In a celebrated series of experimental observations, starling flocks have been shown to be characterized by scale-free, long-ranged spatial correlations in their velocity fluctuations. While this is expected for velocity orientation correlations on the basis of simple symmetry-breaking arguments, the same scaling-free behaviour for speed (i.e. the absolute value of birds’ velocity) correlations cannot be explained by the same symmetry-based argument. Possible explanations so far put forward required the implicit or explicit fine-tuning of a speed control parameter. In this work we explore a different possibility, investigating the effects of the experimental discrete temporal sampling of individual bird trajectories. We argue that observed velocity may well be a time coarse-grained observable, that is, the sum over many faster course corrections taken by the bird. A simple argument shows such a time coarse-grained speed to be linked with the squared fluctuations of (soft modes) transversal velocities, which may thus acquire a long-range correlation. Our idea is numerically tested by measuring spatial correlations between coarse-grained speeds in the on-lattice equilibrium XY model and the off-lattice out-of-equilibrium Vicsek model in two dimensions. Saturation of the speed correlation length is found in the equilibrium XY model, while in the non-equilibrium Vicsek model ordered symmetry-broken phase shows scale-free behaviour with a correlation length ξ is found to be proportional to system size L. We conclude that in non-equilibrium flocking models, the temporal coarse-graining procedure is able to reproduce scale-free behaviour at system sizes which are relevant to the experimental observations. We believe that this mechanism might find applications beyond the case of starling flocks and perhaps be relevant for other experimental observations of collective motion.

The study of the process of speciation is instrumental to understanding the species diversity observed today. Diverging populations are intriguing, because speciation has not reached an endpoint, yet the process that may eventually lead to distinct species can be studied. Systems that contain many putative species and/or parallel divergences, such as many species flocks and species pairs, are extraordinary examples of divergence and therefore are critical to the understanding of the speciation process. A "miniature" species flock of pupfish (Cyprinodon variegatus) discovered in lakes on San Salvador Island, Bahamas has evolved in less than 6 000 years, and is, therefore, important to the study of the pace of evolutionary processes. The San Salvador Island pupfish flock is composed of a normal form, which resembles coastal C. variegatus, and bulldog and bozo morphs, which diverge ecologically and morphologically from the normal morph. In Chapter 1, I sequenced the mtDNA control region and used haplotype frequency analyses to assess the level of differentiation between sympatric normals and bulldogs sampled from Osprey Lake and Little Lake on San Salvador Island. The bozo morph was too rare to include in the study. I also included samples of normals that occur in lakes without bulldog and bozo morphs to assess any differences between lakes on the island. All haplotype frequency comparisons for sympatric normals and bulldogs were highly significant, which suggests these morphs are distinct populations in sympatry and, therefore, have characteristics of biological species. Further, an estimation of Time for Speciation supports geological data that suggest this fauna is very young (6 000 years). The San Salvador Island pupfish species flock is, therefore, the youngest known species flock and presents an important model system for the study of how morphological and ecological divergence can promote speciation in Cyprinodon. In Chapter 2, I first compared the San Salvador Island pupfishes to other Bahamian C. variegatus populations to assess the level of inter- and intra-island pupfish population differentiation in the Bahamas. The mtDNA control region was sequenced for bulldogs and normals from San Salvador Island and normals sampled from New Providence and Exuma Islands. San Salvador Island bulldogs were found to be distinct from all normal populations sampled, and comparisons of shared haplotypes suggest they originated on San Salvador Island rather than any of the other islands sampled. This was intriguing, because a "bulldog-like" morph has recently been observed in a lake on New Providence Island, which suggests parallel divergences may be occurring throughout the Bahamas. I also sequenced the mtDNA cytochrome b gene to assess the phylogeography of C. variegatus. Populations were sampled from the Bahamas and the east coast of North America, and the results suggest the Bahamas were only recently colonized by the Southern coastal lineage of C. variegatus. A distinct Northern lineage of C. variegatus, which may warrant species designation, was also supported by the cytochrome b data. Overall, the results supported a San Salvador Island origin for the Little Lake and Osprey Lake bulldog morphs, and also suggest the Bahamian C. variegatus populations are very young. / Master of Science

Os bandos mistos de aves são agregações sociais complexas estáveis durante o tempo e espaço. Até hoje, a estrutura social dessas espécies foi descrita a partir de estudos subjetivos de campo ou a partir de compilações do número e intensidade das interações a nível de todo o grupo, sem considerar as interações par-a-par individualmente. Nossos objetivos foram buscar evidências de associações não-randômicas entre pares de espécies de aves, delimitar os grupos a partir das espécies com as associações mais fortes e verificar se há diferenças na estrutura social entre os habitat de floresta primária e secundária. Utilizamos dados de ocorrência das espécies coletados a partir de redes de neblina e gravação de vocalizações para identificar pares de espécies que foram co-detectadas mais frequentemente do que o esperado a partir do modelo nulo e compararamos a força dessa interação entre as florestas tropicais primária e