No description available.

Thesis (PhD)--Stellenbosch University, 2010. / ENGLISH ABSTRACT: Biological invasions are a growing threat to biodiversity, trade and agriculture in South Africa. Though alien reptiles and amphibians (herpetofauna) are not currently a major issue, escalating problems worldwide and increased trade in South Africa suggest a possible increase in future problems. In this thesis I explore practical measures for risk assessment implementable under national legislation. I began by documenting record-keeping and legislative differences between provinces in South Africa. This revealed some serious deficiencies, complicating attempts to compile accurate inventories and discern import trends. International trade data, however, revealed an exponential increase in the number of imports to South Africa over the last 30 years. Characterising the abundance of species in this trade is important as species introduced in large numbers pose a higher establishment risk. In South Africa, I found a tendency for venomous and expensive species to be traded in low numbers, whereas species that are easy to breed and handle, or that are colourful or patterned are traded in higher numbers. Unlike South Africa, California and Florida have had a large number of well-documented herpetofaunal introductions. These introductions were used to verify the role of several key predictors in species establishment. I first evaluated the role of each variable separately. I examined different approaches for bioclimatic modelling, the predictive power of different sources of distribution data, and methods of assigning a climate-match score. I also present the first test of Darwin’s naturalization hypothesis for land vertebrates using two new phylogenies inferred for native and introduced reptiles in California and Florida. I then used boosted regression trees (BRT) to infer the relative contribution of each factor to species establishment success. Results from the BRTs were incorporated into a user-friendly spreadsheet model for use by assessors inexperienced in complex modelling techniques. Introduction effort was found to be the strongest contributor to establishment success. Furthermore, species with short juvenile periods were more likely to establish than species that started breeding later, as were species with more distant relatives in regional biotas. Average climate match and life form were also important. Of the herpetofaunal groups, frogs and lizards were most likely to establish, while snakes and turtles established at much lower rates, though analysis of all recorded herpetofaunal introductions shows slightly different patterns. Predictions made by the BRT model to independent data were relatively poor, though this is unlikely to be unique to this study and can be partially explained by missing data. Though numerous uncertainties remain in this field, many can be lessened by applying case by case rules rather than generalising across all herpetofaunal groups. The purpose for import and potential trade volume of a species will influence the threat it poses. Considering this in conjunction with a species’ environmental tolerances and previous success of species with similar life histories, should provide a reasonable and defendable estimate of establishment risk. Finally, a brief summary of the potential impacts of introduced alien herpetofauna is provided in the thesis. / AFRIKAANSE OPSOMMING: Indringer spesies hou ‘n al groter bedreiging in vir die biodiversiteit, handel en landbou van Suid- Afrika. Alhoewel uitheemse reptiele en amfibieërs (herpetofauna) tans nie ‘n groot bedreiging in Suid-Afrika is nie, dui groeiende probleme wêreldwyd asook 'n toename in plaaslike handel op moontlike toekomstige probleme. In hierdie tesis, ondersoek ek praktiese metodes vir risikobepaling wat onder nasionale wetgewing toegepas kan word. Ek begin deur die verskille in stoor van rekords en wetgewing tussen provinsies te dokumenteer. Hierdie proses het ernstige tekortkominge uitgewys, wat pogings om akkurate inventarisse saam te stel en invoertendense te bepaal, bemoeilik. Internasionale handelsdata het egter getoon dat daar ‘n eksponensiële toename in die hoeveelheid invoere na Suid-Afrika oor die laaste 30 jaar was. Die hoeveelheid spesies in hierdie handel is belangrik omdat spesies wat in groot hoeveelhede ingevoer word, ‘n hoër vestigingsrisiko het. In Suid-Afrika is ‘n tendens gevind vir handel in giftige en duur spesies teen lae hoeveelhede, terwyl spesies wat maklik teel, maklik hanteer kan word en kleurvol is of mooi patrone het, in groter hoeveelhede mee handel gedryf word. Kalifornië and Florida, in teenstelling met Suid-Afrika, het ‘n hoë aantal goed-gedokumenteerde gevalle van herpetofauna wat in die natuur vrygestel is. Hierdie introduksies was gebruik om die rol van verskeie belangrike faktore in die vestiging van populasies te bepaal. Eerstens het ek die rol van elke faktor apart ondersoek. Ek het verskillende benaderinge vir bioklimatiese model-bou ondersoek, die akuraatheid van verskillende bronne van distribusiedata getoets en drie metodes om ‘n “climate match score” te bereken, voorgestel. Ek bied ook die eerste toets van Darwin se naturalisasie-hipotese vir landwerveldiere aan, deur gebruik te maak van twee nuwe filogenieë wat ek gebou het vir