White rhinos in the Umfolozi Game Reserve (UGR) are managed with a 'Sink Management Policy' which utilises natural dispersal patterns of white rhinos from an area of high density (the core), to areas of low density (vacuum zones). This study was initiated to determine how white rhinos utilised the vacuum zones, as the management staff of the UGR felt that white rhinos were not dispersing into them but utilising them only as areas of trophic resources. Significantly different white rhino densities were established in the Makhamisa and Masinda vacuum zones by removing white rhinos from Masinda. Changes in the density of white rhinos were monitored and the age and sex compositions determined. No significant changes in the density of white rhino groups were recorded during the study, however, the density of individuals in Makhamisa increased significantly in the dry season, while the density in Masinda did not change. The reason for this increase could not be determined, however, the most likely explanation was that rhinos moved toward the Makhamisa study area in search of surface water which was present in the White Umfolozi River. During the dry season the water level in the Umfolozi River was low so the white rhinos where able to cross into the study area. However, owing to the above average rainfall, resources were not limited in either study area, and most likely in most of the reserve. Thus, it could not be determined why white rhinos would need to enter the Makhamisa study area. Changes in grass height, grass colour, and the availability of water in pans and streams were monitored throughout the study. The grassland community compositions of the two study areas were found not to be significantly different. The total utilisation of the different grassland communities by white rhinos during the study were determined for both study areas. White rhinos in the two study areas utilised the grassland communities in a similar pattern throughout the study. During the wet and dry seasons, white rhinos primarily foraged in the short grassland community (Panicum coloratum & Themeda-Urachloa), their staple grassland community, and were not observed to switch and start foraging in the tall grassland community (Themeda & Themeda-Panicum). their reserve community. White rhinos in both study areas were able to forage in their stable grassland community throughout the dry season because of the above average rainfall experienced during the study The results of the study suggest that white rhinos outside the study areas did not utilise the vacuum zones for trophic resources. However, as the study was conducted in a year with above average rainfall, these findings may not represent how white rhinos utilise the vacuum zones in years with average or below average rainfall. / Thesis (M.Sc.)-University of Natal, Pietermaritzburg, 1998.

Although the leopard (Panthera pardus) is one of the more successful large carnivores, challenges for leopard conservation emulate those of other more endangered carnivores, and they are a model species for investigating issues affecting carnivore persistence worldwide. This thesis represents a six year study of leopard on the Karongwe Game Reserve, South Africa, which provided a unique opportunity to observe various aspects of behavioural ecology in the absence of prey availability constraints or human persecution. Small, enclosed reserves such as Karongwe make up 16.8% of the total land in South Africa and undertaking sound ecological research in these areas provides valuable data for evidence-based conservation and management. The leopard is notoriously shy and difficult to study and I used free darting and habituation to enhance visual observation, in order to understand the ecological processes influencing leopard reproductive success and survival. My results show that leopard in the high prey area studied, consume almost double the number of ungulates as leopard in similar habitats elsewhere. This generalist predator improved its hunting success by selecting vulnerable prey and selectively hunted in habitats of intermediate density, where preferred prey were most abundant. Ample nutrition played a key role in reproductive health and reduced the duration of reproductive parameters below that previously recorded in the literature. Any additional nutritional input could not translate into increased population growth as females were already reproducing optimally. Female territorial size and habitat selection were determined by the availability of riparian habitat and resources of their preferred prey. Territoriality however was governed by prey biomass. Neighbouring leopards were territorial, sharing little space (average 11% territorial overlap) and hunting five times more often in the core than in the rest of their territory. During periods of prey richness, females became more territorial and there was a positive “bottom up” effect through subadult recruitment. Density-dependent intraspecific and interspecific competition for limited space regulated the population around carrying capacity, and constrained population growth. These results provide fundamental baseline data about leopard in the absence of human disturbance, or prey constraints. They highlight that, although the influence of optimal nutrition is important in the reproductive health and territoriality of leopard, habitat quality and quantity are ultimately what govern leopard carrying capacity and population size. I provide baseline reproductive, carrying capacity and territorial data for agencies developing policy, and for setting priorities in conservation and management, as well as habitat protection and restoration, for not only this species but other threatened species as well. / Thesis (Ph.D.)-University of KwaZulu-Natal, Westville, 2013.