secundária Amazônicas. Nós também utilizamos as associações par-a-par para construir as redes de interação social e suas mudanças entre os tipos de habitat. Nós encontramos muitas interações positivas fortes entre as espécies, mas nenhuma evidência de repulsão. As análises das redes de interação revelaram vários grupos de espécies que corroboram com grupos ecológios descritos na literatura. Além disso, tanto a estrutura da rede de interação como a força da interação se alteraram drasticamente com a perturbação do habitat, com formação de algumas associações novas, mas uma tendência geral para quebra de associações entre as espécies. Nossos resultados mostram que as interações sociais entre essas aves podem ser fortemente afetados pela degradação do habitat, sugerindo que a estabilidade das interações desenvolvida entre espécies é ameaçada pelos distúrbios causados pelo homem. / Inter-specific associations of birds are complex social phenomena, frequently detected and often stable over time and space. So far, the social structure of these associations has been largely deduced from subjective assessments in the field or by counting the number of inter-specific encounters at the whole-group level, without considering changes to individual pairwise interactions. Here, we look for evidence of non-random association between pairs of bird species, delimit groups of more strongly associated species and examine differences in social structure between old growth and secondary forest habitat. We used records of bird species detection from mist-netting capture and from acoustic recordings to identify pairwise associations that were detected more frequently than expected under a null distribution, and compared the strength of these associations between old-growth and secondary forest Amazonian tropical forest. We also used the pairwise strength associations to visualize the social network structure and its changes between habitat types. We found many strongly positive interactions between species, but no evidence of repulsion. Network analyses revealed several modules of species that broadly agree with the subjective groupings described in the ornithological literature. Furthermore, both network structure and association strength changed drastically with habitat disturbance, with the formation of a few new associations but a general trend towards the breaking of associations between species. Our results show that social grouping in birds is real and may be strongly affected by habitat degradation, suggesting that the stability of the associations is threatened by anthropogenic disturbance.

Os bandos mistos de aves são agregações sociais complexas estáveis durante o tempo e espaço. Até hoje, a estrutura social dessas espécies foi descrita a partir de estudos subjetivos de campo ou a partir de compilações do número e intensidade das interações a nível de todo o grupo, sem considerar as interações par-a-par individualmente. Nossos objetivos foram buscar evidências de associações não-randômicas entre pares de espécies de aves, delimitar os grupos a partir das espécies com as associações mais fortes e verificar se há diferenças na estrutura social entre os habitat de floresta primária e secundária. Utilizamos dados de ocorrência das espécies coletados a partir de redes de neblina e gravação de vocalizações para identificar pares de espécies que foram co-detectadas mais frequentemente do que o esperado a partir do modelo nulo e compararamos a força dessa interação entre as florestas tropicais primária e secundária Amazônicas. Nós também utilizamos as associações par-a-par para construir as redes de interação social e suas mudanças entre os tipos de habitat. Nós encontramos muitas interações positivas fortes entre as espécies, mas nenhuma evidência de repulsão. As análises das redes de interação revelaram vários grupos de espécies que corroboram com grupos ecológios descritos na literatura. Além disso, tanto a estrutura da rede de interação como a força da interação se alteraram drasticamente com a perturbação do habitat, com formação de algumas associações novas, mas uma tendência geral para quebra de associações entre as espécies. Nossos resultados mostram que as interações sociais entre essas aves podem ser fortemente afetados pela degradação do habitat, sugerindo que a estabilidade das interações desenvolvida entre espécies é ameaçada pelos distúrbios causados pelo homem. / Inter-specific associations of birds are complex social phenomena, frequently detected and often stable over time and space. So far, the social structure of these associations has been largely deduced from subjective assessments in the field or by counting the number of inter-specific encounters at the whole-group level, without considering changes to individual pairwise interactions. Here, we look for evidence of non-random association between pairs of bird species, delimit groups of more strongly associated species and examine differences in social structure between old growth and secondary forest habitat. We used records of bird species detection from mist-netting capture and from acoustic recordings to identify pairwise associations that were detected more frequently than expected under a null distribution, and compared the strength of these associations between old-growth and secondary forest Amazonian tropical forest. We also used the pairwise strength associations to visualize the social network structure and its changes between habitat types. We found many strongly positive interactions between species, but no evidence of repulsion. Network analyses revealed several modules of species that broadly agree with the subjective groupings described in the ornithological literature. Furthermore, both network structure and association strength changed drastically with habitat disturbance, with the formation of a few new associations but a general trend towards the breaking of associations between species. Our results show that social grouping in birds is real and may be strongly affected by habitat degradation, suggesting that the stability of the associations is threatened by anthropogenic disturbance.

