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Includes amendments (8 p.) in pocket / Bibliography: leaves 306-331 / iv, 349 leaves, [34] leaves of plates : ill ; 30 cm. / Title page, contents and abstract only. The complete thesis in print form is available from the University Library. / Thesis (Ph.D.)--University of Adelaide, 1988

The ecology, life history, and population dynamics of the sagebrush defoliator, Aroga websteri Clarke, were studied in the field and laboratory. The defoliator has one generation a year at the Curlew Valley site. It overwintered in the egg stage and passed through five larval instars. Ten parasite species attacked the defoliator at the study site . Four species, Orgilus ferus, Phaeogenes sp., Spilochalcis leptis, and Apanteles cacoeciae, contributed over 75 percent of the total incidence of parasitism. Parasitism ranged from 20 to 76 percent in 1971, but only ranged from 6 to 29 percent in 1972. This decrease in total parasitism in 1972 coincided with a five-fold increase in the defoliator population. In 1972, many mature larvae died as a result of food shortage. A microsporidian infection and a predaceous beetle also caused variable mortality during the two years. Methods for determining defoliation to sagebrush plants were also studied in the field. The consumption and utilization of food by the fourth and fifth larval instars were determined. The fecundity, rate of development, and behavior of the insect were also investigated under laboratory conditions. Partial life tables were constructed from the findings of 1971 and 1972 to assess the r ole of various mortality factors in regulating the sagebrush defoliator numbers.

The annual demography of a population of antelope ground squirrels in Curlew Valley, Utah was studied by measuring population density, natality, and mortality. Capture- recapture techniques yielded lower estimates than the Hayne strip- census. The estimates suggested 1968 was a year of population decline. One hundred seventy- three squirrels were collected and autopsied to obtain sex ratio , age structure, natality, and mortality data . The seasonal sex ratios for adult and yearling squirrels showed 82 percent females in spring 1968, this gradually changed to 56 percent by winter . The seasonal sex ratios of the young squirrels showed 59 percent females upon emergence from natal burrows in summer 1968, increasing to 79 percent by winter. The squirrels collected were aged by cementum annuli . The oldest were believed to be 5 years old . They composed 1.2 percent of the collection while young- of- the-year composed 38. 7 percent. The estimate of mean corpora lutea was 7.6, the mean embryo count was 7.2, and the mean post-partum placental scar count was 5.8. The mean corpora lutea count for yearlings was statistically lower than the count for adults and the mean ovulation rate for 1968 statistically lower than the rate for 1969. An estimate of litter size from four nests gave a mean of 4.5. In 1968, the conception rate was 91 percent with yearlings comprising 75 percent of those not conceiving. In 1969, the rate was 100 percent. Pre-Emergence mortality of the young was 41 percent. post-emergence mortality was 86 percent, and mortality, March - November, 1968 was 92 percent. Adult and yearling spring-to-spring mortality for study area squirrels calculated from retrap data was 81 percent. Spring-to-spring mortality calulated from age distribution data and Ricker's formula was 71 percent. The spring-to-fall mortality for 1968 calculated from density estimates and age distribution data was 53 percent.

Stable carbon isotope ratios of soils, roots and litter along transects stretching from nearly monospecific stands of Ceratoides lanata , a species possessing c3 photosynthesis, to nearly monospecific stands of Atriplex confertifolia, a species possessing c4 photosynthesis, were analyzed to determine i f changes in the relative dominance of the shrubs have occurred in salt-desert communities. The a13c value , which reflects the proportion of 12c and 13c in a sample of plant tissue, can be used to distinguish between c3 and c4 species. Atriolex confertifolia and Ceratoides lanata have a13c values of -13.0 ofoo and - 25 ofoo, respectively. The a13c value of litter and roots was used as a measure of current community dominance. The a13c value of soil organic matter was employed as a measure of past community dominance . The differential between a13c values of roots and soils was consistently about 3.0 ofoo in Ceratoides- dominated stands. Root a13c values were always more negative than soil a13c values. The striking uniformity in the rootsoil differential l in Ceratuitles- duninateci stands is most likely the result of fractionation of carbon isotopes during decomposition. The differential between a13c values of roots and soils in Atriplexdominated stands was more variable. This most likely indicates a lack of long- term community stability in areas na The distribution of a13c values in relation to depth and among soil organic matter fractions was also studied. There was a trend for a13c values to become slightly less negative with depth . The a13c value of humic acid was most similar to the o13c value of the dominant vegetation. Fulvic acid was isotopically heavier than humic acid in all analyses. The implications of these trends are discussed.

