The variable plumage characteristics of the sub-adult male cohort of the red-winged blackbird (Agelaius phoeniceus) are described. At one extreme of the plumage variation, there are a group of sub-adult males that are indistinguishable from the adult males. These birds can only be correctly aged by cloacal examination for the bursa of Fabricius in the autumn. At the other extreme, approximately 4% of the population are near perfect female-mimics. A simple scoring system based on the interspersion of dark feathers in the epaulet is presented, which permits classification of the sub-adult males into 1 of 6 plumage classes. These epaulet classes are significantly correlated with other traits of the plumage. / Age when entering the prebasic molt, and the physical condition of the sub-adult male may influence the development of the varied plumage characteristics. / The distribution of the plumage characteristics of a population of sub-adult males collected at a major blackbird roost in the province of Quebec is described for birds captured in the fall and spring. The spring plumage characteristics are more variable than the fall, a phenomenon which is not consistent with plumage wear as has been previously reported, but which may be related to a prealternate molt which the birds undergo. The prealternate molt is prevalent in some but not all of the contour feather tracts, and is restricted to females and the sub-adult male cohorts. / An age- and sex-specific spring migration of red-winged blackbirds is examined. Adult males arrive to the spring roosts first, followed by yearling males and then females. A similar pattern of dispersal to the breeding territories is described. Within the subadult male cohort, the birds with the most adult-male like plumage traits arrive at the vernal roosts first. / The characteristics of the prealternate molt and differential spring migration are discussed in relation to the pressures of sexual selection on the respective age and sex cohorts. / The adaptive significance of variable sub-adult male plumages and delayed plumage maturation in passerines is evaluated. Several competing hypotheses have been advanced to describe the phenomenon of delayed plumage maturation. These hypotheses are reviewed, and a test is proposed which unequivocally differentiates between the various alternatives. (Abstract shortened with permission of author.)

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My thesis seeks to examine the relationship that exists between queer selfidentification and heterosexual hegemonic/heteronormative power in three works of and about the AIDS era. Working from feminist and queer theory perspectives, I first chart the way in which a problematic identity—be that identity a non-identity of utter invisibility, a sick identity, a dangerous identity, or (most commonly) an identity of utter hedonism disconnected from any notions of attachment, affection, or love beyond the physical sexual act—has been and is still wholly adopted by some. I do this principally with a close reading of Renaud Camus' 1981 novel Tricks, as well as with substantial historical grounding. I assert that this is not just a problem in queer literature, but in queer life which queer literature deeply reflects. Through a close reading of Tony Kushner's play Angels in America: A Gay Fantasia on National Themes, I seek to illustrate the consequences of accepting entirely and without question a constructed and problematic identity for gay men. Historical examination also comes strongly into play through correspondence and personal narratives of men who lived through (and died in) the AIDS era, casualties of war of queer self-definition. Employing a close literary analysis of Larry Duplechan's 1986 novel Blackbird, my thesis seeks to chart a way to a stable, holistic, queer identity negotiated from a position of strength. In a larger sense my thesis explicates constraints upon queer identity intended to limit queer people to a heteronomous, damaged, vulnerable social position. I raise awareness of these constraints in attempt to navigate a way around them with the ultimate destination of this navigation being a perpetually increasing humanization of a historically and institutionally dehumanized population.

