Thesis (MSc)--University of Stellenbosch, 2005. / ENGLISH ABSTRACT: This study investigated the composition of Themeda triandra Renosterveld in part of the Grootvadersbosch Conservancy and the effects of selected environmental and management variables to provide guidelines for promoting the presence of Themeda triandra in the veld. The Zürich-Montpellier phytosociological method was used to determine the composition of the Renosterveld communities. The point quadrat method was used to determine the cover of Themeda triandra at three grass dominated sites and compare cover from one site with past cover measurements at the specific site. Ordination was used to examine the effects of the environmental and management variables on the plant communities. Two community groups, five communities and five subcommunities were identified and described. The Themeda triandra – Stoebe phyllostachys Grassland Community Group consists of two communities of which one has two subcommunities. The Themeda triandra – Elytropappus rhinocerotis Shrubland Community Group consists of three communities of which one has three subcommunities. The vegetation units described in this study have not been described previously in the literature. One community belongs to Silcrete Fynbos while another subcommunity is transitional between Renosterveld and thicket and gallery forest. The rest of the plant communities fit the definition for Renosterveld (the local Renosterveld type is Eastern Rûens Shale Renosterveld). Cover of Themeda triandra did not differ significantly either between sites or between years. The comparison of Themeda triandra cover between years was done at a site that had been burnt between the last two sampling times yet the cover was not significantly different. This indicates that fire and other management practices did not have a negative impact on Themeda triandra at the site. Unconstrained ordination of the dataset in which cover/abundance values were included, grouped the relevés by moisture and disturbance but the presence/absence dataset indicates that the two community groups have a slight transitional overlap. Constrained ordination of both datasets with a) soil variables and b) management variables, both showed a tendency to separate the relevés into community groups, that did not happen with topographic and vegetation variables. Ordination did not separate the community groups into their subdivisions. The soil variables (both nutrients and texture) influence the vegetation structure and the community distribution. Under the existing grazing management regime, fire at three to five year intervals promoted the dominance of Themeda triandra by affecting the structure of the plant communities, the abundance of species other than Themeda triandra, and influencing which plant community is present. The use of fire as a management tool was regulated by the importance of the natural veld pastures to the farmers. The natural veld pastures are not suitable for dairy cattle in milk. Thus dairy farmers are less likely to burn the natural veld (no planned burns only chance fires) than those who farm with mutton or beef (planned burns on a three to five year interval). / AFRIKAANSE OPSOMMING: Die studie het die samestelling van Themeda triandra Renosterveld in ’n deel van die Grootvadersbosch-Bewaria en die uitwerking van geselekteerde omgewings- en bestuursveranderlikes ondersoek om riglyne vir die bevordering van Themeda triandra in die veld daar te stel. Die Zürich-Montpellier fitososiologiese metode is gebruik om die samestelling van die Renosterveldgemeenskappe te bepaal. Die puntkwadraat-metode is gebruik om ’n skatting van die dekking van Themeda triandra by drie gras-gedomineerde persele te bepaal en om ’n vergelyking te maak tussen die huidige en vorige dekking van ’n enkele perseel. Ordinasie is gebruik om die invloed van omgewings- en bestuursveranderlikes op die plantgemeenskappe te bepaal. Twee gemeenskapsgroepe, vyf gemeenskappe en vyf subgemeenskappe is geïdentifiseer en gedefinieer. Die Themeda triandra – Stoebe phyllostachys Grasland-gemeenskapsgroep bestaan uit twee gemeenskappe waarvan een in twee subgemeenskappe onderverdeel is. Die Themeda triandra – Elytropappus rhinocerotis Struik-gemeenskapsgroep bestaan uit drie gemeenskappe waarvan een in drie subgemeenskappe onderverdeel is. Die