourists is essential information for the development of a tourism product and its marketing strategy. Tourists to different destinations have different internal reasons for doing so, and wish to fulfil certain needs. These needs motivate tourists to choose a certain destination and, therefore, identifying the travel motives of tourists to tourism destinations will help the destination to better target their marketing strategies to tourists with common motives. The aim of this study is to determine the travel motives of tourists to selected national parks in South Africa as well as to determine whether there are differences and/or similarities between these motives. The parks concerned are the Addo Elephant-, Augrabies Falls- and Golden Gate Highlands National Parks. This will assist SANParks in their future marketing efforts and help to better understand why the tourists choose to visit the respective parks. A literature review regarding tourist behaviour was carried out to achieve Objective 1, which was to conduct a theoretical analysis of tourist behaviour. The primary reasons for understanding tourist behaviour are to be able to retain your customer for life, to fulfil the tourists’ needs and to develop better target marketing strategies. To understand tourists’ travel motives better, studying tourist behaviour becomes important. The tourist’s decision-making process (that forms part of the tourism behaviour model) is divided into six stages. These are, awareness of a need; seeking information; evaluation of information; decision making; action or buying and postpurchase evaluation. The tourist behaviour model consists of a number of factors that influence this decision-making process, including internal (for example perception, learning and motivation) and external (for example culture, social class and family) influences as well as personal (race, gender and age) and market-related (climate, economy and government) influencers. The most important influence relevant to this study is motivation. To achieve Objective 2, which was to conduct a theoretical analysis of travel motives of tourists to nature based products, a literature review concerning the travel motives of tourists was carried out. This revealed that a number of different travel motives to tourism destinations have been identified. Internal needs motivate tourists to travel to specific destinations, in the hope of fulfilling these needs. Research on travel motives is based on a number of theories, and some of the important motives that arose were: nature, wildlife, escape, relaxation, education, novelty, socialisation, family recreation and destination attractions. In conclusion, there are similarities as well as differences in the travel motives of tourists to different natural areas. Objectives 3 and 4 included determining the profiles as well as the travel motives of tourists to the selected national parks. Secondary data was used to determine the profiles and travel motives of the tourists. The first section focused on the profiles of the tourists, which proved to be quite similar in all three parks, with minor differences regarding province of residence and the languages spoken. The second section focused on the travel motives of these tourists, and a factor analysis was used to extract six factors from the data. These factors were: relax and socialise, park attractions, nature and wildlife, knowledge-seeking, photography and park activities. The factors of tourists to each park were compared by means of a practical significance test (ANOVA) to determine the differences in travel motives to each park. The results revealed that certain travel motives were more important to certain of the three selected national parks, for example: knowledge-seeking is more important to tourists visiting the Addo Elephant National Park, relax and socialise is more important to tourists visiting the Augrabies Falls National Park, and park activities are more important to tourists visiting the Golden Gate Highlands National Park. By determining the differences and similarities in travel motives to the three selected national parks, the aim of this study was achieved. The results are in agreement with other researchers’ findings that there are a number of travel motives motivating tourists to visit certain destinations, and this study determined that these motives differ from destination to destination. / Thesis (M.Com. (Tourism))--North-West University, Potchefstroom Campus, 2012.