The biology of the red-winged blackbird (Agelaius phoeniceus) was studied in relation to variations in agricultural landscapes of southwestern Quebec, eastern Ontario, and southwestern Ontario by examining the relationships between corn (Zea mays) production and population size, distribution of territorial males, and morphology. Dietary studies showed consistent use of corn during their 6-9 month residency in these regions, and during winter months. / A positive correlation was found between independent estimates of the density of males and counts of red-wings obtained from the North American breeding bird survey (BBS). Density of males was a useful indicator of the number of red-wings in an area, whereas the BBS was a useful indicator of change in regional population level. / Historical increases in populations of red-wings in these regions were correlated with increases in waste grain (corn) following harvest. Greater rates of population increase in Quebec (compared with Ontario) corresponded to higher rates of increase in waste corn abundance and landscape heterogeneity. In Quebec, cultivation (mainly corn) was the most important habitat influencing density of males. Wetland and hay field abundance were also important. Heterogeneous landscapes incorporating crops, wetlands and hay fields attracted the most male red-wings regardless of population level, and breeding males preferred wetlands and hay fields for territory establishment. Fields were used in proportion to abundance, whereas forest, crops and areas of human occupation were avoided. At high population density, proportionately more males settled in areas of human occupation and in fields; these habitats were less suitable than wetlands or hay fields. Packing of males into a preferred hay field habitat occurred. / Overwinter mortality did not favor small body size in male red-wings but evidence of stabilizing selection on female body size was found. Sexual dimorphism was slightly more pronounced in spring than in fall. Historical changes in the body size of red-wings were not clearly related to increasing abundance of food (corn). If population size tracks increases in food abundance, then inter-male competition for breeding space may intensity; thus, large size may be limited by energy contraints imposed on males by sexual selection pressure regardless of food availability.

Obligate brood parasites rely solely on other species, the hosts, to incubate their eggs and raise their offspring, which often reduces the host’s reproductive output. This reproductive cost has led to the evolution of anti-parasite adaptations among hosts, which in turn, has led to better trickery by parasites, a process termed an evolutionary arms race. The objective of this thesis was to investigate host-parasite coevolutionary arms races to address questions of host-use diversity. Host diversity varies dramatically among brood-parasitic species, but reasons for variations in host-use among brood parasites are not well understood. In Chapter 2, I address questions on host diversity specifically, whereas I address questions about coevolutionary interaction between hosts and parasites in Chapters 3, 4 and 5 using two host-parasite systems, one in New Zealand and one in North America. Chapter 2 investigates if host diversity is constrained by aggressive nest defence behaviour. I compared the nest defence behaviour of the exclusive host of the shining cuckoo Chrysococcyx lucidus lucidus on the main islands of New Zealand, the grey warbler Gerygone igata, to two other potentially suitable hosts that are not currently parasitised, the fantail Rhipidura fuliginosa and the silvereye Zosterops lateralis. The results suggest that grey warblers are as aggressive as fantails and silvereyes towards shining cuckoos at the nest and thus, host specialisation in shining cuckoos in New Zealand, at least, does not appear to be the result of nest-defence constraints imposed by potential but unused host species. Chapter 3 investigates if red-winged blackbirds Agelaius phoeniceus, a species that typically accepts the eggs of parasites, recognises, as indicated by changes in incubation behaviour, when they have been parasitised by brown-headed cowbirds Molothrus ater. Recognition without rejection suggests that rejection may be context-dependent but the results suggest that red-winged blackbirds do not recognise when their nests have been parasitised by brown-headed cowbirds, at least at the egg stage. This study was the first to investigate if hosts that almost invariably accept the eggs of parasites recognise when they have been parasitised. Chapter 4 investigated the possibility of coevolutionary arms races occurring through olfactory channels in contrast to earlier work that focussed only on visual and auditory cues. Recent research has revealed that olfactory abilities in birds are more common than previously thought. Uropygial gland secretions are posited to be a key source of avian body odour and its composition has been found to vary among species and individuals as well as between the sexes. I compared gas-chromatography (GC-FID) traces of shining cuckoo preen wax to the GC-FID traces of the grey warbler, the only host of the shining cuckoo in mainland New Zealand, as well as the preen wax of seven other species for evidence of mimicry. Preliminary results suggest there is evidence for mimicry and the potential for odour-based nestling discrimination in grey warblers. Further tests recording the response of grey warblers to odour-manipulated nestlings are necessary. Finally, in Chapter 5, I investigated the response of the song thrush Turdus philomelos, a species that rejects the eggs of the common cuckoo Cuculus canorus and conspecifics at intermediate and low frequencies, respectively, to nest-odour manipulations using the preen wax of conspecifics and heterospecifics. The results suggest song thrush do not use odour to assess the risk of parasitism at least as indicated in terms of changes in incubation behaviour. Investigations of the role of olfaction in avian brood parasite systems can provide a better understanding of brood-parasite coevolution. Only by considering all channels of communication can we be sure to completely understand the coevolutionary dynamics between brood parasites and their hosts.

