Disturbances impact abiotic and biotic components within forested ecosystems. This dissertation identifies the impacts of recreational crossing disturbances on water quality, uses dendrochronological techniques to estimate sediment deposition and identify impacts hydrology and climate on radial growth in riparian forested wetlands, and quantifies influences of biotic and abiotic factors on the occurrence of frost-induced cambial damage. In southwestern Virginia, modeled soil erosion rates for multiple-use (hiking, mountain biking, and horseback riding) recreational trail approaches to stream crossings were found to be 13 times greater than rates for undisturbed forests. Downstream changes in macroinvertebrate-based indices indicated water quality was negatively affected downstream from culvert and ford crossings. These findings illustrate recreational stream crossings have the potential to deliver sediment into adjacent streams, particularly where best management practices are not being rigorously implemented, and as a result can negatively impact water quality below stream crossings. Impacts of hydrologic regime were apparent on sediment deposition and on green ash (Fraxinus pennsylvanica Marsh.) and water tupelo (Nyssa aquatica L.) radial growth along the Tensaw River in southwest Alabama. Annual dendrogeomorphic sediment accretion rates were significantly greater for a recent time period (25 years) when compared to longer term rates (131 years) along a natural levee and backswamp. Radial growth in green ash along the natural levee and backswamp was found to be significantly correlated with days flooded and average daily stage level during April while water tupelo further in the backswamp appeared resistant to hydrologic and climatic fluctuations. Results illustrate the importance of riparian wetlands in trapping sediment from adjacent waterways and highlight the role hydrologic regime plays in bottomland succession and productivity. Across northern lower Michigan, late spring frost-induced cambial damage in jack pine (Pinus banksiana Lamb.) occurred more frequently in younger trees and in trees with smaller diameters. Biotic and abiotic factors were found to influence the occurrence of earlywood frost rings. Frequent occurrences of frost-rings can be used to identify frost-prone environments and geographical boundaries for plant species. This dissertation bridged gaps in knowledge of recreational, hydrogeomorphic, and climatic disturbances in forested ecosystems which can be used to develop management strategies. / Ph. D.

Ecosystems in the modern world face a vast array of disturbances, from globally shifting abiotic conditions, to increasingly variable extreme natural events, to high intensity discrete human-caused disturbances. Well-developed, applicable theoretical frameworks on how ecosystems can respond to and withstand these disturbances are needed for adequate management of valued ecological systems. To date, the most promising theoretical development for understanding ecological response to complex sets of disturbances is resilience. Ecological resilience acknowledges non-linear ecosystem behavior, incorporates the role of slowly changing environmental parameters in ecological dynamics, and offers one of the few potential methods to predict, and avoid, impending ecological collapse. However, as ecological resilience has evolved conceptually to include social, political, and economic fields, it has become increasingly difficult to clearly define in, and apply to, managed ecosystems. This dissertation pairs ecological resilience with other, well-established attributes of ecological response to disturbance, namely resistance, persistence, and recovery. By doing so, we can clearly define and quantify each attribute in a range of ecosystem types and over a variety of ecological scales. In Chapter 1, we use microcosm communities to test the relationship between one potential mechanism, landscape connectivity, and multiple attributes of ecological response to disturbance including resistance, resilience, and recovery. We find that each attribute responds uniquely to connectivity, and that generalizing the role of connectivity over all three may give an inaccurate prediction of how ecosystems may respond to individual disturbances. In Chapter 2, we experimentally investigate the presence of early warning indicators of approaching critical thresholds. Using water table drawdown treatments in bog, we test for critical slowing and increased autocorrelation as the bog approaches a transition to forest. We find that critical slowing is clear in composition and moss cover, but that autocorrelation is not apparent. The decoupling of critical slowing and increased autocorrelation could be due to a number of complex ecosystem dynamics, all of which are common in ecosystem management globally. Thus, early warning indicators likely need further development if they are to become applicable. In Chapter 3, we observationally study how conservation management actions may increase or decrease ecological resilience. In particular, we explore how invasive species management intensity correlates with changes in functional redundancy, response diversity, and spatial occurrence of regime shifts in Garry oak meadows. We find that more intense management correlates with less area lost to woody encroachment and increases in functional redundancy through time. However, the relationship was strongly mediated by individual landscape settings. Finally, in Chapter 4, we scale up to a provincial study, investigating persistence of ecosystems and large mammal species in the face of the continuous pressures of land use change. In the results from all four chapters, it is clear that individual attributes of ecological response to disturbance, i.e. resistance, persistence, resilience, or recovery, all play unique roles in ecosystem dynamics. Additionally, the metric chosen to quantify each attribute can play a pivotal role in how we interpret observed dynamics. The work in this dissertation highlights that we cannot understand or predict ecological response to disturbance without clear, measurable concepts. Around a single state of interest, resilience is only one among a suite of attributes that are important to understand. Its additional strength, of potentially predicting the occurrence of ecological thresholds, is still being developed as we explore methods of quantification and application in individual ecosystems. / Graduate