inheemse en ingevoerde reptiele in Kalifornië en Florida. Ek het verder gebruik gemaak van “boosted regression trees” (BRT) om die relatiewe bydrae van elke faktor tot die vestigings-potensiaal van spesies te bepaal. Resultate van hierdie BRTs was ingekorporeerd in ‘n gebruikersvriendelike ontledingstaat wat deur bestuurders, onervare in komplekse modelboutegnieke, gebruik kan word. Invoer-hoeveelheid was die faktor wat die sterktste bygedra het tot suksesvolle vestiging. Verder is spesies met kort jeugperiodes en dié met verlangse familie in streeks-biotas, meer geskik om suksesvol te vestig. Gemiddelde klimaatooreenstemming en lewensvorm was ook belangrik. Paddas en akkedisse was die mees waarskynlikste van die herpetofauna groepe om populasies te vestig, terwyl slange en skilpaaie teen laer tempo’s populasies gevestig het, alhoewel analise van alle gedokumenteerde gevalle van herpetofauna-invoerings wêreldwyd effens verskillende tendense toon. Voorspellings wat deur die BRT-model vir onafhangklike data gemaak is was redelik swak, alhoewel hierdie resultaat onwaarskynlik nie uniek aan die studie is nie, en word gedeeltelik verduidelik deur die gebrek aan data. Alhoewel talle onsekerhede steeds bestaan, kan dié verminder word deur geval-tot-geval reëls toe te pas eerder as om vir herpetofauna as ‘n groep te veralgemeen. Die doel van invoer en potensiële handel-volumes van ‘n spesie, sal die bedreiging wat die spesie toon, bepaal. Hierdie faktore moet saam met omgewingstoleransie en voorafgaande sukses van spesies met soortgelyke lewenswyses oorweeg word, om ‘n aanvaarbare en verdedigbare beraming van vestigingsrisiko te gee. Laastens, word ‘n kort opsomming van die effekte wat uitheemse herpetofauna mag hê, verskaf.

Thesis (MSc (Geography and Environmental Studies))--University of Stellenbosch, 2007. / The Agulhas Plain, an area rich in fynbos, was monitored within six months after the February 2006 fire. The potential of using medium resolution imagery, specifically from the Moderate Resolution Imaging Spectroradiometer (MODIS) in determining the re-growth rates of indigenous and alien vegetation types after fire was explored. Pixels representing dense areas of each vegetation type were selected. There was a significant difference in the pixels selected for each vegetation type. A time series of Normalized Difference Vegetation Index (NDVI) data was derived and fitted to functions, such as Double Logistics and Asymmetric Gaussian as implemented in the TIMESAT software. The results show that alien vegetation grows faster after a fire occurrence than in its absence. Within the specified months of monitoring, it was observed that fynbos grew faster than the alien vegetation. Also, the re-growth rates of vegetation on the coastal soils were higher than those of vegetation on the inland soils. The determination of the re-growth rate was necessary to assist resource managers determine the appropriate time for follow-up of clearing invaded sites after fire.

There is little understanding of the implementation efficiency and effectiveness of restoring plant invaded landscapes within operational contexts. South Africa's Working for Water (WfW) programme is arguably the most ambitious alien plant control programme in the world, yet little is known about its cost-effectiveness and the challenges it faces in linking poverty and environmental objectives. My first aim was to assess the cost-effectiveness of invasive plant removal, and the factors that underpin its effectiveness over large spatial and temporal scales. The second aim was to compare the accuracy of evidence-based findings with managers' experience-based beliefs, and to assess whether managers are willing to change their beliefs after being exposed to it. The third aim was to assess the costs and benefits of removal versus removal followed by active native re-vegetation. My final aim is to assess the challenges and lessons learnt by managers linking ecological restoration with poverty alleviation objectives, specifically within the public works model. My study area was focused primarily on two WfW river catchment projects in the western region of the Eastern Cape province. I adopted an interdisciplinary approach drawing from a range of methods such as observational studies, statistical modelling and interviews with managers. The key findings were that control efforts in the two catchment projects are largely inadequate owing to many sites being re-invaded and not enough resources being allocated to the catchments. It would take between 54 and 695 years to clear the respective catchments. In terms of cost-effectiveness, my results exceeded previous estimates by 1.5 to 8.6 times for each catchment project. After being exposed to the evidence-based findings, the managers did not change their beliefs when it came to forecasting the future effectiveness. I found that active native re-vegetation after removal of invasive plants is very costly and that priority should be given to understanding the effectiveness of the removal treatments on native species recovery. The managers cited significant challenges in effectively and efficiently meeting the programmes dual objectives. Based on a broader review of the public works literature I recommend WfW re-examine the type of public works they currently use.

