Palaeobotanical records of Holocene climate change in the southern Coast Mountains identify a cooler/wetter Neoglacial period subsequent to 6600 BP. Geomorphic evidence of alpine glacier advance suggests that there were three distinct cooler/wetter periods during the Neoglacial, but this pattern has not been identified in palaeobotanical studies. By careful selection of a sensitive alpine site this thesis has recognised this structure in a palynological record of Neoglacial climate. This continuous record of Neoglacial climate which has the same basis as records of early Holocene climate (i.e. palynological) and hence allows more direct comparisons of the two periods. Pollen spectra, conifer needle macrofossils, organic matter content, and magnetic susceptibility were assessed for a 4800 year continuous sequence of sediment from an alpine lake. Calibration of the Picea/Pinus pollen ratio by using an altitudinal transect of surface pollen samples allowed partial quantification of shifts in treeline. Treeline at the site was at least 85 m above the present level from 4800-3800 BP, suggesting that summer temperatures were at least 0.6°C above the present. High treeline until 3800 BP indicates a relatively late date for the Hypsithermal/Neoglacial transition at this site. Alternatively, the apparent complexity of this transition in the Coast Mountains may be due to difficulties of separating temperature and precipitation signals in many climatic records. Treeline declined to near present levels by 2500 BP and was lower than present from 2500-1500 BP and from 1200 BP until close to the present. Estimates of equilibrium line altitude depression for Coast Mountain glaciers during the Little Ice Age suggest that these periods of lower treeline were due to a cooling of up to 0.8°C. During the last 5000 years the Southern Coast Mountains have experienced fluctuations on the order of 1.5°C.

Palaeobotanical records of Holocene climate change in the southern Coast Mountains identify a cooler/wetter Neoglacial period subsequent to 6600 BP. Geomorphic evidence of alpine glacier advance suggests that there were three distinct cooler/wetter periods during the Neoglacial, but this pattern has not been identified in palaeobotanical studies. By careful selection of a sensitive alpine site this thesis has recognised this structure in a palynological record of Neoglacial climate. This continuous record of Neoglacial climate which has the same basis as records of early Holocene climate (i.e. palynological) and hence allows more direct comparisons of the two periods. Pollen spectra, conifer needle macrofossils, organic matter content, and magnetic susceptibility were assessed for a 4800 year continuous sequence of sediment from an alpine lake. Calibration of the Picea/Pinus pollen ratio by using an altitudinal transect of surface pollen samples allowed partial quantification of shifts in treeline. Treeline at the site was at least 85 m above the present level from 4800-3800 BP, suggesting that summer temperatures were at least 0.6°C above the present. High treeline until 3800 BP indicates a relatively late date for the Hypsithermal/Neoglacial transition at this site. Alternatively, the apparent complexity of this transition in the Coast Mountains may be due to difficulties of separating temperature and precipitation signals in many climatic records. Treeline declined to near present levels by 2500 BP and was lower than present from 2500-1500 BP and from 1200 BP until close to the present. Estimates of equilibrium line altitude depression for Coast Mountain glaciers during the Little Ice Age suggest that these periods of lower treeline were due to a cooling of up to 0.8°C. During the last 5000 years the Southern Coast Mountains have experienced fluctuations on the order of 1.5°C. / Arts, Faculty of / Geography, Department of / Graduate

A well-preserved Paleocene flora of 39 species from the Fort Union Formation near Baggs, Carbon County, Wyoming has been studied. This is the first study of the plant megafossils in this area. The most abundant species collected are: Carya antiquorum Platanus nobilis, Cercidiphyllum arcticum and Platanus raynoldsi. Together they represent 74% of the flora. The flora has been dated as Upper Middle Torrejonian. This report is also the first paleoecological study of any North American Paleocene flora. Two methods were used to determine the paleoclimatic conditions of the flora. The first was an analysis of leaf margin characteristics, comparing them to characteristics of modem floras. The second method was an analysis of leaf size comparing the average leaf length and leaf size profile to that of modern floras. Both of these studies suggest that this flora lived in a warm-temperate to subtropical, seasonably dry to moist climate. A correlation of the identified species to their nearest living relatives and their favored climates supports this conclusion.