plantegroei-eenhede wat in die studie beskryf is, is nie voorheen in die literatuur beskryf nie. Een gemeenskap behoort aan Silkreet-fynbos en ’n ander subgemeenskap is ’n oorgangsfase tussen Renosterveld en struikbosveld of woud, terwyl die res van die plantgemeenskappe binne die definisie van Renosterveld val (die plaaslike Renosterveld tipe staan bekend as Oostelike Rûens Skalierenosterveld). Die bedekking van Themeda triandra het nie betekenisvol gevarieer tussen óf die verskillende lokaliteite óf die verskillende jare nie. Die vergelyking van Themeda triandra-bedekking oor tyd is onderneem in ’n gebied wat tussen opnames gebrand is. Die bedekking het nie betekenisvol verskil nie. Dit dui aan dat vuur en ander bestuurspraktyke nie ’n negatiewe invloed op Themeda triandra in hierdie gebied het nie. Onbeperkte ordinasie van die datastel met die vergelyking van bedekking/volopheidwaardes, groepeer die relevés volgens vogtigheid en versteuring, terwyl die datastel ten opsigte van teenwoordigheid/ afwesigheid aandui dat die twee gemeenskapsgroepe ’n effense oorgangs-oorvleueling het. Beperkte ordinasie van beide datastelle met a) grondveranderlikes en b) bestuursveranderlikes, toon albei ’n neiging om die gemeenskapsgroepe te skei, wat nie gebeur het met die topografiese- en plantegroeiveranderlikes nie. Ordinasie het nie die gemeenskapsgroepe onderverdeel in gemeenskappe of subgemeenskappe nie. Die grondveranderlikes (beide voedingstowwe en tekstuur) beïnvloed die struktuur en die verspreiding van die plantegroei. Met die bestaande weidingsbestuur bevorder brande met ’n interval van tussen drie tot vyf jaar die oorheersing van Themeda triandra deur die struktuur van die teenwoordige plantgemeenskap te beïnvloed, deur die getal van die verskillende plantsoorte te beïnvloed en selfs deur die plantgemeenskap se voorkoms te beïnvloed. Die waarde wat die boer aan die natuurlike veld as weiveld heg bepaal die mate waartoe hulle veldbrand gebruik as ’n deel van bestuur. Die natuurlike veld is nie geskik vir melkkoeie wat in die melkproduksiestadium is nie. Die melkboere is dus minder geneig om die natuurlike veld te brand (geen beplande brande nie net kans brande) as die boere wat met vleisbeeste of skape boer (beplande brande elke drie tot vyf jaar).

The Kafue Flats floodplain wetland system in southern Zambia is under increasing climate and human pressures. Firstly, drought episodes appear more prevalent in recent years in the region and secondly, two dams were built on the lower and upper ends of the wetland in 1972 and 1978, respectively, across the Kafue River which flows through the wetland. The study uses multi-temporal remote sensing to assess change in extent and vigour of green vegetation, and extent of water bodies and dry land cover on the Kafue Flats. The change detection's management value is assessed. Four normalised, co-registered digital Landsat images from 24 September 1984, 3 September 1988, 12 September 1991 and 20 September 1994 were used. The main change detection method used was comparison of classifications, supplemented by Normalised Difference Vegetation Index (NDVI) and Principal Component Analysis (PCA) change detection. Ancillary land use and environmental data were used in interpreting the change in the context of cause and effect. The results indicate inconsistent trends in the changes of most land cover classes, as a result of manipulation of the wetland by man through annual variations in the timing and magnitude of regulated flows into the wetland, as well as burning. However, the results also show spatial reduction in the wetland's dry season dense green reed-grass vegetation in upstream sections which are not affected by the water backing-up above of the lower dam. Sparse green vegetation is replacing the dense green vegetation in these upstream areas. It is inferred that this dry season degradation of the wetland threatens bird species which may use the reeds for dry season nesting. It is proposed that ground surveying and monitoring work at the micro-habitat level is necessary to ascertain the implications of the losses. It is concluded that, in spite of difficulties, multi-temporal remote sensing has a potential role in wetland change assessment on the Kafue Flats at the community level, but that it needs to be supplemented by targeted, micro-habitat level ground surveys.