Wildlife viewing is a form of recreation that is becoming increasingly popular throughout the world, particularly in African protected areas. In order for protected area managers to cater for this demand effectively, managers need to incorporate wildlife viewing recreation into the planning and development of protected areas. Protected area management has traditionally focused on the management of wildlife populations and habitats to the exclusion of visitor recreational needs. Where visitor needs have been incorporated into the planning and development of protected areas, this has been through the provision of inputs such as facilities and wildlife. The experience-based management (EBM) approach to recreation however proposes that people engage in particular recreation opportunities in order to attain certain desired benefits or outcomes. Madikwe Game reserve provides visitors with the opportunity to view a wide variety of game. The aims of this study were to (1) provide an understanding of what visitors sought from their experience~ regarding wildlife viewing in Madikwe Game Reserve (2) classify the types of experiences desired by visitors to the reserve using the EBM model as a framework and (3) examine management implications of results. A survey of visitors was conducted in the reserve using a Pre-visit and a Postvisit questionnaire. Results from 178 respondents indicated that well-known species as well as rare/endangered specie: were the most popular among visitors. Respondents were generally very satisfied with their wildlife viewing experiences in terms of species abundance and variety, and information received about animals. The results also suggest that additional information about items other than wildlife could enhance the experiences of visitors to Madikwe. Three distinct wildlife viewing experiences desired by visitors were identified, namely a High Involvement Experience, which had the highest interest in almost all recreational opportunities, a Generalist Experience characterised by a moderate interest in recreational opportunities and an Occasionalist Experience that displayed the least interest. While the Occasionalist Experience is presently adequately catered for in Madikwe, lodge and park manager can provide for the High Involvement and Generalist Experiences more efficiently by expanding the wildlife viewing experience that is currently offered in the reserve. This would be done primarily through the expansion of informational items provided, and the development of activities associated with wildlife viewing. The success of such measures would be dependent on the adoption of a cooperative strategy between lodge managers, park managers and other relevant stakeholders. / Thesis (M.Env.Dev.)-University of KwaZulu-Natal, Pietermaritzburg, 2004.

In recent years several regions in South Africa including KwaZulu-Natal (KZN) have experienced a significant shift in their farming practice, moving away from conventional commercial farming to the apparently more lucrative private game farming. This is clearly evident in the northern parts of KZN, where most Private Game Reserves (PGRs) occur in semi-arid areas, which are poorly suited to agriculture or livestock farming. These conversions of land cover or change in land use (i.e. from conventional commercial farming to private game farming) is occurring fairly swiftly even though the extent (area of change), impacts and implications of this trend have not been established. Very limited scientific research has been carried out on the environmental effects, impacts and implications of these land cover/use changes related to PGRs. In an attempt to document these changes as well as the environmental implications, three PGRs i.e. Kube Yini, Thanda and Phinda and the Mkhuze Game Reserve within northern KZN were selected for in-depth study. Mkhuze, which is owned and managed by the state, was selected as the control for this study, as it has remained relatively undisturbed for a long period of time. A comparison between the PGRs and Mkhuze was carried out to compare the land cover/use changes within the study area since the establishment of the PGRs. Remote sensing software was used to achieve the aim and objectives of this study. Using satellite images from 1990 and 2007, a change detection analysis was performed to determine the land cover/use changes that occurred within the study area during the period 1990-2007. Sixty-four land cover/use classes were generated from the analysis. These sixty-four classes were reclassified into five broad land cover/use classes which were identified as (1) water and riverine vegetation, (2) sand forest, forest and rock faces, (3) woodlands, (4) old fields and disturbed areas, (5) and grasslands. The results illustrated that during the period 1990 to 2007, the land cover/use of the entire study area changed significantly, with the overall land cover/use changes illustrating a decrease in the water bodies and riverine vegetation (6 percent to 3 percent); sand forest, forest and rock faces (19 percent to 16 percent); woodlands (26 percent to 25 percent); and old fields and disturbed areas (26 percent to 25 percent) whilst an increase in the grasslands (23 percent to 31 percent) was evident. Within each individual PGR, significant land cover/use changes related to private game farming were evident. In Kube Yini PGR, a decrease in the water bodies and riverine vegetation (4 percent to 1 percent); old fields and disturbed areas (23 percent to 11 percent); and grasslands (6 percent to 5 percent) were observed whilst an increase in the sand forest, forest and rock faces (31 percent to 47 percent); and grasslands (6 percent to 5 percent) land covers were seen. There has been no change in the woodlands cover (i.e. remains 36 percent). Old fields and disturbed areas have decreased due to the clearing programme that was implemented by Kube Yini, which has also contributed to the increased sand forest, forest and rock faces; and grassland land covers. At Thanda PGR, water bodies and riverine vegetation remained the same overall, while an increase in the grasslands cover (19 percent to 49 percent) was clearly evident. A decrease in the sand forest, forest and rock faces (13 percent to 4 percent); woodlands (29 percent to 18 percent); and old fields and disturbed areas (38 percent to 28 percent) was also noted. This was due to Thanda‟s Management Plan, whereby management had cleared the old fields and disturbed areas to accommodate the increased tourist accommodation, increased game species as well as the increased space that is required for these species. Furthermore over the recent years, Thanda has been practicing bush clearing and thinning of vegetation, which has resulted in the creation of a grassland landscape. A grassland landscape was required for viewing the game species, which resulted in a decreased sand forest, forest and rock faces; woodlands; and old fields and disturbed areas. In Phinda too, the water bodies and riverine vegetation remained the same (i.e. 1 percent). The sand forest, forest and rock faces (32 percent to 19 percent); and woodlands (43 percent to 41 percent) decreased whilst an increased old fields and disturbed areas (15 percent to 26 percent); and grasslands (9 percent to 13 percent) were observed. Sand forests are sensitive and decrease in extent rather easily as a result of disturbances experienced in the area. The decrease in sand forest, forest and rock faces during this period may be due to the increased wildlife and especially elephant population experienced on this reserve. The woodlands land cover decreased in size as a result of clearing to accommodate tourist infrastructure as well as creating a grassland landscape i.e. open space and clear viewing for the game species. The changes in Phinda‟s land cover resulted can be explained by the different management practices that were practised on the reserve. The results from the PGRs were compared against Mkhuze so as to determine whether private game faming was in fact changing the landscape of the study area. During this period 1990 to 2007, Mkhuze experienced only slight changes within it‟s land cover/use classes: water and riverine vegetation (2 percent to 1 percent); sand forest, forest and rock faces (16 percent to 15 percent); and grasslands (29 percent to 26 percent) decreased whilst an increase in the woodlands (27 percent to 30 percent); and old fields and disturbed areas (26 percent to 28 percent) were noted. Since Mkhuze is managed by the state with its priority being conservation first and not increased monetary gain, competition between species and habitants occurred with the least amount of human influence. It is most likely that the water and riverine vegetation; and grasslands decreased due to the drought experienced in the area between 2001-2006, which may have led to the increased old fields and disturbed areas. In addition woodlands may have increased due to its resistance to harsh conditions. Also contributing to the increased old fields and disturbed areas could be the increased wildlife numbers experienced in the reserve. The slight drop in the sand forest, forest and rock faces land cover most likely resulted due to natural competition between the vegetation species as well as management realising the significance of the sand forest, thus assisting in the preservation of the sand forest, forest and rock faces cover. An accuracy assessment was carried out to determine the accuracy of the image classification, and a 75 % accuracy was achieved for the overall analysis. The results illustrate that during the period 1990-2007, the study area‟s land cover changed significantly and that this was mostly like due to the changes within the land cover of the individual PGRs. Climatic data for this period was also taken into consideration to see whether climate variations had occurred and whether this had impacted the change in land cover/use of the study area. The results from the PGRs were compared against the state reserve results as well as the climatic data. The results suggest that the majority of the land cover/use changes within the study area had occurred largely due to private game farming rather than natural processes. The climatic variables such as temperature of the area remained fairly unchanged whilst humidity remained fairly high. With the exception of the drought experienced in the area between the period 2001-2006, it is most likely that the changes in the area are the result of the change to private game farming. Private game farming is having a significant effect on the landscape of the study area and this is largely due to the different management practices practised on each individual PGR. It is recommended that PGRs be monitored and regulated consistently in order to prevent the degradation of the environment and its ecosystem. The study exhibits the value and usefulness of satellite imagery in detecting land cover/use changes related to private game farming. Landsat 5 TM imagery was the most suitable and accurate in identifying and mapping broad scale land cover/use changes related to private game farming. If a more detailed level of investigation is required, the use of higher resolution imagery (e.g. Spot/Landsat 7) is advised. / Thesis (M.Sc.)-University of KwaZulu-Natal, Westville, 2010.

