Chondrichthyans are cartilaginous fishes that include the extant chimaeras, sharks, and batoids, and their extinct relatives. In this dissertation, I examined the growth, development, and relationships of extinct and extant chondrichthyans. I reexamined the skeleton of fossil and fetal specimens of the angel sharks (Squatiniformes) and reanalyzed the current morphologically based hypothesis of chondrichthyan evolution, which conflicts with the current molecular based hypothesis. I did this by including extinct taxa and new characters based on dentition, and the ethmoid, occipital, pectoral, and vertebral skeleton. My results supported the pre-existing morphological hypothesis that angel sharks, saw sharks, and batoids form a clade. However, some of my new characters, particularly those based on the vertebral morphology, indicate that fetal and juvenile angel sharks do not share as many apomorphies with batoids as previously hypothesized from the examination of adults. I also examine the relationships of major groups within batoids, beginning with the construction of a hypothesis of the evolutionary history of electric rays (Torpediniformes). My results are consistent with previous rank-based classifications. However, they deviate from previous classifications depending on the criteria used to generate the hypothesis and on which taxon, or combination of taxa, were used to root the ingroup phylogeny. Because pectoral and vertebral morphology are so critical to morphological hypotheses, I also examined the growth and development of the synarcual cartilage in batoid fishes, with particular emphasis on the synarcual of skates (Rajiformes). My results demonstrate that calcification and chondrification do not proceed in the same order, temporally and spatially. Finally, I redescribe the extinct batoid †Cyclobatis, known only from the Cretaceous, and evaluate its phylogenetic position. My results indicate that †Cyclobatis, the oldest known rajid, is also the sister taxon to a clade of all known extant members of Rajidae. Furthermore, the inclusion of new characters, mainly derived from the synarcual, help to resolve the interrelationships of Rajidae. / text

South Africa is well known for its many, important fossil bearing deposits of Plio-Pleistocene age. Many of these sites contain abundant remains of extinct and extant animals including hominins. Non-herpetological fauna have been used to reconstruct important information about past environments and for chronological data. This study tried a different method; using herpefauna to establish past environments of Cooper’s Cave. Herpetological fauna potentially have numerous advantages in their use for such questions, including typically limited ranging behaviors, and specific temperature requirements. Despite these advantages only a few studies have even mentioned the presence of squamate fauna in the fossil assemblages of these sites. This study has demonstrated that herpefauna exists, in reasonable levels of abundance and with adequate preservation within the fossil record of the dolomitic region now known as the Cradle of Humankind. It has also demonstrated for the first time the presence of a relative abundance of herpefauna at numbers far greater than any previous study has recognized. The present study was however, handicapped in not being able to fully utilize the material at hand to interpret past environments, owing to the lack of comparative material needed to move beyond the family or generic level in most cases with any degree of confidence. However, Agamids and Pseudocordylids were described to generic level. However, the many shortcomings that this work highlighted should not be seen as reason not to pursue the study of herpefauna, but to improve present comparative collections and collecting methods around the fossils themselves.