The South African grassland biome is disappearing rapidly through advancing development and change in agricultural land use. One of the most threatened grassland types, Midlands Mistbelt, in the KwaZulu-Natal Midlands is an extremely diverse and home to many endemic species across an array of taxa. Three taxa, namely, grasses, grasshoppers and butterflies represent various trophic levels, which are important to the functioning of the grasslands. Ten grasslands were sampled by walking ten fifty metre transects for a twelve-month period. The grasslands were selected as they represented a range of management practices and varying environmental conditions. Using Indicator Species Analysis (ISA) twenty-two species of grasshopper were identified as indicators of environmental variables and management practices. The abundances of the various species indicated the intensity of the management regimes or disturbances. Using the twenty-two grasshopper species abundances and a three hundred point sampling assessment of the grasses creates an assessment tool that can rapidly appraise the management of the grassland, but due to lack of data for other taxa, cannot assess whether management practices for the focal taxa create congruent results for non-focal taxa. Two of the three taxa proved to be good indicators of grassland health, whilst the third, butterflies were ineffectual, due to low abundance and richness. From the results it was concluded that burning was taking place to frequently, and required a reduction to every four years, as this would improve butterfly richness and abundance, and increase abundance of endemic and flightless grasshopper speCies. A rotational grazing system needs to be implemented at sites where continual grazing takes place, wildlife or livestock, impacts on the grassland condition and species diversity. Increasing habitat heterogeneity increases species diversity, and allows later successional species to be included in the grasshopper assemblage. Management of the grasslands in the KwaZulu-Natal Midlands needs to be more responsive and adaptive. In addition, small fragment management needs to be intensified to provide a range of habitats and refugia that will suit all species. This study advocates the use of grasshoppers and grasses as suitable biotic indicators of grasslands in the KwaZuluNatal Midlands. / Thesis (Ph.D.) - University of KwaZulu-Natal, Pietermaritzburg, 2005

Aristida junciformis subsp. junciformis Trin. et Rupr. is an indigenous, densely tufted, evergreen perennial grass associated with the degeneration of grasslands over large areas of South Africa. More than two hundred thousand square kilometres of veld, c. 17% of the total land area of South Africa, contains A. junciformis. The aim of this study was to improve our understanding of the mechanisms by which this species invades and dominates grassland, specifically in this study, Southern Tall Grassveld. Aristida junciformis also has a low nutritional value resulting in a very low grazing value. The unpalatability of the plant is due to the high tensile strength of the leaves, whose hard, fibrous laminas are very difficult for a grazing animal to crop once their length attains 30 cm or more. This species lack of response to conventional grazing practices has often resulted in a grassland in which the carrying capacity has diminished to such an extent as to be virtually useless for grazing in a relatively short time. The extent of encroachment of veld by A. junciformis appears dependant on the frequency of disturbance and rest afforded to the veld. Annual burning and mowing maintained the species composition of A. junciformis at levels <10% whilst protection from fire, burning or grazing allows this species to dominate the herbaceous layer at levels approaching 90%. Aristida junciformis does not become moribund and is unlikely to die if left undefoliated. As the abundance of A. junciformis increases, veld deterioration gradually accelerates through increased selective grazing on remaining palatable species. Competition from adult A. junciformis plants increases the mortality of seedlings and constrains growth of surviving seedlings. Tiller production of Aristida junciformis seedlings declined from an average of 5.2 tillers per seedling in a no competition situation to 2.2 tillers per seedling when subjected to full competition. Tiller production of T. triandra seedlings decreased from 9.6 tillers per seedling free from competition to 3.3 tillers per seedling subject to full competition for resources. Once these seedlings have become adult plants they are avoided by grazing animals and grazing pressure on the remaining palatable species consequently increases. This allows the A. junciformis plants to increase their