Species that become naturalized in a new geographical range, subsequently multiply and spread, and persist to the detriment of resident communities, are known as alien invasive species. Two aspects of species invasion – spread and ecological impact – were examined using Lantana camara L. (henceforth lantana) as the study system, specifically in the context of a seasonally dry tropical forest ecosystem of the Mudumalai Wildlife Sanctuary and National Park. Lantana is a thicket-forming woody shrub of South and central American origin, which is now widespread across the tropics. The thesis is divided broadly into two parts -the first part examines the influences of environmental factors on the distribution and spread of lantana while the second part focuses on the effects of lantana on the distribution, survival and growth of native woody species. Much of the work presented in this thesis was conducted within a 50 ha permanent plot (the Mudumalai Forest Dynamics Plot, MFDP hereon) in Mudumalai, chiefly because the history of invasion by lantana has been recorded here since 1989. The influence of changing resources on lantana invasion was explored at two scales -small spatial but fine temporal scale in the MFDP and at the level of the landscape. Available data on an 18 year chronosequence of changes in the qualitative density of lantana from the MFDP and field studies between the years 2009 and 2010 were used to determine the environmental correlates of lantana spread in time and space. It was found that biotic factors such as the presence of the shrub Helicteres isora and abiotic factors such as proximity to drainages and the combination of fire and drought promoted the intensification of lantana invasion in time while proximity to streams, higher total annual rainfall and low fire frequency contributed to lantana invasion at the landscape level. The impacts of lantana on the seedlings of native woody species were assessed in 10m x 10m plots within the MFDP. An initial enumeration of 60 such plots revealed that animaldispersed, dry forest habitat preferring species were most affected by the presence of dense lantana. A follow-up study comprising of growth and survival measurements made on 1105 seedlings over two years (2008-2010) further confirmed that dry forest preferring species were most affected by the presence of dense lantana and that this response at the community level was most likely influenced by the most abundantly sampled species in this habitat preference guild – Catunaregam spinosa. In conclusion, while the environmental correlates of lantana most likely promoted its invasion, only certain guilds of native species seemed to be negatively affected by the presence of lantana.

Thesis (PhD (Conservation Ecology and Entomology))--University of Stellenbosch, 2010. / ENGLISH SUMMARY: Climate change and biological invasions are major components of global change induced by human activity and are considered major drivers of global biodiversity decline in terrestrial ecosystems. These drivers interact synergistically and render ecosystems vulnerable to invasion by invasive alien species. Grasses are a group of invasive plants that easily respond to global changes and alter native plant community structure and ecosystem processes, such as fire frequency. To date there have been few studies that assess the ecological drivers and effects of invasive grass species on arid and semi-arid ecosystems of South Africa. Fountain grass (Pennisetum setaceum) is a widely distributed invasive alien perennial grass from North Africa and the Middle East, valued by horticulturalists worldwide. It spreads along the edges of roads on the outskirts of most towns, and is common on mine spoil in many areas throughout South Africa. Occasionally, it escapes into natural vegetation along drainage lines or after fires. This grass can be a costly problem for agriculture and biodiversity conservation as it is unpalatable and increases fire risk. Understanding its distribution and invasive dynamics will contribute to better management and control practices. The main objective of this study was to understand the ecology and invasion processes of P. setaceum across South African environmental gradients, and use it as a model to understand the synergistic relationships between biological invasions and other global change (climate and land use) scenarios. This study provides the first assessment of how P. setaceum overcomes different invasion barriers in South Africa as an emerging invader, in comparison with other parts of the world where it has already become problematic. A number of management and control options for this grass and other similar perennial grasses result from this study. Specifically, the study provides comprehensive understanding of: 1) the distribution and habitat preferences of