In dieser Arbeit wurde die Systematik und Evolution der Viviparidae (Mollusca, Gastropoda, Caenogastropoda) mit einem Fokus auf den asiatischen Gattungen untersucht. Es wurde die erste umfassende gattungsübergreifende Phylogenie der Gruppe mit Vertretern von mehr als 70% der bekannten Viviparidengattungen durch die Analyse eines molekularen Datensatzes aus mitochondrialen wie auch nukleären Sequenzen rekonstruiert. Die Einteilung in die aufgrund von anatomischen Merkmalen beschriebenen Unterfamilien konnte durch die genetischen Untersuchungen bestätigt werden. Neben der molekularen Systematik stand auch die Aufklärung der historischen Biogeographie der Viviparidae im Fokus. Mithilfe von Fossilbelegen und der Anwendung einer molekularen Uhr wurde die Diversifikation dieser Familie in Raum und Zeit untersucht. Abschätzungen über das Alter der Abspaltungen der wichtigsten Abstammungslinien innerhalb der Viviparidae mittels der molekularen Uhr lassen sowohl Vikarianz- als auch Dispersal-Ereignisse am Zustandekommen heutiger Verbreitungsmuster erkennen. Die bisherige systematische Gliederung der Viviparidae ist ausschließlich durch einen conchologischen Ansatz geprägt. Bei der Untersuchung zur Evolution der Schale, unter Berücksichtigung der molekularen Ergebnisse, zeigt sich jedoch, dass eine klare Ordnung innerhalb der Vivipariden, bezogen auf die Entstehung von Farbbändern oder Skulpturierungen auf der Schalenoberfläche, nicht zu erkennen ist. Scheinbar komplexe Schalenmerkmale entstanden mehrfach unabhängig in unterschiedlichen Linien. Es konnte hier aber auch gezeigt werden, dass die Untersuchung von anatomischen Merkmalen im Vergleich zur Schale wesentlich besser geeignet ist, natürliche Verwandtschaftsbeziehungen zu erkennen. Die in dieser Arbeit gewonnenen Ergebnisse schaffen die Grundlage für eine in Zukunft nötige Revision der Familie Viviparidae auf Gattungsebene. / This thesis focusses on the systematics and evolution of Viviparidae (Mollusca, Gastropoda, Caenogastropoda), a large and putatively ancient group of ovoviviparous freshwater gastropods with an almost worldwide distribution. With more than 200 described species with highly disparate shells, Asia and Australia have the highest viviparid diversity. Three genetic markers where used to calculate the first molecular phylogeny comprising more than 70% of all extant viviparid genera which provides new insights into the systematics and biogeography of this diverse group of freshwater snails. The traditional division into three subfamilies (Lioplacinae Gill, 1863, Viviparinae Gray, 1847 and Bellamyinae Rohrbach, 1937), based on anatomical data, is supported by the molecular phylogeny. The taxonomy of the Asian taxa is still largely based on shell morphology and rather inconsistent, though, making it difficult to gain deeper insights into their evolution and biogeography at present. The molecular phylogeny presented here indicates that the current shell morphology-based viviparid taxonomy does not reflect the phylogenetic relationships of species and clades. In addition to the focus on molecular systematics, a major part of this thesis is devoted to elucidating the historical biogeography of viviparids. A molecular clock analysis based on a calibration scheme using fossils of varying age and taxonomic affinity was used to yield divergence time estimates for the major viviparid lineages. Ancestral areas were inferred by parsimony and likelihood analyses. Viviparid distribution patterns have been shaped by both dispersal and vicariance events. The study of anatomical features turned out to be considerably better approach for establishing phylogenetic relationships of viviparids, manifested in good support of genera established by molecular data. The results obtained in this work provide the basis for a much needed revision of the Viviparidae at the generic level.

Bibliography: leaves [206]-244.

Bibliography: leaves [206]-244. / xxviii, 244 leaves, [31] leaves of plates : ill. (some col.) ; 30 cm. / Title page, contents and abstract only. The complete thesis in print form is available from the University Library. / Thesis (Ph.D.)--University of Adelaide, Dept. of Geology and Geophysics, 2001

<p>Small shelly fossils are common in sediments of Early Cambrian age and include the earliest common representatives of metazoan animals with mineralized hard parts. The group includes fossils of very different morphology, composition and ultrastructure. They seem to represent skeletal remains of numerous animal groups, the biological affinities of which are largely unresolved. However, the wide geographic range of many forms has the potential to enhance biostratigraphic and palaeogeographic resolution in the Early Cambrian. </p><p>The late Early Cambrian sequence of North-East Greenland has yielded an assemblage of more than 88 species of small shelly fossils, brachiopods and trilobites, indicative of a middle Dyeran age (Botoman equivalent). The recovered fossils include a number of species that are known from other Early Cambrian palaeocontinents, and particularly strong ties to late Early Cambrian faunas of Australia are documented. The many cosmopolitan taxa thus identified suggests a close juxtaposition of palaeocontinents at this time. </p><p>The systematic affinity of many of these small shelly fossils is poorly understood, partly because of their fragmentary nature and poor preservation. However, new data from North-East Greenland improves our understanding of the function and biological affinity of certain taxa. Collections of the problematic fossil <i>Mongolitubulus</i> from North and North-East Greenland exhibit characters indicative of a defensive function as spines of bivalved arthropods, while species of the problematic genus <i>Triplicatella</i> represent the opercula of an unknown tubular shell, probably related to orthothecid hyoliths. The bivalved fossil <i>Mickwitzia</i> from North-East Greenland combines characters of linguliform brachiopods and sclerites of <i>Micrina</i>, a non-bivalved problematic form (halkieriid) from Australia. The combination suggests that <i>Mickwitzia</i> is a stem group brachiopod and strengthens arguments for a halkieriid ancestry of the brachiopod phylum.</p>