Plant indicator species and longitudinal paleobotanical data were used as independent measures to document the human ecological record of the contact period in southern Marin County, California. These data suggest that archaeological and documentary records are insufficient for examining changes in land management and use during the contact period. Prior to A.D. 1579, Western Miwok peoples had not encountered Europeans face to face. This early phase of the contact period is marked the possible introduction of New World species through passive cultural vectors. Two brief encounters occurred between the Miwok and Europeans between A.D. 1579 and 1775. Introduced and weedy plant species from fossil samples appear to confirm these encuentros and confirm the archeological evidence for intermittent contact during the second phase of the contact period. Modern and fossil pollen samples suggest that the intensity of human disturbance is geographically stratified and related to exploration, procurement, and management of specific resources. Coastal prairie, the redwood forest, and Bay salt marshes were the most affected by the second phase of the contact period. Shifts in vegetation diversity and increases in the numbers of introduced and weedy species were compared between ruderal and undisturbed contexts. These data were used as analogs to monitor the final phase of contact between A.D. 1775 to 1817. Hypotheses derived from ethnohistoric and ethnographic sources that suggest rapid shifts in land management practices and changes in plant representation were corroborated by some pollen data. Specifically, the ecological responses to the suppression of anthropogenic burning, changes in land tenure and parcelization, and the initiation of grazing and logging practices were examined. The cumulative impacts of introduced plants, shifts in land management from Miwok to Euroamerican-dominated resource procurement and subsistence practices, and ecological responses of plant species suggests that the contact period might better be defined on ecological terms rather than by purely material cultural or ethnographic definitions. In southern Marin, paleobotanical data provide a measurable indication of the ecological character of the pre-contact landscape and the cultural processes that effectively altered its character during the contact period.

Bibliography : leaves 120-144. / 2v. : ill. ; 30 cm. / Title page, contents and abstract only. The complete thesis in print form is available from the University Library. / Thesis (Ph.D.)--University of Adelaide, Dept. of Botany, 1994

The fossil flora and depositional setting of the early Eocene Falkland site in the southern interior of British Columbia, Canada is reported here in detail for the first time. Falkland is part of a series of fossil localities that occur in a region known as the Okanagan Highlands. These sites represent relatively cool upland environments in the greenhouse world of the early Eocene. Macrofossil collections were obtained from Falkland using an unbiased census approach with systematic sampling through three informal units in the exposed outcrop. A stratigraphic log reveals a lacustrine sequence dominated by finely laminated mudstone or shale with periodic influx of coarser material, punctuated by thin volcanic ash layers. Paleoelevation of the site is estimated based on paleobotanical evidence to have been similar to or slightly higher than modern levels (¡Ý1.3 km) during the early Eocene.<p> Paleoclimate is assessed using both physiognomic and floristic approaches as applied to the Falkland flora. Physiognomic approaches correlate aspects of leaf morphology with climate, while floristic approaches use the tolerances of modern nearest living relatives to infer a climate envelope for the fossil flora. Overall, the different methods give broadly consistent results, with an identifiable zone of overlap in the estimates for mean annual temperature at ~10.5¡ãC, cold month mean temperature at 2.3¨C6.3¡ãC, warm month mean temperature at 20.2¨C23.7¡ãC, and a minimum mean annual precipitation of 82¨C120 cm/yr. Assessment of paleoclimate for the three individual units indicates a cooling trend over time, consistent with a radiometric date of 50.61¡À0.16 Ma that places the site in the waning phase of the Early Eocene Climatic Optimum (EECO).<p> The stomatal frequency of fossil Ginkgo adiantoides from Falkland is used to estimate paleoatmospheric carbon dioxide (pCO2). Results from Falkland indicate that pCO2 was significantly higher than modern (>2x) in the early Eocene, although the upper limit of the estimate is unconstrained due to limitations with modern calibration datasets. Analysis of specimens from the three units indicates that climate and pCO2 were coupled during the EECO. Examination of modern Ginkgo biloba leaves suggests that stomatal density is more likely to be accurately measured than stomatal index. In addition, there is a significant difference between stomatal frequencies of long- and short-shoot leaves, suggesting that this factor needs to be taken into account in modern calibration datasets.<p> The Falkland flora was described in two phases. In the first phase, specimens were assigned to morphotypes, informal categories that ideally correspond to species-level organization. In total, 1561 specimens were assigned to 138 morphotypes encompassing foliage and reproductive structures. The taxonomic literature was then investigated and morphotypes were assigned to formal taxa wherever possible. Gymnosperms are dominated by taxa in Cupressaceae, Pinaceae, and Ginkgoaceae, and there is a diverse angiosperm flora particularly rich in taxa belonging to Rosaceae, Betulaceae, and Sapindaceae. Rarefaction analysis shows Falkland as having diversity comparable to that of the hyper-diverse Laguna del Hunco site in Argentina. These data are consistent with an emerging understanding of high diversity in early Eocene forest communities associated with mild but equable climates. The Falkland flora retains a foundation of common taxa through all three units, including Metasequoia, Ginkgo, and Alnus; however, there is a distinct plant community in the upper unit as angiosperms become more abundant and the assemblage more diverse. Patterns in plant diversity are assessed within a context of changing climate and an active disturbance regime at the Falkland site.