size and density in the sward. Once this density is sufficiently high, grass seedlings of either A. junciformis or more desirable grass species such as T. triandra are unable to establish, eventually resulting in a monospecific stand of A. junciformis if left undisturbed. Large amounts of caryopses are produced by A. junciformis - up to a 19 000 caryopses from a large mature plant (c. 38 000 caryopses/m²). Of these c. 40% is likely to be infertile but the remaining c. 60% viable caryopses are dispersed in a typically leptokurtic distribution, the number of caryopses dispersed rapidly declining within a 10 metre radius. A large proportion of the caryopses was trapped in surrounding foliage but in open swards caryopses had greater opportunity to be blown further distances than in a closed sward. The density of A. junciformis caryopses on the soil surface was positively correlated with the density of flowering adult plants in the area and varied from 400 caryopses/m² (density of parent plants c. 0.6 plants/m²) in less effected areas to 11000 caryopses/m² (density of adult plants c. 6 plants/m²) in severely encroached areas. The primary function of the three awns appears to be orientating the caryopsis correctly for in its descent from the parent plant to expedite germination. Caryopses orientated vertically with the awns uppermost exhibited the highest and most rapid germination (67%) compared to caryopses lying horizontally (35%) whilst only 1% of inverted caryopses germinated. Caryopses trapped in litter and effectively held off the soil surface failed to germinate. Removal of the glumes from A junciformis seed greatly enhanced the rate and overall germination of the seed except for inverted seed of which <1% germinated. The highest numbers of A. junciformis seedlings (32 seedlings/m²) were found in those areas with the highest density of caryopses on the soil surface. Despite the large amounts of caryopses produced, dispersed and landing in apparently suitable micro-sites for germination, comparatively few A. junciformis seedlings (n=992) were found and overall germination ranged between 1 % and 4% of the initial caryopses density on the soil surface. Seedling survival through winter was low with only 13% surviving to the following spring. The basal areas of A. junciformis increased overall by 66% whilst that of other grass species increased overall by only 3% in the time monitored. These results suggest that the primary method of encroachment of A. junciformis in the grass sward appears to be through vegetative expansion and not seedling recruitment. Frequent defoliation of the sward and avoidance of overgrazing to enhance the competitive abilities of palatable species and provide as high a fuel load as possible appear to be the most economically and logistically feasible ways to remove or at least inhibit A. junciformis veld encroachment at present. / Thesis (M.Sc.)-University of Natal, Pietermaritzburg, 1998.

The objective of this study was to describe the impacts of the density of Chromolaena odorata (chromolaena) on species composition in coastal grasslands and to investigate serial changes in the vegetation following the implementation of a burning programme. The thesis deals with key ecological concepts and issues, so a comprehensive literature review is included. Chromolaena invades coastal grasslands that are not burnt regularly (i.e. biennially). Grasslands that were not burnt for 30 years were seral to secondary forest. The successional pathway from open grassland to closed canopy forest varied according to soil type. Coastal grasslands on Glenrosa soils were characterised by savanna at an intermediate stage between the grassland and forest states. Shading ended the persistence of savanna species (e.g. Combretum molle, Dichrostachys cinerea and Heteropyxis natalensis) in forest, whereas forest precursors (e.g. Canthium inerme, Maytenus undata and Protorhus longifolia) only established where fire was absent. Chromolaena infestations were characterised by multi-stemmed adult plants of variable height (i.e. 1-3 m), depending on soil type. Regic sands did not support stratified woody vegetation and chromolaena infestations were self-supporting, reaching a maximum height of 1.5 m. Glenrosa soils supported tree communities and chromolaena reached more than 3 m in places. The density of chromolaena affected species composition in grasslands with moderate to dense stands (> 5 adult plants m ¯² or >50000 shrubs ha ¯¹). Chromolaena stands became monospecific when the number of adult plants exceeded 7 m ¯². Succession to forest also ceased once chromolaena became thicket-forming. Fire-induced mortality of the chromolaena depended on grass fuel loads. Grass cover of 30% (c. 1 000 kg ha ¯¹) was