P. setaceum in arid and semi-arid parts of South Africa, 2) environmental resources and habitat conditions that promote its invasive potential, 3) growth and reproductive performance across environmental gradients, 5) the response of an arid ecosystem to P. setaceum invasion and fire promotion, and 6) the predicted future distribution of this grass in South Africa under scenarios of climate change and spread. The study has found that road-river interchanges are ideal sites for P. setaceum growth and that these sites should be targeted for management and control of this species. P. setaceum was found to persist under diverse biome environments, which is attributed to local adaptation. Disturbance was found to be a major factor promoting fountain grass invasion into semi-natural areas away from roadsides. Major clean-ups of this grass should focus on disturbed areas, especially in the fertile parts of the fynbos region where the grass has high climatic suitability. Management and control should also focus on areas with high nutrients and extra water, as these areas facilitate growth and reproduction. The dynamics of P. setaceum invasion and spread makes it a good model for management of similar emerging invasive perennial grasses in similar ecosystems. / AFRIKAANSE OPSOMMING: Klimaatsverandering en biologiese indringing is grootskaalse komponente van mensgedrewe globale veranderinge, en kan ook gesien word as prominente drywers van die huidige afname in globale biodiversiteit van terrestriële ekosisteme. Hierdie drywers werk saam in sinergie, en laat sodoende ekosisteme kwesbaar vir indringing deur uitheemse indringerplante. Grasse is ‘n groep indringerplante wat maklik reageer ten opsigte van globale veranderinge en verander inheemse plantgemeenskapstruktuur en ekosisteem prosesse, soos byvoorbeeld veldbrand frekwensie. Tot op hede is daar wynig studies gedoen wat die ekologiese drywers en effekte van indringergras spesies in droë en semi-droë ekosisteme van Suid-Afrika assesseer. Pronkgras (Pennisetum setaceum) is ‘n wyd verspreide meerjarige uitheemse indringergras, oorspronklik vanaf Noord-Afrika en die Midde-Ooste, en word waardeer deur tuinboukundiges wêreldwyd. Dit versprei al langs padrande aan die buitewyke van meeste dorpe, en is algemeen op ou mynhope in verskeie dele van Suid-Afrika. Somtyds ontsnap hierdie gras langs dreineringskanale of na veldbrande, en beland so in die omringende natuurlike plantegroei. Hierdie gras kan ‘n duur probleem word vir landbou asook biodiversiteit bewaring omdat dit onsmaaklik is vir diere, en dit verhoog veldbrand risiko. ‘n Beter begrip van pronkgras verspreiding en indringingdinamika sal bydra tot verbeterde bestuur en kontrole praktyke. Die hoofdoel van hierdie studie was om die ekologie en indringing prosesse van P. setaceum regoor Suid-Afrikaanse omgewingsgradiente te begryp, en dit dan te gebruik as ‘n model om die sinergistiese verhoudings tussen biologiese indringing en ander globale veranderinge (klimaat en grondgebruik) te verstaan. Die studie verskaf die eerste assessering van hoe P. setaceum verskillende indringing hindernisse in Suid-Afrika oormeester as ‘n opkomende indringer, in vergelyking met ander dele van die wêreld waar dit al klaar problematies is. Hierdie studie verskaf uiteindelik ‘n aantal bestuur en kontrole opsies vir hierdie en ander soortgelyke meerjarige grasse. Die studie verskaf spesifiek ‘n deeglike verstaan van: 1) die verspreiding en habitat voorkeure van P. setaceum in droë en semi-droë areas in Suid-Afrika, 2) omgewingshulpbronne en habitat toestande wat hierdie plant se indringing potensiaal verhoog, 3) groei- en voortplantingsvertoning oor verskillende omgewingsgradiente, 4) die reaksie wat ‘n droë ekosisteem toon teenoor P. setaceum indringing en veldbrand verhoging, en 5) die toekomstige verspreiding van hierdie gras in Suid-Afrika onder voorspellings van klimaatsverandering en verspreiding. Die studie het gevind dat pad-rivier tussengange ideale liggings is vir P. setaceum vestiging, en dat hierdie liggings dus geteiken moet word vir bestuur en kontrole doeleindes. P. setaceum toon volharding onder diverse bioom omgewings, wat toegeskryf word aan plaaslike aanpassing. Daar is gevind dat versteuring ‘n groot faktor is in die verhoging van pronkgras indringing in semi-natuurlike areas weg van padskouers. Grootskaalse pronkgras uitroeiing projekte moet fokus op versteurde gebiede, veral in die vrugbare dele van die fynbos streek, waar hierdie gras hoë klimaatsgeskiktheid toon. Bestuur en kontrole programme moet ook fokus op areas met hoë nutriente inhoud en ekstra water, aangesien hierdie areas groei en voortplanting vergemaklik. Die dinamika van P. setaceum indringing en verspreiding maak dit ‘n goeie model vir die bestuur van soortgelyke opkomende meerjarige indringergrasse in soortgelyke ekosisteme.