M.Sc. / Mabula Game Reserve initiated an extensive grass-monitoring programme ten years ago. At the same time, management kept records of factors such as rainfall, burning, game numbers and game composition that might have an influence on grass production. They also conducted an extensive mapping programme over a number of years to familiarise themselves with the physical phenomena of Mabula, namely geology, soils, slope and aspect, that influence grass growth. The most important aspect of their mapping programme was to divide the reserve into different vegetation communities. The objective of the study is to supply Mabula Game Reserve with clear-cut evidence of the condition of the grazing resources at Mabula. The aim of the study is to assist the management of Mabula with future guidelines in the veld management of the reserve. The overall purpose of this study was to investigate the veld management efficiency of Mabula Game Reserve by establishing whether the ecological status of the grazing had been degraded or improved over the study period. This researcher analysed the grass monitoring data to determine veld condition indices, grazing values, perenniality and to establish the amount of climax and pioneer grass. He sequenced his research by: • classifying grass species in terms of the ecological index method to determine whether grass had been utilised in a sustainable manner or over-utilised;. • calculating the grazing capacity for each vegetation community over the past ten years; • highlighting those grass species in the analyses that showed abnormal growth or decline over the ten years; and • interpreting the results in terms of the ecological status of the relevant area.. The study was conducted firstly by categorising the grass monitoring stations into different vegetation communities. Ecological indices for each vegetation community was determined. The grass species that showed the biggest increase/decrease in the ten years of monitoring were seperated to determine the ecological significance of the increase/decrease. The ecoloical indices of the different vegetation communities were compared with each other to determine the overall improvement or deterioration of the veld quality. By re-calculating the game numbers for every year, this researcher was able to determine the Game Livestock Unit for the area. With this figure, he was able to differentiate between the grass utilisers, comparing them from year to year in terms of their grazing capacity. The study revealed that in order to obtain an objective view of the condition of the veld, it is necessary to evaluate the condition of the veld in terms of each vegetation community, as well as the total area. Such an approach would enable the manager to plan according to a specific vegetation community. The study also made it clear that it is necessary to reduce the number of game, but that this must be done by culling the appropriate group of utilisers. The necessity of introducing a browsing monitoring programme soon became evident. The study showed how the grass and game monitoring analyses for the area can be applied in association with the rainfall figures in forward planning in order to manage the sustainable utilisation of biological resources and to minimise the effects of drought conditions. The researcher concluded this study by highlighting the shortfalls of the ecological index method in determining the condition of the veld, and recommended other methods i.e. the Benchmark Method, Key Species Method, and the Weighted Key Species Method to supplement the present method. Another recommendation was to collect two different sets of data during monitoring to avoid conflicting results, as was the case with the 2004 survey. One data set should include herbs and forbs while the other set should only contain grass species.