The objectives of the study were to determine the year-round distribution and habitat selection of four marsh passerine species, as described by vegetation type and its physical characteristics, and to investigate interspecific competition as a factor in the habitat selection of the Long-billed Marsh Wren and the Red-winged Blackbird. Fulfillment of these objectives should help the B.C. Fish and Wildlife Branch to assess the effects of their management plans on marsh passerines. I determined the abundance of four passerine species in different vegetation types via marsh census transects over the period of one year. Nesting habitat and breeding information on marsh wrens and redwings was obtained via nest study plots. The results were: Areas traversed from the central and deepest marsh outward towards higher ground are inhabited by redwings, marsh wrens. Common Yellowthroats, and Song Sparrows, respectively. Habitats of wrens and redwings are spatially and temporally segregated. Wrens use the "Dense Spiraea" (Hardhack shrub) habitat extensively, while redwings use it hardly at all in the marsh. Similarly, redwings are abundant in Scirpus cyperinus (Hairy-seeded Bulrush), whereas wrens seldom use this habitat. Both wrens and redwings use other vegetation types, but redwings use them in the late spring, whereas wrens use them in the summer. Both wrens and redwings have lower reproductive success when nesting in close proximity than when nesting apart. This supports the hypothesis that interspecific competition exists and indicates that selection may favour habitat segregation. I also found mutual suppression of reproductive success when the effects of different vegetation types were removed. The results were not statistically significant however, possibly due to small sample sizes. Of the observed differences between the physical nest site parameters of wrens and redwings, vegetation height and presence and/or depth of water require further investigation as factors in the habitat selection of the two species. Bren-wren competition does not appear to affect wren nesting success in the present study marsh. Management recommendations are given based on the objective of maintaining and possibly enhancing bird species diversity in the study marsh. / Science, Faculty of / Zoology, Department of / Graduate

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The epaulets of male Red-winged Blackbirds (Agelaius phoeniceus) function in both intersexual and intrasexual contexts, but there is little evidence that they covary with reproductive success in this well-studied species. We used path analysis of male/territory traits, mating success, and reproductive success of unmanipulated males to estimate current directional selection. Territory size had a positive effect on number of within-pair mates. Number of extra-pair mates had a positive effect on number of extra-pair fledglings, and number of within-pair and extra-pair fledglings had positive effects on total number of fledglings. We also reddened epaulets of free-living territorial males to determine whether manipulated and control males differ in territorial behavior, mating success, or reproductive success. Compared with control males, males with reddened epaulets incurred elevated rates of trespassing and territorial challenges, lost their territories more often, and were unable to produce extra-pair offspring. Despite these differences, however, the realized reproductive success of experimental and control males did not differ significantly, perhaps because males with reddened epaulets devoted more time to anti-predator vigilance and were more aggressive toward a simulated predator than were control males. An apparent lack of current sexual selection on epaulet color may be a form of counter-balancing sexual selection in which male aggression against redder epaulets opposes female preference for redder epaulets, or it might be the result of males compensating for lost extra-pair fertilizations by increasing their parental care.

In an effort to determine type, extent, and distribution of agricultural damage by blackbirds, county-wide damage appraisals were conducted in the fall of 1960. Crops checked included corn, peanuts, and milo. Based on the 96 fields examined throughout Nansemond County, the average monetary loss to the corn crop was slightly less than $.05 per acre. This loss can be reduced by planting a variety with a high degree of ear drooping and planting as early as possible. The peanuts appeared to be the crop most heavily damaged. An estimated 50 per cent of the crop was picked before blackbirds arrived. Of the part of the crop left exposed after the arrival of the blackbirds, only a small portion sustained damages amounting to above $5 per acre. By harvesting peanuts at an early date, losses to blackbirds can virtually be eliminated. The 1960 milo crop was not damaged by birds nearly so severely as previous crops were reported to have been. Anthracnose, a fungal disease, accounted for about 86 per cent of the damage previously called "bird damage." An estimated 15 per cent of this crop was destroyed by anthracnose, insects, and birds. In the case of all three crops, it was noted that the first and most severe damage was to those fields nearest nesting and roosting habitat. / Master of Science