Small shelly fossils are common in sediments of Early Cambrian age and include the earliest common representatives of metazoan animals with mineralized hard parts. The group includes fossils of very different morphology, composition and ultrastructure. They seem to represent skeletal remains of numerous animal groups, the biological affinities of which are largely unresolved. However, the wide geographic range of many forms has the potential to enhance biostratigraphic and palaeogeographic resolution in the Early Cambrian. The late Early Cambrian sequence of North-East Greenland has yielded an assemblage of more than 88 species of small shelly fossils, brachiopods and trilobites, indicative of a middle Dyeran age (Botoman equivalent). The recovered fossils include a number of species that are known from other Early Cambrian palaeocontinents, and particularly strong ties to late Early Cambrian faunas of Australia are documented. The many cosmopolitan taxa thus identified suggests a close juxtaposition of palaeocontinents at this time. The systematic affinity of many of these small shelly fossils is poorly understood, partly because of their fragmentary nature and poor preservation. However, new data from North-East Greenland improves our understanding of the function and biological affinity of certain taxa. Collections of the problematic fossil Mongolitubulus from North and North-East Greenland exhibit characters indicative of a defensive function as spines of bivalved arthropods, while species of the problematic genus Triplicatella represent the opercula of an unknown tubular shell, probably related to orthothecid hyoliths. The bivalved fossil Mickwitzia from North-East Greenland combines characters of linguliform brachiopods and sclerites of Micrina, a non-bivalved problematic form (halkieriid) from Australia. The combination suggests that Mickwitzia is a stem group brachiopod and strengthens arguments for a halkieriid ancestry of the brachiopod phylum.

Bibliography: leaves [206]-244. / xxviii, 244 leaves, [31] leaves of plates : ill. (some col.) ; 30 cm. / Title page, contents and abstract only. The complete thesis in print form is available from the University Library. / Thesis (Ph.D.)--University of Adelaide, Dept. of Geology and Geophysics, 2001

Research Doctorate - Doctor of Philosophy (PhD) / The records of environmental change in Australia’s arid zone can be greatly enriched by employing a multi-proxy approach and landscape-scale analysis. This research uses these tools to construct a palaeoenvironmental history of the Paroo/Warrego Region. While the Region’s flow regimes and water balances are characterised by medium-term (decadal) variability (Young, 1999), its hydrological records are inadequately brief. Subsequently, land and water management decisions are based on short term data, risking irreversible damage, desertification or loss of diversity. A better understanding of this highly dynamic landscape can thus improve the land and resource management outcomes. While dating was constrained by a lack of funds, the Paroo/Warrego history reconstructed from fluvial and aeolian deposits correlated well with events recorded from other inland regions of the Australian continent. In summary, this new research provided evidence of high lake water levels prior to the Last Glacial. The extreme aridity at the onset of Last Glacial caused long term drying of the lakes and mobilisation of the red sand dunes. In latter stages of the glacial phase the aridity gave way to periodic fluctuations between flood and drought events that probably lasted until 16 000 - 14 000 BP. The new climatic regime resulted in formation of gypsum lunettes and later, following reduction in gypsum supply, clay lunettes. The orientation of red sand dunes and lunettes indicates a more northerly extent of the westerlies than in modern times. Around the late Pleistocene-early Holocene boundary the climate became more stable and wetter, but still somewhat drier than during the pre-Last Glacial lacustrine phase. As a result, the region’s lakes reverted to a permanent and semi-permanent status. A strong aridity signal, comparable to the semi-regular droughts of the Last Glacial, was recorded in the Paroo/Warrego lakes during the late 1890s-1940s period of below average rainfall. It was followed by 50 years of wetter conditions with two extremely wet phases in the 1950s and the 1970s. Finally, the most recent records suggest a new drying trend. The semi-arid vegetation appears to have adapted to climate variability, with herbs and grasses expanding with the onset of wet conditions before being replaced by Chenopodiaceae as the landscape started to dry. The fresher lake basins and water courses were likely to provide refuge during prolonged arid phases and dispersal foci during intervening wetter periods, thus enabling greater flexibility in response to changes and enhancing resilience. The European land use interfered with the natural cycles and balances, leading to decrease in ground cover, suppression of fire, increase in runoff and catchment erosion, acceleration of sediment accumulation rates in wetlands, resulting in decline of their water holding capacity, and expansion of woody vegetation. The research improved the processing protocols, reference databases, and transfer of methods to enable greater sample processing efficiency and improve results. The use of multiple proxies (including biotic and abiotic components) and sites, as well as different depositional features, provided access to a broader picture of environmental change than was previously possible. It also facilitated multi-scale resolution, allowing discrimination between localised responses of individual lakes and regional trends. The full value of this research will come from informing natural resource managers, whose actions will shape the future landscapes of the Paroo and Warrego Region.