Thesis (MScConsEcol (Conservation Ecology and Entomology))--University of Stellenbosch, 2010. / ENGLISH ABSTRACT: Riparian ecosystems of the Mediterranean south-western Cape region are projected to experience significant decreases in streamflow due to climate change and increased demands for water associated with human demographic trends and increasing living standards. Aggravating this problem are woody invasive alien plants, such as Acacia mearnsii, whose impacts, including those on catchment water yields, have justified extensive eradication programmes such as Working for Water (WfW). WfW has been highly successful in managing invasive alien species in some areas, while at the same time enhancing the quality of life of poor South Africans. Unfortunately, the managers of these eradication initiatives often experience a lack of information on the species- and site-specific ecological properties that may aid in prioritising sites more prone to invasion, essentially inhibiting accurate management output. Knowledge, such as that associated with the possible future distribution of invasive species in a changing environment, may help to optimise eradication initiatives. I therefore determined whether woody plants portray different physiological (vulnerability to cavitation) and wood anatomical traits (wood density, vessel resistance to implosion, vessel lumen and wall diameters) across three prominent riparian zones in the south-western Cape that each differ in streamflow quantity – to gain a mechanistic understanding of how woody species, especially invasive species, adapt their hydraulic strategy across this proxy for water availability. Using factorial ANOVA’s, to distinguish any differences in plant physiological and wood anatomical responses to water availability within and between species, showed Acacia mearnsii having consistently higher drought-tolerance (lower P50 values and denser wood) compared to native species under reduced water availability. These results supported the significant variation in drought-tolerance strategies that exist within and between taxonomically different species across different environments. Water availability thus has a strong selective effect on functional traits of species; however, minimum water potentials were more useful in describing in situ hydrological conditions than streamflow. Additionally, a non-causal relationship between wood anatomical traits and drought-induced cavitation was observed. Therefore, using only wood anatomical measurements to assess drought-tolerance of species might not be accurate across all species. This mechanistic approach to assess the invasive potential of species under projected drier conditions has great practical value. It can be used to improve species selection for restoration initiatives, and is of great value for future use in prioritizing eradication programmes. / AFRIKAANSE OPSOMMING: Rivieroewer ekosisteme in die Mediterreense Suidwes-Kaap streek kan groot afnames in stroomvloei verwag, wat meestal toegeskryf kan word aan klimaatsverandering en die verhoogde aanvraag na water geassosieerd met stygende populasiegrootte. Uitheemse indringerplante soos Acacia mearnsii vererger hierdie probleem. Laasgenoemde spesies plaas groot druk op opvanggebiede se water opbrengs, en vorm grootliks die rede waarom uitroeiing programme soos Werk vir Water (WfW) gestig is. WfW het al groot suksesse behaal in die bestuur van uitheemse indringerplante, en terselfdertyd die lewenskwaliteit van arm Suid-Afrikaners verhoog. Ongelukkig sukkel hierdie indringer uitroeiing inisiatiewe telkens met ongenoegsame bewyse wat kan help om leiding te verskaf by die prioritisering van areas meer geneig tot indringing. Die gevolg is dat onvoldoende bestuurs besluite gemaak kan word. Gevolglik sal kennis geassosieerd met die moontlike toekomstige verspreiding van hierdie indringerplant in ʼn veranderlike omgewing help om uitroeiing programme meer akkuraat te stel en gevolglik waardevol wees t.o.v. prioritisering van kwesbare areas. Die doel van hierdie studie was om die kwesbaarheid van sleutel inheemse houtagtige rivier spesies en veral die dreigende indringerplant Acacia mearnsii, te bestudeer in terme van verlaagde stroomvloei soos voorspel vir die Wes-Kaap fynbos ekosisteem. Deur te bepaal of houtagtige plante verskillende fisiologiese (kwesbaarheid teen xileem blokkasie) en anatomiese (hout digtheid, vat weerstandbiedendheid teen inmekaarkolking, vat holte en wand deursnee) hidrologiese eienskappe tentoonstel gemeet tussen drie prominente rivieroewer sones in die suidwes Kaap wat variëer in stroomvloei kwantiteit, verskaf ʼn meganistiese begrip van hoe spesies, veral die indringerplante, hul hidrologiese strategieë ten opsigte van droogtes kan aanpas. Faktoriële ANOVAS’s tussen spesies en liggings het getoon dat beide plant fisiologiese en hout anatomiese data beduidend variëer tussen taksonomies verskillende spesies. Veral Acacia mearnsii het deurgaans laer P50 waardes asook hoër houtdigtheid getoon, ‘n teken van hierdie spesies se verhoogde droogte toleransie in vergelyking met die inheemse spesies. Wat beduidend is is dat spesies-spesifieke lewensstrategieë ten opsigte van droogte toleransie in hierdie rivieroewer sones voorkom. Resultate het bevestig dat water beskikbaarheid ʼn groot rol speel in die seleksie van funksionele karaktereienskappe in plante, maar, dat minimum water potensiaal meer akkuraat was om in situ hidrologiese toestande te beskryf as wat stroomvloei kon. Die verhouding tussen plant fisiologiese en hout anatomiese karaktereienskappe was indirek as gevolg van spesies-spesifieke lewensstrategieë. Gevolglik sal die allenige gebruik van hout anatomiese eienskappe om droogte toleransie in plante te assesseer onakkuraat wees omdat daar ʼn swak verhouding tussen hout anatomie en droogte-geïnduseerde xileem blokkasie bestaan. ʼn Meganistiese benadering om die indringing potensiaal van spesies onder voorspelde droër kondisies te assesseer, asook om spesies seleksie vir restourasie doeleindes op te skerp, is prakties uitvoerbaar, en van groot waarde vir toekomstige gebruik in uitroeiing en restourasie inisiatiewe.