required to achieve 80% mortality of the parent infestation after the initial burn. Dense infestations could only be killed by running head-fires from adjacent grasslands into thickets. Under conditions where head-fires could not be used, infestations were slashed and burnt at the height of the dry season (July to August) to achieve an 80% kill rate. Seedlings were killed (99%) by annual burning in sparse (≤ 10000 shrubs ha ¯¹) to moderate < 50 000 shrubs ha ¯¹) infestations. The suppression of chromolaena and other alien species, establishing on bare ground after clearing dense infestations, required chemical control until grass cover was sufficient (i.e. 1 000 kg ha ¯¹) to effect uniform burning. Certain secondary alien invaders (e.g. Lantana camara, Psidium guajava and Solanum mauritianum) persisted by coppicing profusely after fire and herbicides need to be integrated into burning programmes when these species occur. Grasslands on regic sands (e.g. Ischaemum fasciculatum, Panicum dregeanum and Themeda triandra) were more resilient to the modifying effects of woody vegetation, than grasslands on Glenrosa soils. Grasslands on Glenrosa soils did not revert to an open state but persisted as ruderal savanna grassland (e.g. Eragrostis curvula, Hyparrhenia tamba and Cymbopogon validus) once fire-resistant tree species (e.g. Combretum molle and Heteropyxis natalensis) had established. Depending the objectives for land management and the vegetation's condition, coastal grasslands can be rehabilitated and managed in multiple states, i.e. grassland, savanna or forest communities. A state-and-transition model based on the empirical data recorded in the study is presented and shows chromolaena altering vegetation states from open grassland to chromolaena dominated thicket. The model illustrates chromolaena thickets as the dominant phase of a moist coastal forest/savanna succession, irrespective of soil type, in absence of appropriate land management practices (e.g. control burning and integrated control of alien vegetation). This model should aid in planning strategies for the control of chromolaena in subtropical grasslands in South Africa. / Thesis (M.Sc.)-University of Natal, Pietermaritzburg, 2000.

Eighty three of Acocks's sites, originally surveyed about 50 years ago, were resurveyed in 1996 to determine the extent of grassland change in the humid grasslands of KwaZulu-Natal. Sites were relocated using 1:10 000 scale ortho-photos and present land cover was determined for each site. Forty six of the sites that were still under original grassland were further examined to determine present species composition. A survey method was designed that would emulate Acocks's data and comparisons were drawn between original and present species composition. These differences were then analysed together with some environmental variables to try to determine the factors which had the most influence on the change and which environment and management factors are related to the present variation in composition among sites. Of the 83 sites, 26 had changed from natural vegetation to some other form of agriculture such as forestry or cultivation. Most of this change had occurred in the Natal Mistbelt Ngongoni Veld where large areas are forested. Cultivation is found predominantly in the communal areas where subsistence, cultivation practices are employed. The remainder of the sites had changed significantly in terms of their species composition. The most pronounced change had occurred in areas under communal tenure although significant changes had occurred in the commercially farmed areas. The direction of change was also more consistent towards species that commonly predominate in heavily grazed areas in the communal areas compared to the commercial areas. The exact reasons for this were unclear but this could possibly be attributed to heavier stocking rates in the communal areas. Change in floristic composition was also more pronounced at lower altitudes where the mean annual rainfall is lower and the mean annual temperature higher. This could possibly be a result of the vegetation at lower altitudes being less stable and thus less resistant to change. Basal cover differed significantly between commercial and communally grazed sites. Lower basal cover was found in the communal sites where intensive grazing limits the growth of individual tufts. Number of species found at each site did not differ significantly between communal and commercially grazed sites. This study was also a practical implementation of the resurveying of Acocks's sites and the original data set was found to be a useful baseline data set to determine coarse long-term changes in the vegetation. / Thesis (M.Sc.)-University of Natal, Pietermaritzburg, 1997.