M.A. (Geography) / In the last century, the establishment of protected areas has become an important part of managing South Africa’s wilderness and wildlife. The notion of untouched protected areas is becoming contested in a time when developing nations are seeking to improve quality of life for their citizens. With the promulgation of the NEM: PAA of 2003, resource sharing with local rural communities has become an important policy guideline for protected area management authorities. An example of a protected area where management has sought to facilitate resource access in neighbouring rural communities is Mkuze Game Reserve (MGR), now part of the greater Isimangaliso Wetland Park. This study uses a mixed methods approach to combine local rainfall records, census archives, and remotely sensed data with qualitative interview data in order to investigate spatial, social and quantitative aspects of anthropogenic land-cover change between 1979 and 2008. If the proposed balance between development and conservation is to be achieved in this ecologically diverse locale, a deeper understanding of contextual relationships between human activity and environmental change will be vital. Results showed that while rainfall was cyclic, natural land-cover decreased consistently in densely populated rural areas. In contrast, protected areas such as MGR showed little change in land-cover indicating that human activity and cattle have a significant impact on the land surface in the Mkuze Region. While many local residents in the rural communities living adjacent to MGR recognise their role in the achievement of development-conservation objectives, a history of exclusion from MGR has left a legacy of negative perceptions towards MGR in these communities. In order to mitigate natural land-cover loss, local communities must have positive perceptions about MGR and become involved in its management. Useful inferences have been made from the results regarding the management of human population and activities around the borders of protected areas in South Africa.

Remaining viable cheetah (Acinonyx jubatus) populations in Africa are threatened by direct persecution through conflict with farmers and habitat degradation and fragmentation. Botswana is considered a stronghold for free roaming cheetahs in Africa, yet the country has had relatively limited research on its cheetahs, and information from the east of the country is lacking. Data on the current status of populations is thus required to make informed management decisions. My study provides estimates of population density, abundance, distribution and status for the demographically open cheetah population of the Northern Tuli Game Reserve (NOTUGRE) in Botswana. The effectiveness of two population monitoring methods, namely camera trapping and a photographic survey, were also investigated. Moreover, I report on the level of conflict between livestock farmers and predators on rural communal farmlands within and adjacent to NOTUGRE. Data were collected between May 2012 and November 2013. Results indicate a low population density of 0.61 ± 0.18 adult cheetahs per 100 km² and a minimum population size of 10 individuals (nine adults and one cub). Camera traps placed at cheetah scent-marking posts increased detection rates and provided ideal set up locations. This approach, together with Spatial Explicit Capture- Recapture (SECR) models, is recommended for future studies. The long-term studies that are required to better understand the status of cheetahs in Botswana do not exist. Thus, photographic surveys may provide an alternative method for providing baseline data on population numbers, distribution and demography. The third aspect of my study gathered information on levels of livestock loss and human tolerance of predators through the use of interviews (n = 80). Conflict with subsistence farmers is a concern as livestock depredation is relatively high (9.1% of total livestock owned) and farmers had an overall negative attitude towards conservation of large predators. My results suggest that human-predator conflict in this area is more complex than the direct financial loss from depredation. Hence, reducing depredation rates alone is unlikely to change farmer tolerance of wildlife on farmlands. Improved, responsible farm management, including self-responsibility for livestock rearing, and positive appreciation for wildlife are necessary. The NOTUGRE cheetah population requires further research to understand possible threats to the population. Furthermore, a better understanding of the connectivity between cheetahs of NOTUGRE, South Africa and Zimbabwe is required. The number of cheetahs within NOTUGRE is too small to sustain a viable population, hence conserving cheetahs outside of the protected area should be a priority for the conservation of the population. This can only be achieved through assistance and involvement from national authorities, local people and conservation organisations.