Thesis (MScConEcol (Conservation Ecology and Entomology))--University of Stellenbosch, 2011. / ENGLISH ABSTRACT: Invasive alien plants (IAPs) reduce streamflow and threaten the biodiversity of South Africa’s Cape Floristic Region. Up-to-date information on invasive vegetation is required for land management agencies to formulate policies and make appropriate resource management decisions. Invasion maps are typically not updated often enough because of the time and expenses required to do so. As a result, invasion maps for South Africa are limited to coarse resolution data or isolated small scale studies. Invasive alien plants change the landscape by destabilizing catchments and thereby increasing soil erosion, altering fire regimes and hydrology, as well as changing the physical and chemical composition of the soil. Information on IAPs is needed at a landscape scale. Remote sensing is a powerful tool that can be used to characterise landscapes in a biologically meaningful manner. The Normalised Difference Vegetation Index (NDVI) was used to create an up-to-date invasion map of the Agulhas Plain, lying at the heart of the species rich Cape Floristic Region. This information was combined with actual evapotranspiration data from the Surface Energy Balance Algorithm for Land (SEBAL) study done by Water Watch and the Council for Scientific and Industrial Research. The results showed that invasive vegetation uses more water than natural fynbos vegetation and that the greatest amount of water would be made available by clearing the invaded deep sands on the Agulhas Plain. These deep sand areas conflict with the priority areas of the Working for Water programme. This IAP eradication programme targets sparsely invaded upland areas for long-term sustainability. The recommendation of this study is to clear invaded wetland and riparian areas as these zones yield the greatest hydrological benefit per hectare and meet the priorities of Working for Water. Overall, 36 million cubic meters of water would be made available by clearing the Agulhas Plain. It can be concluded that there is a significant hydrological benefit to clearing invasive alien vegetation on the Agulhas Plain. / AFRIKAANSE OPSOMMING: Indringerplante (IP) verminder stroomvloei en bedreig die biodiversiteit van Suid-Afrika se Kaapse Floristiese Streek. Die nuutste inligting oor uitheemse plantegroei is nodig vir grondbestuuragentskappe om beleide te formuleer vir die neem van toepaslike hulpbronbestuur besluite. As gevolg van die tyd en uitgawes wat nodig is om indringingskaarte op te dateer, word dit gewoonlik nie dikwels genoeg gedoen nie. Dus is indringingskaarte vir Suid-Afrika beperk tot growwe resolusie data of geïsoleerde kleinskaal studies. Indringerplante verander die landskap deur opvangsgebiede te destabiliseer en sodoende te lei tot gronderosie, verandering van vuurregimes en hidrologie, sowel as die verandering in die fisiese en chemiese samestelling van die grond. Inligting oor IP is nodig op 'n landskapskaal. Afstandswaarneming is 'n kragtige tegniek wat gebruik kan word om landskappe op 'n biologies betekenisvolle manier te karakteriseer. Die Normalised Difference plantegroei-indeks (NDVI) is gebruik om 'n opgedateerde indringingskaart van die Agulhas-vlakte, wat in die hart van die spesiesryke Kaapse Floristiese Streek lê, te skep. Hierdie inligting is gekombineer met die werklike evapotranspirasie data vanaf die Surface Energy Balance Algorithm for Land (SEBAL) studie gedoen deur Water Watch en die Raad vir Wetenskaplike en Nywerheidnavorsing. Die resultate het getoon dat uitheemse plantegroei meer water gebruik as natuurlike fynbosplantegroei en dat die grootste hoeveelheid van hierdie water beskikbaar gestel sal word deur IP op diepsand op die Agulhas-vlakte skoonte maak. Hierdie diepsand areas is in konflik met die prioriteitsgebiede van die Werk vir Water-program. Hierdie IP uitroeiingsprogram fokus op yl ingedringde berggebiede vir langtermyn volhoubaarheid. Die aanbeveling van hierdie studie is om duidelik ingedringde vleilande en oewergebiede skoon te maak, siende dat hierdie sones die hoogste opbrengs en die grootste hidrologiese voordeel per hektaar bied, en voldoen aan die prioriteite van Werk vir Water. In totaal sou 36 miljoen kubieke meter water beskikbaar gestel word deur die skoonmaak van die Agulhas-vlakte. Dus kan dit afgelei word dat die verwydering van hidrologiese indringerplante op die Agulhas-vlakte 'n beduidende voordeel sal inhou.