Between March, 1969 and July, 1970, 16 badgers (Taxidea taxus) were caught and fitted with radio transmitters in the southern part of Curlew Valley. The animals were followed telemetrically; seven animals contributed sufficient data from which home-range, movement and activity patterns could be discerned. The aver age annual home-range size of five females was 664 acres (± s.d. 99 .5 acres). Female home-range sizes were approximately the same within crested wheat-grass (Agropyron aristatum) and sagebrush (Artemisia tridentata) vegetation types, with greater distances traveled each night by females in the crested wheat-grass. Both home-range size and total movement were less during winter in the one badger observed during more than one season. The average home range of two males followed between September and mid-December was 1,440 acreas, twice the average female home-range size. The necropsy of badgers in the study collection and a scat collection yielded information on the food habits and breeding biology of badgers in Curlew Valley. Many prey species were used, but mice were the most frequently eaten food item. Badgers bred between mid-July and the end of August. Delayed implantation PE sisted until approximately January 26. Pregnant females gave birth to an average of 2.2 young about April 1.

The interrelations of black- tailed jackrabbits and the desertshrub vegetation on which they were feeding were studied in Curlew Valley, Northern Utah. The vegetation was described as a threecornered continuum, the corners being types dominated respectively by Artemisia tridentata, Atriplex ~ onfertifolia, and Sarcobatus vermiculatus. Jackrabbit diet was studied by microscopic analysis of plant fragments in stomachs from shot animals. The method was inaccurate, apparently because the ratio of identifiable tissues to all ingested tissues was very low, and varied between plant taxa, and seasonally. This problem seems intractable for desert shrub vegetation. The diet was similar to that reported by other workers on this species, with perennial grasses and forbs most important in sprlng and summer, shrubs in autumn and win ter. Features new to this vegetat ion were large percentages of Halogeton glomeratus, particularly in autumn and winter, and intense selection for Kochia americana. Attempts to explain the foods chosen ln terms of t heir nutrient contents were partically successful. Diet selection by large generalist herbivores was conceptualized as optimization of nutrient intake, mediated by long-delay learning, and constrained by food availability only at very low levels of availaoility. Spatial variation in jackrabbit diets confirmed this "cut-offll response to ava i 1 all i 1 i ty . Percentage utilization was estimated indirectly as jackrabbit density, times yearly food consumption per jackrabbit, times yearround percentage of each taxon in the diet, div i ded by available biomass of each taxon. Less abundant plants were more intensely used, which is expected if consumption does not vary continuously with availability. Perennial grasses, Kochia americana and possibly Grayia spinosa seemed to be under damaging pressure at high jackrabbit densities. Kochia had almost disappeared from outside a sheep- and jackrabbitproof exclosure since the 1950 1 s. In other exc1osures, the presence or absence of jackrabbits seemed to make no difference to the rate of vegetation recovery over 5-7 years after exclusion of sheep. Jackrabbit use of a crested wheatgrass seeding was concentrated ln a 300 m band around its edge.

Basalt outcrops in Curlew Valley consist of vii several flow remnants and eruptive centers situated along the valley margins. Basalt is also found in association with salic rocks that erupted in the central portion of The basalts are of probable Pliocene age and the valley. were erupted during a period of active extensional tectonics. Since their emplacement, and downfaulted. the basalts have been extensively eroded The present morphology reflects the erosional and depositional processes of prehistoric Lake Bonneville. Twelve samples were analyzed chemically for major oxides, trace elements, and rare-earth elements. The basalts form a hypersthene normative series ranging from olivine tholeiite to tholeiite. Based on chemical data, the basalts form three distinct groups. Comparison of the major oxides shows two of the groups forming a differentiation sequence separate from the third group. Trace element and rare-earth element data indicate that the three chemical groups are related to a common source but that two batches of magma probably emanated from this source. Pyrolite, spinel lherzolite, and garnet lherzolite were evaluated as hypothetical mantle materials from which a parent magma might be derived through partial melting. Based on rare-earth element profiles, garnet lherzolite appears to be the most likely source material for deriving the basalts. The trace element and rare-earth element data do not show any anomalies that would suggest contamination from crustal material. Comparison of chemical data shows that the Curlew Valley basalts are genetically similar to basalts from the Kelton and Rozel Point-Black Mountain areas, southwest and southeast of the study area, respectively. The Curlew Valley basalts are chemically similar to olivine tholeiites from the Snake River Plain and Blackfoot Reservoir areas in Idaho, but they do not show much similarity to basalts near Snowville, Utah, northeast of the study area. The Curlew Valley basalts are generally more iron rich and less alkalic than other basaltic rocks from the eastern Basin and Range Province.

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Range improvements for livestock were analyzed for the Curlew National Grassland and Saddle Creek Allotment. Increases in aum's which were a result of range improvements were calculated and valued at $4.00 each. Internal rates of return for both study areas were computed with a 15-year project life span for estimated grazing capacity and permitted grazing. Rate of return was then set at 10 percent to calculate project life span for both permitted grazing and estimated grazing capacity for the two areas. Effects of range improvements for livestock on wildlife habitat were studied. Due to a lack of quantitative data, values could not be placed upon benefits and detriments which improvements for livestock had on wildlife. Positive and negative effects which range improvements for livestock had on wildlife are given for both study areas. An alternative grazing policy is suggested for both study areas. Internal rates of return for permitted grazing are calculated for a 15-year project life span. Internal rate of return was then set at 10 percent to calculate project life span necessary to provide that rate of return.