Os bandos mistos de aves são agregações sociais complexas estáveis durante o tempo e espaço. Até hoje, a estrutura social dessas espécies foi descrita a partir de estudos subjetivos de campo ou a partir de compilações do número e intensidade das interações a nível de todo o grupo, sem considerar as interações par-a-par individualmente. Nossos objetivos foram buscar evidências de associações não-randômicas entre pares de espécies de aves, delimitar os grupos a partir das espécies com as associações mais fortes e verificar se há diferenças na estrutura social entre os habitat de floresta primária e secundária. Utilizamos dados de ocorrência das espécies coletados a partir de redes de neblina e gravação de vocalizações para identificar pares de espécies que foram co-detectadas mais frequentemente do que o esperado a partir do modelo nulo e compararamos a força dessa interação entre as florestas tropicais primária e secundária Amazônicas. Nós também utilizamos as associações par-a-par para construir as redes de interação social e suas mudanças entre os tipos de habitat. Nós encontramos muitas interações positivas fortes entre as espécies, mas nenhuma evidência de repulsão. As análises das redes de interação revelaram vários grupos de espécies que corroboram com grupos ecológios descritos na literatura. Além disso, tanto a estrutura da rede de interação como a força da interação se alteraram drasticamente com a perturbação do habitat, com formação de algumas associações novas, mas uma tendência geral para quebra de associações entre as espécies. Nossos resultados mostram que as interações sociais entre essas aves podem ser fortemente afetados pela degradação do habitat, sugerindo que a estabilidade das interações desenvolvida entre espécies é ameaçada pelos distúrbios causados pelo homem. / Inter-specific associations of birds are complex social phenomena, frequently detected and often stable over time and space. So far, the social structure of these associations has been largely deduced from subjective assessments in the field or by counting the number of inter-specific encounters at the whole-group level, without considering changes to individual pairwise interactions. Here, we look for evidence of non-random association between pairs of bird species, delimit groups of more strongly associated species and examine differences in social structure between old growth and secondary forest habitat. We used records of bird species detection from mist-netting capture and from acoustic recordings to identify pairwise associations that were detected more frequently than expected under a null distribution, and compared the strength of these associations between old-growth and secondary forest Amazonian tropical forest. We also used the pairwise strength associations to visualize the social network structure and its changes between habitat types. We found many strongly positive interactions between species, but no evidence of repulsion. Network analyses revealed several modules of species that broadly agree with the subjective groupings described in the ornithological literature. Furthermore, both network structure and association strength changed drastically with habitat disturbance, with the formation of a few new associations but a general trend towards the breaking of associations between species. Our results show that social grouping in birds is real and may be strongly affected by habitat degradation, suggesting that the stability of the associations is threatened by anthropogenic disturbance.