<p align="justify"><i>Typologie des cours d’eau en Wallonie et caractérisation par la faune invertébrée</i><br>Vingt-cinq types de cours d’eau ont été définis en Wallonie d’après les termes de la Directive Cadre pour l’Eau en tenant compte des facteurs obligatoires et optionnels rassemblés sous trois critères : la taille (du « ruisseau » à la « très grande rivière »), la pente (faible, moyenne et forte), et cinq régions naturelles (la région limoneuse, le Condroz, la Famenne, l’Ardenne et la Lorraine belge, résumant les facteurs obligatoires altitude, latitude, longitude et géologie). Une approche par analyses multivariées appliquées à un grand nombre d’échantillons (listes taxonomiques) a permis de définir de façon plus précise certains types de rivières et de distinguer sept groupes typologiques montrant des assemblages faunistiques similaires.</p> <p align="justify"><i>Conditions de référence types et définition de l’état écologique des rivières en Wallonie à l’aide de métriques biocénotiques basées sur les invertébrés</i><br> Les listes faunistiques et les valeurs métriques de l’IBGN (Indice biologique global normalisé) issues du réseau d’évaluation de la qualité biologique en Wallonie ont été utilisées pour discriminer les types de rivières et pour définir les sites de référence, les conditions de référence et les limites des classes des états écologiques. Au sud du sillon Sambre et Meuse, impliquant quatre groupes typologiques de rivières, les valeurs de référence et la définition des états écologiques ont été basées et calculées sur un réseau de référence de sites de très bon état. Les « valeurs de conditions de référence » ont été définies par les médianes des métriques des sites de « très bon état ». Lorsque aucun site de très bon état n’était disponible, c’est-à-dire pour les groupes typologiques au nord du sillon Sambre et Meuse et pour la Meuse et la Sambre, les conditions de référence (ou le « potentiel écologique maximal ») et les limites de classes ont été basées sur des coefficients appliqués aux sites de « bon état » et sur jugement d’expert.</p> <p align="justify"><i>Exercice pilote d’inter-étalonnage</i><br> Cinquante échantillons ont été sélectionnés parmi les rivières du type d’inter-étalonnage européen R-C3 (petite taille, altitude moyenne, substrat siliceux) en Wallonie (correspondant aux « ruisseaux ardennais ») dans toute la gamme de qualités écologiques, du « très bon » au « mauvais » état. Chaque métrique de l’indice « Intercalibration Common Metrics » (ICM) a été comparée (par régression linéaire et polynomiale) aux valeurs de l’IBGN. La plus forte corrélation a été établie avec l’indice synthétique de l’ICM dont le R² très élevé (0,95) permet un inter-étalonnage précis et fiable. A un EQR (Equivalent Quality Ratio) de l’IBGN = 1 correspond un indice ICM très proche (= 1,004). En conséquence, la métrique « cote de l’IBGN » est proposée comme métrique pour l’évaluation de l’état écologique de la faune invertébrée en Wallonie.</p> <p align="justify"><i>Espèces exotiques et invasives</i><br>L’ouverture récente du canal à grand gabarit Main – Danube en 1992 et la navigation qui en résulte a permis à plusieurs espèces de macroinvertébrés ponto-caspiennes d’envahir successivement les bassins du Rhin et de la Meuse. De 1998 à 2000, le réseau de mesure a enregistré quatre nouvelles espèces exotiques dans la Meuse en Belgique : une polychète (<i>Hypania invalida</i>) et trois crustacés (<i>Hemimysis anomala, Dikerogammarus villosus, D. haemobaphes</i>). Peu avant, en 1995, le bivalve asiatique <i>Corbicula fluminea</i> et l’amphipode nord-américain <i>Crangonyx pseudogracilis</i> ont également été enregistrés pour la première fois. La dynamique des invasions a été étudiée et discutée.</p> <p align="justify">L’<i>état biologique actuel</i> (2000-2002) des masses d’eaux de surface en Wallonie et l’évolution des états biologiques sur une décennie (1990-2002) figurent dans l’étude et sont discutés. Une amélioration globale de la qualité de 6 % est enregistrée pour cette période. L’amélioration de la qualité se manifeste principalement pour les états « mauvais » et « médiocre » s’élevant à l’état « moyen ».