M.Sc. / The grassland biome of South Africa harbours rich ecosystem diversity. Some of the distinctive features of grassland biodiversity in South Africa include globally significant centres of plant endemism, half of the country's endemic mammal species, a third of its endangered butterfly species and 10 of 14 of its globally threatened bird species. Grassland is one of the most inadequately maintained biomes in Southern Africa because 23% is under cultivation, 60% is irreversibly transformed and most of the remaining natural area is used as rangeland for livestock. Only 2% of the grassland biome is currently protected. Grasslands provide essential ecosystem services for economic development, but this biome also supports a large human population whose resource demands have serious environmental implications that threaten the grasslands‘ biodiversity. Urbanisation is possibly one of the major immediate threats to the grassland ecology in South Africa. This is also the case in the Cradle of Humankind World Heritage Site (COHWHS) and adjacent areas. New housing complexes and informal housing are encroaching on the COHWHS. Indigenous fauna and flora are being affected by ecologically insensitive urban development. This poses a major threat to the fauna of this region including the insects that occur in grassland habitats. The insects play a vital role as pollinators in grassland habitats and form an essential food source to a range of predators, including grass owls, shrews, bull frogs, lizards and bats. In order to conserve the insects and therefore the food web of which they form part, it is necessary to understand the diversity of the insects in the grassland in the dolomitic areas. The COHWHS is a world renowned heritage site devoted to the origin of humankind and is characterised by dolomitic caves. These caves are also the home of a large population of bats consisting of several species. The negative impact on the grasslands in the COHWHS and surroundings pose a threat to the survival of these bat populations if the food source they depend on is negatively affected. For this reason it is important to determine which flying nocturnal insect species are available in the grasslands surrounding bat roosts in the COHWHS and surroundings. 3 The choice of location for the primary trap site was made on the basis of its proximity to known bat roosts and the fact that it is situated in a nature reserve that, although the river is polluted, contains an otherwise relatively unspoilt grassland habitat. Sampling took place over a period of 14 months during which fluctuations in the insect population was observed. The fluctuations can be ascribed to seasonal climate changes and the three veld fires that occurred during this period. This fluctuation was most evident in the representatives of the Orders Lepidoptera and Coleoptera sampled.

This paper deals with a study of the rangelands in Coleman County just west south and west of the Western Cross Timbers of Texas. This problem consists of classifying certain rangelands in this area according to whether they are excellent, fair, good, or poor, the chief basis for which is their deterioration.

Luis T. Gutierrez, Willard R. Fey. / Includes index. / Bibliography: p. [220]-228.

South African grasslands are under-conserved and there is a need to expand conservation efforts beyond the boundaries of protected areas. While communal grasslands have conservation potential they are generally over-utilized and the impact of communal land-use on biodiversity is poorly studied. At the same time there is pressure on protected areas to allow for the sustainable utilization of biodiversity. The aim of this study was to examine the impact of communal land-use on various components of biodiversity and to make recommendations regarding communal use of protected areas. A fence-line study was conducted to assess the impact of eight years of controlled communal land-use on biodiversity in the uKhahlamba-Drakensberg Park. The communally used land (referred to as the lease land) which was used for controlled grazing as well as plant collection was compared with land under formal conservation. Vegetation was sampled using the importance score method and veld condition assessments. Selected invertebrate taxa were sampled using sweep netting, colour pan traps and transects and were identified to morphospecies level. Multivariate statistics revealed that sites generally grouped according to landscape position rather than land-use. No significant differences were found in diversity, evenness, richness or veld condition between the lease and conservation land. However, more than twenty-five percent of vegetation and invertebrate species were found exclusively in the lease or conservation land, suggesting that different suites of species were supported by the two landuses. Four alien plant species were found exclusively in the lease land, while one vulnerable and one rare plant species were found only in the conservation land. Further research is required to assess whether biodiversity was diminished by controlled communal. While the lease concept may offer potential as a low-use buffer zone, localised damage from cattle paths and weak enforcement of grazing agreements were areas of concern. Keywords: communal grass)ands, grassland flora, grassland invertebrates, transfrontier park. / Thesis (M.Env.Dev.)-University of KwaZulu-Natal, Pietermaritzburg, 2004.