This work is devoted to the numerical solution of equations describing the dynamics of flocks of birds. Specifically, we pay attention to the Euler equations for compressible flow with a right-hand side correction. This model is based on the work Fornasier et al. (2010). Due to the complexity of the model, we focus only on the one-dimensional case. For the numerical solution we use a semi-implicit discontinuous Galerkin method. Discretization of the right-hand side is chosen so that we preserve the structure of the semi-implicit scheme for the Euler equations presented in the work Feistauer, Kučera (2007). The proposed numerical scheme was implemented and numerical experiments showing the robustness of the scheme were carried out. Powered by TCPDF (www.tcpdf.org)

Made available in DSpace on 2015-03-05T16:20:40Z (GMT). No. of bitstreams: 0 Previous issue date: 27 / Universidade do Vale do Rio dos Sinos,Coordenação de Aperfeiçoamento de Pessoal de Nível Superior / O presente trabalho teve como objetivos caracterizar a estrutura, composição e dinâmica da comunidade de aves em um fragmento de Mata Atlântica localizado no extremo norte do estado do Rio Grande do Sul, bem como realizar análises comparativas entre interior e borda de floresta em relação a (1) composição de espécies, (2) riqueza, (3) abundância de grupos funcionais de aves e (4) abundância de espécies endêmicas de Mata Atlântica. Foram definidos 22 grupos funcionais de aves para descrição geral do fragmento de floresta estudado e comparação entre interior e borda de floresta, entretanto apenas 11 puderam ser comparados a nível estatístico, os demais grupos não obtiveram suficiência amostral para serem incluídos nas análises. Os grupos funcionais utilizados na comparação de abundância entre borda e interior de floresta foram: granívoros/insetívoros generalistas, granívoros/frugívoros de solo, insetívoros de sub-bosque, insetívoros de sub-bosque e estrato médio, insetívoros escaladores de troncos e galhos, ins / The present goal was to characterize structure, composition and dynamics of bird community at an Atlantic Forest remnant at the extreme north of Rio Grande do Sul State as well to compare interior and forest edge related to (1) species composition, (2) richness, (3) abundance of functional groups and (4) Atlantic Forest abundance of endemics bird species. General description of the fragment and comparisons between forest interior and edge were based on 22 defined functional groups, however only 11 could be compared to statistical level, the other groups did not obtain sufficient sample data to be included in the analysis. Functional groups used in the comparison between the abundance of forest edge and interior were: generalist seedeater/insectivores, ground seedeater/frugivores, understory insectivores, mid and understory insectivores, tunk and twig insectivores, bamboo or forest tangle insectivores, canopy insectivores, mid and understory insectivore/frugivores, canopy insectivore/frugivores, nectarivore/in

Orientador: Marcos César de Oliveira Santos / Banca: Marco Aurelio Pizo Ferreira / Banca: Pedro Ferreira Develey / Resumo: A maria-da-restinga (Phylloscartes kronei) é uma espécie da família Tyrannidae descrita recentemente e considerada como globalmente ameaçada de extinção, cuja história natural é pouco conhecida. Visando preencher parte desta lacuna, o objetivo deste estudo foi de caracterizar o comportamento de forrageamento da maria-da-restinga, descrever os bandos mistos aos quais ela se associa, bem como descrever alguns outros aspectos de sua história natural como seus hábitos alimentares, o comportamento de dormir, insights sobre a biologia reprodutiva, o alcance de vôo, encontros agonísticos interespecíficos e a muda. No período de agosto de 2008 a julho de 2009 foram realizadas excursões mensais ao município de Ilha Comprida (~24º S), Estado de São Paulo. Aliado ao esforço anual mencionado, uma visita ao município de Guaraqueçaba (25º18'S e 48º19'W), Estado do Paraná, foi conduzida entre os dias 25 de julho e 1º de agosto de 2008. Foram coletadas 613 amostras de forrageamento, através de observações diretas com auxílio de binóculos com aumento de 10 vezes. Durante os períodos de coleta de dados sobre o forrageamento da espécie, para cada encontro com a mesma foi anotado se esta se encontrava só, aos pares, em grupos ou associada a bandos mistos de aves. Quando em bandos mistos, foram anotadas todas as espécies ocorrentes, visando caracterizar os bandos aos quais se associa. Para a descrição do hábito alimentar da espécie foram analisados os conteúdos estomacais de cinco exemplares coletados na década de 1990 no Estado do Paraná, tendo sido identificados 116 ítens