Hunting is often summarised to the act of searching for and killing an animal. Due to this fact, hunting has in recent years, come under scrutiny for its’ morality and lack of environmental consciousness. The thesis will be countering this idea, by demonstrating the entanglement of hunting in specific social and material circumstances what is referred to here as the assemblage of hunting. Following this thread, it will highlight the relationship between hunting, conservation, rewilding, rurality. To illustrate this point, the thesis will be analysing the case of hunting in Alsace. With the help of five interviewees, including three Alsatian hunters, and two members of a local conservation organisation, Alsace Nature, I discuss how the local Alsatian hunting system is experienced, perceived and produced, within a local and more global context. By using theories of Actor-Network-Theory, Assemblage Theory, Landscape Theory, Lefebvre’s production of space, and Disturbance Ecology, I analyse how hunting in Alsace has been shaped by the industrialisation of agriculture in the 20th century, how the activity connects to tradition and the creation of a wider Alsatian identity as Germanic, how hunting, conservation, and agricultural policy intertwine to produce a set of practices – or territorialisations but also how they continuously challenge each other. In addition, I trace the prey animals as deterritorialising agents. Overall results show that hunting in Alsace, while not static, has seen little change when it comes to the local hunting system. This creates conflict, as the local Alsatian landscape has changed heavily. Debates on hunting, such as the return of the lynx, or rural-urban divides can be tied to this perceived divide between hunting and the local landscape. Despite these conflicts, hunting stays an important activity in the Alsatian landscape, and a valuable way of managing the environment according to the interviewees.

The Cerulean Warbler (Setophaga cerulea) is a mature forest obligate and one of the fastest declining songbird species in the United States. This decline may be related to a lack of disturbance within contemporary forests; however, the consequences of disturbance on the species have not been rigorously evaluated. Thus, we assessed multiple responses by Cerulean Warblers to a range of experimental forest disturbances across the core of their breeding range in the Appalachian Mountains. We quantified individual and population responses to these manipulations, and assessed the potential consequences of disturbance on the sexual signaling system. Male ceruleans were strongly attracted to intermediate and heavy disturbances at the stand scale. Despite attraction to disturbed habitats, nest success declined in these conditions, particularly in the highly productive Cumberland Mountains of northern Tennessee. Taken together, these opposing responses suggest that anthropogenically-disturbed forests may act as local ecological traps, but the impact of these local traps on the global population is dependent on several unestimated parameters. At a finer scale, selection for habitat features varied spatially. Males consistently selected for territories near canopy gaps and on productive slopes, but they displayed inconsistent territory selection in regards to tree diameter, basal area, overstory canopy cover, and canopy height. Females were more consistent in their selection of features within territories, selecting nest patches with large, well-spaced trees near disturbances. Floristically, female ceruleans consistently selected for sugar maples (Acer saccharum), white oaks (Quercus alba), and cucumber magnolias (Magnolia acuminata) as nest trees and they selected against red maples (A. rubrum) and red oaks (Q. rubra). Disturbances had little effect on male age structure, but males that occupied disturbed forest habitat were in better condition than those in undisturbed habitat. Parental behavior differed among disturbances, with birds in more highly disturbed habitats provisioning their young at greater rates, but bringing smaller food loads, potentially helping to explain the decrease in nest survival in disturbances. Finally, we found that male ceruleans displayed various plumage ornaments that signaled individual quality. However, the relationship between breast band width and body mass was contingent on habitat, and only existed in intermediate disturbances.