</p> <b>Abstract</b> <p align="justify"><i>River typology in Wallonia and invertebrate fauna characterization</i><br>Twenty-five river-types in Wallonia were defined according to the Water Framework Directive taking into account obligatory and optional factors gathered in three criteria: the size (from “brook” to “very large river”), the slope (gentle, medium and strong) and five natural regions (Loess, Condroz, Famenne, Arden and Jurassic summarising the obligatory altitude, latitude, longitude and geology factors). A multivariate approach applied to a large amount of samples (i.e. taxa lists) led to give a more accurate definition of some river-types and to discriminate seven river-type groups exhibiting similar faunal assemblages.</p> <p align="justify"><i>Type-specific reference conditions and ecological status definition of rivers in Wallonia using invertebrate biocenotic metrics</i><br> Faunal lists and IBGN (i.e. "Standardized Global Biological Index IBGN") metric values, evolving from the biological quality assessment network in Wallonia, were used to discriminate the river types and to define the reference sites, the reference conditions and the status class limits. South of the axis made by the Sambre & Meuse Rivers, involving four river-type groups, reference values and ecological status definition were based and calculated on a reference network of sites of high status. The "reference condition values” were defined as the median values of the metrics in all “high status” sites. Where no site of high status was available, i.e. in the others river-type groups north of the same axis and in the Rivers Sambre and Meuse themselves, reference conditions (or maximum ecological potential) and class limits were based on coefficients applied to “good status” sites and on expert judgement.</p> <p align="justify"><i>Intercalibration pilot exercise</i><br> Fifty samples were selected from R-C3 rivers (i.e. small, mid-altitude, siliceous) in Wallonia (corresponding to the “Arden’s brooks”) showing the widest range of ecological quality from high to bad status. Each Intercalibration Common Metric (ICM) was compared (linear and polynomial regression) to IBGN scores. The highest correlation was found with the synthetic ICM index, whose high R² (0.95) allows accurate and reliable intercalibration. To IBGN EQR (i.e. Equivalent Quality Ratio) = 1 corresponds a very close ICM index value (= 1.004). Consequently, the metric IBGN score is being proposed to act as the metric for the assessment of the invertebrate fauna ecological status in Wallonia, Belgium.</p> <p align="justify"><i>Exotic and invasive species</i><br> The recent opening of the canal Danube – Main in 1992 and the subsequent navigation allowed several Ponto-Caspian macroinvertebrate species to invade successively the Rhine and the River Meuse basins. From 1998 to 2000, the monitoring network recorded four new alien species in the River Meuse in Belgium: one Polychaeta (<i>Hypania invalida</i>) and three Crustacea (<i>Hemimysis anomala, Dikerogammarus villosus, D. haemobaphes</i>). Earlier, in 1995, the Asian Bivalvia <i>Corbicula fluminea</i> and the North American Amphipod <i>Crangonyx pseudogracilis</i> were also recorded for the first time. The invasions’ dynamics were studied and discussed.</p> <p align="justify"><i>Present biological status</i> (2000-2002) of surface water-bodies in Wallonia and biological status evolution over a one-decade period (1990-2002) is included and discussed in the study. A global quality improvement of 6 % is recorded for the period. Quality improvement concerns mainly the “bad” and “poor” status raising up to “moderate” status.</p>

<p>Arundo donax is an invasive species that mostly impacts on sensitive riparian ecosystems. Arundo is also invasive in South Africa, though less is known about its ecology, biology, and impacts. Since California and the Western Cape of South AFrica have similar Mediterranean-type climates, we could assume that the impacts of Arundo on ecosystems in California are likely to be similar in the Western Cape, and that control methods used could be extrapolated for use in South Africa. This thesis attempted to determine what impact Arundo has on biodiversity at the community-ecosystem level.</p>