alimentares. Através da análise dos dados obtidos, conclui-se que P. kronei captura suas presas principalmente em folhas verdes, entre 0,5 e 15m acima do solo, utilizando a manobra "investir-atingir". Alimenta-se predominantemente de artrópodes, sendo que os insetos das ordens Coleoptera... (Resumo completo, clicar acesso eletrônico abaixo) / Abstract: The Restinga Tyrannulet (Phylloscartes kronei) is a recently described and globally threatened species of the family Tyrannidae, whose natural history is poorly known. In this study several information on the natural history of P. kronei were gathered such as several aspects on foraging behavior, association with mixed species bird flocks, feeding habits, the description of the roosting behavior, insights on the reproductive biology, the flying range, interespecific agonistic encounters and moulting. From August 2008 to July 2009, data on this species were collected on a monthly basis in Ilha Comprida (~24º S), São Paulo state, Brazil. In addition, a visit to Guaraqueçaba (25º18'S e 48º19'W), Paraná state, was conducted between 25 July and 01 August 2008. A total of 613 foraging maneuvers were gathered by direct observations with the aid of binoculars with 10x magnification. In all meetings with the species it was reported if individuals were alone, in pairs, in groups or associated with mixed species bird flocks. When mixed species flocks with Restinga Tyrannulets were observed, all species in the flock were identified. Five stomach contents collected in the 1990s in Paraná state were evaluated and 116 food items were identified. The Restinga Tyrannulet captures preys mainly with sally-strikes in green leaves between 0.5 and 15m above the ground. This species feeds mainly on arthropods, especially Coleoptera and Hymenoptera. The observed foraging behaviors and feeding habits were similar to the ones observed in other species of Phylloscartes, but new food items are described for this genus: Anobiidae (Coleoptera), Coccinelidae (Coleoptera), Chalcidoidea (Hymenoptera), Microhymenoptera and two species of fruits (Clusia criuva and Ternstroemia brasiliensis). P. kronei is a sporadic follower of mixed species bird flocks, which had the inclusion of 25 species on their compositions... (Complete abstract click electronic access below) / Mestre

This thesis was conducted to evaluate important epidemiological features of Gastro-Intestinal Nematode (GIN) infections in Ontario sheep flocks; namely, the PeriParturient Egg Rise (PPER), overwintering of GIN free-living stages on pasture, and Anthelmintic Resistance (AR). Three main studies were carried out: a longitudinal study was conducted on six sheep farms to evaluate the PPER in ewes lambing in different seasons and to determine whether total plasma protein (TPP) levels and packed cell volume (PCV) were associated with increased fecal GIN-egg shedding. Secondly, a pilot-study was conducted on three farms to describe pasture-level environmental conditions and over-wintering survival and infectivity of free-living GIN larvae, especially Haemonchus contortus. Lastly, a cross-sectional study was conducted on 47 sheep farms in Ontario, to evaluate the frequency of AR, compare different diagnostic tests for AR, and evaluate management practices associated with AR. In the longitudinal study, the PPER was observed in winter, spring and autumn lambing ewes, though the magnitude and distribution of the PPER varied with season. Lower TPP and PCV values were associated with increased fecal GIN-egg counts. The pilot-study suggested that H. contortus larvae did not overwinter successfully on pasture, while other GINs, such as Teladorsagia sp., Trichostrongylus spp. and Nematodirus spp., were able to overwinter on pasture, and were infective the following spring. Resistance to ivermectin, fenbendazole and levamisole was demonstrated on 97% (28/29), 95% (19/20) and 6% (1/17) respectively of the farms tested; most of the resistance observed was found in Haemonchus sp. The Fecal Egg Count Reduction percentage following treatment was influenced by which mean (i.e. arithmetic vs. geometric) was used in the formula; use of pre-treatment in addition to post-treatment faecal egg counts was not influential. Both the fecal egg count reduction test and the larval development assay diagnosed resistance, but there was poor agreement between the two tests, as indicated by the Kappa test. The prior use of benzimidazoles on farms was associated with higher levels of fenbendazole resistance. The information generated in this thesis will be used to develop a parasite control program for sheep flocks in Ontario and to guide future research on GIN parasitism. / New Directions - Ontario Ministry of Agriculture, Food and Rural Affairs; Ontario Sheep Marketing Agency; Ontario Agriculture Centre Canada; Merial; Animal Health Strategic Initiative - University of Guelph Partnership; Undergraduate Research Assistantship; Gartshore Memorial Sheep Scholarship;