The fossil flora and depositional setting of the early Eocene Falkland site in the southern interior of British Columbia, Canada is reported here in detail for the first time. Falkland is part of a series of fossil localities that occur in a region known as the Okanagan Highlands. These sites represent relatively cool upland environments in the greenhouse world of the early Eocene. Macrofossil collections were obtained from Falkland using an unbiased census approach with systematic sampling through three informal units in the exposed outcrop. A stratigraphic log reveals a lacustrine sequence dominated by finely laminated mudstone or shale with periodic influx of coarser material, punctuated by thin volcanic ash layers. Paleoelevation of the site is estimated based on paleobotanical evidence to have been similar to or slightly higher than modern levels (¡Ý1.3 km) during the early Eocene.<p> Paleoclimate is assessed using both physiognomic and floristic approaches as applied to the Falkland flora. Physiognomic approaches correlate aspects of leaf morphology with climate, while floristic approaches use the tolerances of modern nearest living relatives to infer a climate envelope for the fossil flora. Overall, the different methods give broadly consistent results, with an identifiable zone of overlap in the estimates for mean annual temperature at ~10.5¡ãC, cold month mean temperature at 2.3¨C6.3¡ãC, warm month mean temperature at 20.2¨C23.7¡ãC, and a minimum mean annual precipitation of 82¨C120 cm/yr. Assessment of paleoclimate for the three individual units indicates a cooling trend over time, consistent with a radiometric date of 50.61¡À0.16 Ma that places the site in the waning phase of the Early Eocene Climatic Optimum (EECO).<p> The stomatal frequency of fossil Ginkgo adiantoides from Falkland is used to estimate paleoatmospheric carbon dioxide (pCO2). Results from Falkland indicate that pCO2 was significantly higher than modern (>2x) in the early Eocene, although the upper limit of the estimate is unconstrained due to limitations with modern calibration datasets. Analysis of specimens from the three units indicates that climate and pCO2 were coupled during the EECO. Examination of modern Ginkgo biloba leaves suggests that stomatal density is more likely to be accurately measured than stomatal index. In addition, there is a significant difference between stomatal frequencies of long- and short-shoot leaves, suggesting that this factor needs to be taken into account in modern calibration datasets.<p> The Falkland flora was described in two phases. In the first phase, specimens were assigned to morphotypes, informal categories that ideally correspond to species-level organization. In total, 1561 specimens were assigned to 138 morphotypes encompassing foliage and reproductive structures. The taxonomic literature was then investigated and morphotypes were assigned to formal taxa wherever possible. Gymnosperms are dominated by taxa in Cupressaceae, Pinaceae, and Ginkgoaceae, and there is a diverse angiosperm flora particularly rich in taxa belonging to Rosaceae, Betulaceae, and Sapindaceae. Rarefaction analysis shows Falkland as having diversity comparable to that of the hyper-diverse Laguna del Hunco site in Argentina. These data are consistent with an emerging understanding of high diversity in early Eocene forest communities associated with mild but equable climates. The Falkland flora retains a foundation of common taxa through all three units, including Metasequoia, Ginkgo, and Alnus; however, there is a distinct plant community in the upper unit as angiosperms become more abundant and the assemblage more diverse. Patterns in plant diversity are assessed within a context of changing climate and an active disturbance regime at the Falkland site.

Thesis (Ph. D.)--University of Oregon, 2000. / Typescript. Includes vita and abstract. Includes bibliographical references (leaves 237-250). Also available for download via the World Wide Web; free to University of Oregon users.

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Includes amendments (8 p.) in pocket / Bibliography: leaves 306-331 / iv, 349 leaves, [34] leaves of plates : ill ; 30 cm. / Title page, contents and abstract only. The complete thesis in print form is available from the University Library. / Thesis (Ph.D.)--University of Adelaide, 1988