The savannas of southern Africa are a highly variable and globally-important biome supporting rapidly-expanding human populations, along with one of the greatest concentrations of wildlife on the continent. Savannas occupy a fifth of the earth's land surface, yet despite their ecological and economic significance, understanding of the complex couplings and feedbacks that drive spatiotemporal patterns of change are lacking. In Chapter 1 of my dissertation, I discuss some of the different theoretical frameworks used to understand complex and dynamic changes in savanna structure and composition. In Chapter 2, I evaluate spatial drivers of water quality declines in the Chobe River using spatiotemporal and geostatistical modeling of time series data collected along a transect spanning a mosaic of protected, urban, and developing urban land use. Chapter 3 explores the complex couplings and feedbacks that drive spatiotemporal patterns of land cover (LC) change across the Chobe District, with a particular focus on climate, fire, herbivory, and anthropogenic disturbance. In Chapter 4, I evaluated the utility of Distance sampling methods to: 1) derive seasonal fecal loading estimates in national park and unprotected land; 2) provide a simple, standardized method to estimate riparian fecal loading for use in distributed hydrological water quality models; 3) answer questions about complex drivers and patterns of water quality variability in a semi-arid southern African river system. Together, these findings have important implications to land use planning and water conservation in southern Africa's dryland savanna ecosystems. / Ph. D.

<p>Three prescribed burn sites and seven selective timber harvest sites were surveyed using a UAS equipped with a PPK-triggered RGB sensor to determine optimal image collection parameters surrounding each type of disturbance and land cover. The image coordinates were corrected with a third-party base station network (CORS) after the flight, and photogrammetrically processed to produce high-resolution georeferenced orthomosaics. This addressed the first objective of this study, which was to <i>establish effective data procurement methods from both before and after planned </i>disturbances. <br></p><p>Orthomosaic datasets surrounding both a prescribed burn and a selective timber harvest, were used to classify land covers through geographic image-based analysis (GEOBIA). The orthomosaic datasets were segmented into image objects, before classification with a machine-learning algorithm. Land covers for the prescribed prairie burn were 1) bare ground, 2) litter, 3) green vegetation, and 4) burned vegetation. Land covers for the selective timber harvest were 1) mature canopy, 2) understory vegetation, and 3) bare ground. 65 samples per class were collected for prairie burn datasets, and 80 samples per class were collected for timber harvest datasets to train the classifier. A supported vector machines (SVM) algorithm was used to produce four land cover classifications for each site surrounding their respective planned disturbance. Pixel counts for each class were multiplied by the ground sampled distance (GSD) to obtain area calculations for land covers. Accuracy assessments were conducted by projecting 250 equalized stratified random (ESR) reference points onto the georeferenced orthomosaic datasets to compare the classification to the imagery through visual interpretation. This addressed the second objective of this study, which was to <i>establish effective data classification methods from both before and after planned </i>disturbances.<br></p><p>Finally, a two-tailed t-Test was conducted with the overall accuracies for each disturbance type and land cover. Results showed no significant difference in the overall accuracy between land covers. This was done to address the third objective of this study which was to <i>determine if a significant difference exists between the classification accuracies between planned disturbance types</i>. Overall, effective data procurement and classification parameters were established for both <i>before </i>and <i>after </i>two common types of <i>planned </i>disturbances within the CHF region, with slightly better results for prescribed burns than for selective timber harvests.<br></p>