Analisou-se a composição, estrutura e padrão sazonal de formação dos bandos mistos de aves em um remanescente de floresta paludosa na planície costeira do Rio Grande do Sul, sul do Brasil. Efetuaram-se amostragens mensais de fevereiro de 2005 a janeiro de 2006, com observações ocasionais em 2004 e janeiro de 2005. Pontos de contagem foram utilizados para estimar a abundância relativa das espécies integrantes dos bandos, e transecções foram efetuadas para as observações dos bandos mistos. Quarenta e sete espécies foram registradas como participantes em 92 bandos amostrados. O número de indivíduos foi positivamente correlacionado com o número de espécies dentro dos bandos. O pool dos não-passeriformes foi pobremente representado nos bandos, enquanto passeriformes suboscines e oscines foram igualmente representados na riqueza dos bandos. Onívoros e insetívoros foram bem representados, mas os nectarívoros tiveram apenas uma espécie integrante; frugívoros e granívoros foram ausentes nos bandos mistos.Espécies migratórias foram pouco representadas nos bandos, como havia sido verificado para outras áreas de Floresta Atlântica. A regularidade de uma espécie em integrar bandos mistos foi uma função de sua abundância relativa, e as espécies mais conspícuas tenderam a ser os integrantes mais regulares. Detectaram-se quinze co-ocorrências significativas em 153 combinações possíveis (10%), onze delas positivas e quatro negativas. Nenhum tipo estrutural de bandos foi distinguido por meio de análise de agrupamento. Associações positivas e negativas talvez estejam relacionadas a similaridades e dissimilaridades na distribuição vertical de cada par de espécies. Os atributos de freqüência, tamanho e riqueza dos bandos foram os menores encontrados em qualquer outro estudo na Floresta Atlântica. Os bandos formaram-se o ano inteiro no Faxinal, mas a freqüência, o tamanho e a riqueza dos bandos diferiram significativamente entre os meses e entre as estações. Osvalores máximos destes três atributos foram encontrados na estação não-reprodutiva, lembrando os padrões encontrados em outros locais da Floresta Atlântica. A combinação de um pequeno grupo de espécies abundantes e bem distribuídas, junto com uma baixa riqueza local talvez torne a estrutura dos bandos mistos no Faxinal menos variável do que em outros locais da Floresta Atlântica. / The composition, structure and seasonal pattern of formation of mixed-species bird flocks were analyzed in a swamp forest remnant in the coastal plain of Rio Grande do Sul state, southern Brazil. Field work was carried out monthly from February 2005 to January 2006, plus occasional observations in 2004 and January 2005. Point counts estimated the relative abundance of flocking species, and transects were performed to access data relative to the flocks. Forty seven species were recorded as participants in 92 flocks sampled. The number of individuals was positively correlated with the number of species within the flocks. Non-passerines pool is less represented in the flocks, while suboscines and oscines were equally represented in the flock richness. Omnivores and insectivores were well represented, but nectarivores had only one flocking species while frugivores and granivores were absent in flocks. Migrants had low representation on the composition and structure of flocks, as predicted for lowland Atlantic Forest areas. The regularity of a given species in to integrate flocks was a function of its relative abundance, and the species more conspicuous tended to be the more regular flock joiners. Fifteen significant co-occurrences of 153 possible pairs (10%) were detected, eleven positively and four negatively associated. None structural type of flocks was recognized through cluster analyses. Positive and negative associations were better explained by similarities or dissimilarities on vertical distribution of each species pair. The attributes of frequency, flock size and flock richness were the lowest found in any study on Atlantic Forest. Flocks were formed throughout the year at Faxinal, but frequency, flock size and flock richness differed significantly between months and between seasons. Higher values of these three attributes were found in non-breeding season, reaching the pattern found elsewhere in Atlantic Forest. The combination of a little set of abundant and well distributed speciesand low richness perhaps makes the flock structure at Faxinal less variable than that of Atlantic Forest elsewhere.