Research Doctorate - Doctor of Philosophy (PhD) / The records of environmental change in Australia’s arid zone can be greatly enriched by employing a multi-proxy approach and landscape-scale analysis. This research uses these tools to construct a palaeoenvironmental history of the Paroo/Warrego Region. While the Region’s flow regimes and water balances are characterised by medium-term (decadal) variability (Young, 1999), its hydrological records are inadequately brief. Subsequently, land and water management decisions are based on short term data, risking irreversible damage, desertification or loss of diversity. A better understanding of this highly dynamic landscape can thus improve the land and resource management outcomes. While dating was constrained by a lack of funds, the Paroo/Warrego history reconstructed from fluvial and aeolian deposits correlated well with events recorded from other inland regions of the Australian continent. In summary, this new research provided evidence of high lake water levels prior to the Last Glacial. The extreme aridity at the onset of Last Glacial caused long term drying of the lakes and mobilisation of the red sand dunes. In latter stages of the glacial phase the aridity gave way to periodic fluctuations between flood and drought events that probably lasted until 16 000 - 14 000 BP. The new climatic regime resulted in formation of gypsum lunettes and later, following reduction in gypsum supply, clay lunettes. The orientation of red sand dunes and lunettes indicates a more northerly extent of the westerlies than in modern times. Around the late Pleistocene-early Holocene boundary the climate became more stable and wetter, but still somewhat drier than during the pre-Last Glacial lacustrine phase. As a result, the region’s lakes reverted to a permanent and semi-permanent status. A strong aridity signal, comparable to the semi-regular droughts of the Last Glacial, was recorded in the Paroo/Warrego lakes during the late 1890s-1940s period of below average rainfall. It was followed by 50 years of wetter conditions with two extremely wet phases in the 1950s and the 1970s. Finally, the most recent records suggest a new drying trend. The semi-arid vegetation appears to have adapted to climate variability, with herbs and grasses expanding with the onset of wet conditions before being replaced by Chenopodiaceae as the landscape started to dry. The fresher lake basins and water courses were likely to provide refuge during prolonged arid phases and dispersal foci during intervening wetter periods, thus enabling greater flexibility in response to changes and enhancing resilience. The European land use interfered with the natural cycles and balances, leading to decrease in ground cover, suppression of fire, increase in runoff and catchment erosion, acceleration of sediment accumulation rates in wetlands, resulting in decline of their water holding capacity, and expansion of woody vegetation. The research improved the processing protocols, reference databases, and transfer of methods to enable greater sample processing efficiency and improve results. The use of multiple proxies (including biotic and abiotic components) and sites, as well as different depositional features, provided access to a broader picture of environmental change than was previously possible. It also facilitated multi-scale resolution, allowing discrimination between localised responses of individual lakes and regional trends. The full value of this research will come from informing natural resource managers, whose actions will shape the future landscapes of the Paroo and Warrego Region.