During the last 25 years, the Eastern Cape Province of South Africa has seen the establishment of many small to medium sized (≤ 440km2) game reserves. These reserves have reintroduced many of the larger indigenous wildlife that had been extirpated by the early 20th century. As such, these reserves and wildlife introductions have created many research opportunities, including investigations on the ecology of reintroduced carnivores in the Thicket biome. Brown hyaenas (Hyaena brunnea) are one of the large carnivore species that have been reintroduced into the area. As these animals have predominantly been studied in more open, arid systems, their reintroduction has provided an excellent opportunity to study the species in an alternate natural habitat. Information gathered from such investigations adds to our knowledge of the species and also provides information for the management of brown hyaenas within small, enclosed reserves. Data were collected over the period of one calendaryear, from February 2013 to February 2014 at Kwandwe Private Game Reserve in the Eastern Cape, South Africa. Brown hyaena population estimates were calculated using capture-recapture methods from individually identifiable images captured during a three month camera trapping survey. Images of brown hyaenas were separated into left- and right-side profiles. Twenty-eight individuals were positively identified from left-side images and 27 from right-side images. Non-spatial and spatially explicit capture-recapture analyses were both run in the program DENSITY 5.0. Density estimates ranged from 14 to 20 individuals/100km2 (equivalent to a total abundance of 26-37 individuals) depending on the method used. Despite the range of estimates, all are considerably higher than in other areaswhere densities have been calculated. Satellite/GPS collars were fitted to three individual brown hyaenas (two males and one female) to measure their home range size and use of space. Home range size was calculated using two different methods, Kernel utilisation distributions and Brownian bridges. Home range estimates were similar using both methods; however Brownian bridge methods appeared to exaggerate the use of space by individuals. Kernel home range sizes for the three individuals ranged between 42.62km2 and 79.88km2. These estimates are considerably smaller than previous findings from other parts of Africa and suggest that sufficient resources may be available within this enclosed system. The results from this study suggest that brown hyaenas are successful generalists in this enclosed system and are able to persist at high densities and occupy relatively small home ranges. This information is important for the managers of small reserves who wish to reintroduce brown hyaenas. Should brown hyaenas be introduced into reserves in the Thicket biome with sufficient resources, the numbers are likely to increase rapidly. If high numbers are not desired then preventative measures (e.g. contraception) should be investigated before release.

Anthrax was associated with the death of a large number of animals in the Selous Game Reserve. The significance of this disease to the populations of these animals was of concern to the reserve's management. Models are presented of the evolutionary effects of four host-pathogen relationships. Based on a demographic study of the wildebeest and a study of the characteristics of the disease, the anthrax-wildebeest relationship was compared with the models. The results of the study indicate that even though anthrax was responsible for the death of approximately 10% of the wildebeest, it had a balanced relationship with the population. The pathogen was probably an asset to its host population's continued existence rather than a hazard, as its mortality was associated with animals that appeared to be a disadvantage to the wildebeest population. If the management of the reserve wished to reduce the occurrence of anthrax, it is suggested that they reduce the number of wildebeest older than calves. In addition to maintaining a stable age configuration, this would improve the quality of the habitat, reduce the number of animals in poor condition, and thus reduce the number dying of anthrax. In addition, I deal with several other topics in Appendices. / Science, Faculty of / Zoology, Department of / Graduate

Between 1970 and 1975 Natal Parks Board hel .icopter counts of waterbuck ( Kobus ellipsiprymnus) in the Umfolozi Game Reserve (UGR) declined from 1 098 to 494. The purpose of this project was to study the ecology of waterbuck with an emphasis on population regulating mechanisms, so that the reality and significance of this decline could be assessed. Density estimates for 1976-1977 were over twice the 1976 helicopter count of 319, but the population was confirmed to be declining. The ecological density of waterbuck represented 11,9 per cent of the metabolic biomass of grazers, which compared well with data from other areas. However, the other species' proportionate contributions to biomass were far from optimal, with an excess of short grass feeders. High juvenile mortality (80,7 per cent), attributable mainly to severe tick (Rhipicephalus appendiculatus) infestations, was the principal cause of the present decline. These severe infestations were probably symptomatic of poor condition of the whole waterbuck population, which meant that the ultimate cause of juvenile mortality lay elsewhere. The hypothesis that the waterbuck population was in a stressed condition was supported by their habitat utilization patterns, their feeding ecology and aspects of their behaviour. The dynamics of interspecific associations suggested that the primary habitat of Waterbuck was that now occupied by nyala (Tragelaphus angasi) and what had been recorded were the results of competition pushing waterbuck into an unfavourable habitat. Feeding studies indicated that nutritious food for herbivores represented a limited resource in UGR during winter, for which waterbuck may be largely outcompeted and that nyala and impala ( Aepyceros melampus) were the rnain competitors . Interspecific competition was the probable ultimate cause of the decline of waterbuck in UGR. It was recommended that the numbers of nyala and impala be reduced and that this action be integrated into a more precise management plan for UGR than exists at present. / Thesis (DSc)--University of Pretoria, 1978. / gm2014 / Zoology and Entomology / unrestricted