Angl. resumé

No description available.

At the area of CHKO Trebonsko are known three localities with major appearance of Coast Douglas-fir in four groups. First two groups appear in forest ground of St. Barbora. In the first group are 18 and in the second group 23 individuals. These groups appear in the forest stand no. 408 B 13 (Cisarska group) and in the forest stand no. 457 C 12a (in a fencing at Majdalena). Surveying of these groups took place in 2006 by Mrs. Balounova and Mr. Smahel.Two odd groups appear near by Stankovsky pond and they consist of 13 and 31 individuals. They appear in the forest stand no. 245 C 11 at the area of so-called {\clqq}Stankovske woods`` between Stankov and Mirochov villages, at intersection Rovenska way and the way which leads from Mirochov to Stankov. In a neighbourhood of examined vegetation occurs USES element, it is a regional importance biocenter {--} Elk{\crq}s moors reserve at Mirochov which is also declared as {\clqq}Ramsar{\crq}s area of Trebon lowland moors``.

La formation du duramen est un processus physiologique clé impliqué dans la qualité du bois puisqu'il contribue notamment à sa durabilité naturelle. L'objectif de cette thèse est de comprendre les mécanismes mis en jeu lors de la formation du duramen chez le douglas. L'étude a été menée aux niveaux phénotypiques, biochimiques et moléculaires sur plusieurs génotypes de douglas. Des études phénotypiques, il ressort que la proportion de duramen serait sous influence génétique et très peu environnementale, et que l'expansion du bois de coeur se déroule principalement en automne-hiver. Afin de caractériser la composition biochimique du bois, une nouvelle méthode a été développée. Elle repose sur la dissolution du bois dans des liquides ioniques, les solutions obtenues sont ensuite immuno-marquées à l'aide d'anticorps dirigés contre des épitopes de polymères pariétaux. Cette méthode a permis d'observer l'évolution, cerne par cerne, de la composition pariétale du bois de l'aubier externe au coeur du duramen. Certains polymères sont plus abondants dans l'aubier (arabinanes), tandis que d'autres dans le duramen (pectines, xylanes et galactanes). En parallèle, les gènes impliqués dans la formation du duramen ont été étudiés par RNA-Seq à partir de douglas appartenant à un seul génotype et abattus en hiver. Les résultats montrent que des gènes codant des facteurs de transcription, des protéines de défense, des enzymes de la voie de biosynthèse des phénylpropanoides et des enzymes impliquées dans le remodelage de la paroi sont surexprimés dans la zone de transition par rapport à l'aubier. Des hormones, l'éthylène et le jasmonate notamment, semblent jouer un rôle important dans la maturation de l'aubier. / The heartwood formation is a key physiological process involved in wood quality because it contributes to its natural durability. The goal of this thesis is to understand mechanisms involved in the heartwood formation in douglas fir. This study has been carried out at phenotypic, biochemical and molecular levels from several douglas-fir genotypes. Thanks to phenotypic analysis, we showed that heartwood proportion is probably under genetic control, and little influenced by the environment. In douglas fir, heartwood expansion mainly occurs during autumn and winter. To characterize the biochemical composition of wood, a new method has been developed. The method implies the wood dissolution in ionic liquid, the solution obtained are then analyzed by immunodetection with monoclonal antibodies against plant cell wall glycan epitopes. Thanks to this method, the wood cell wall composition has been studied, ring-by-ring, from the outer sapwood to the inner heartwood. Some polymers are more abundant in the sapwood (arabinans) while others in the heartwood (pectins, xylans and galactans). Then, genes involved in the heartwood formation have been studied by RNA-Seq from trees belonging to one genotype sampled during winter. Results show that genes encoding transcription factors, defence related proteins, enzymes involved in phenylpropanoid biosynthesis and plant cell wall modification are upregulated in transition zone compared to sapwood. Hormones, ethylene and jasmonate especially, seem to play an important role during sapwood maturation.

Time domain properties of 23 tree-ring chronologies derived from a large sample of Douglas-fir and ponderosa pine trees growing in closed-canopy forests of Colorado and New Mexico were analyzed using Box-Jenkins models. A variety of statistical criteria were employed during the identification and validation stages for evaluating the performance of different significant models, and the "best" Box-Jenkins model and its immediate "competitor" were reported for each tree-ring chronology. All series were stationary, and only one was approximately a white noise series. Overall, the ARMA(1,1) model was judged the best for 11 series, and the second for 7 of the remaining 12 series. The AR(2) model was considered the best for 6 series, and the second for 4 of the remaining 17 series. No statistical evidence was found for moving average models, nor for models with more than three different parameters. However, both cyclical (or seasonal) models and third-order autoregressive models with a null second-order parameter were chosen for some series. Fitted models explained from 7 to 51% of the variance of the original ring-index series, with an average of about 22 %. All parameter estimates were positive, and they varied within a relatively small range. From a comparison of all employed criteria, Akaike's Information Criterion (AIC) was the one that performed best in identifying Box-Jenkins models for tree-ring chronologies. Possible distinctions were recognized that would separate the selected models according to species and /or standardization option. Among the 12 chronologies from Colorado sites, all derived using the same standardization option, most Douglas-fir series were best fitted by the ARMA(1,1) model, while most ponderosa pine series were best fitted by the AR(2) model, suggesting a difference in the biological persistence of the two species. On the other hand, most of New Mexico chronologies, developed using various standardization options, were best fitted by the ARMA(1,1) model, and no difference was found between Douglas-fir and ponderosa pine series. Also, models fitted to Colorado chronologies explained a lower amount of variance than those for New Mexico chronologies (averages of 17 versus 29% respectively), and cyclical models were mainly selected for New Mexico series. Although periodicities in Douglas-fir series were probably caused by western spruce budworm outbreaks, similar periodic patterns in ponderosa pine series were more difficult to explain because pine trees in the study area had not been defoliated by that insect. Compared to the original tree-ring chronologies, prewhitened series showed similar short-term growth patterns, reduced long-term growth fluctuations, lower standard deviations, and higher mean sensitivities. Also, cross-correlations between chronologies from the same area usually increased after prewhitening. Since the autocorrelation problem is crucial in analyzing the relationships between different time series, and in removing the biological persistence included in tree-ring chronologies, the Box-Jenkins approach should facilitate the analysis of the dynamic relationships between tree growth and environmental variables.

We examined tree rings from Douglas-fir (Pseudotsuga menziesii var. glauca (Beissn.) Franco) timbers in the Salt Lake Tabernacle, constructed from 1863–1867 in Salt Lake City, Utah. A seismic upgrade to the Tabernacle initiated in 2005 required the replacement of wooden timbers with steel beams. Our objectives were to 1) determine cutting dates for the timbers to identify logs that may have been salvaged from previous structures, and consequently would have greater historical significance, 2) identify the species and provenance of the timbers, and 3) develop a chronology that could extend or strengthen the existing tree-ring record for environmental and historical applications in northern Utah. We built a 162-year floating chronology from 13 cores and 15 cross-sections, crossdated visually using skeleton plots and verified statistically with COFECHA. Statistically significant (p , 0.0001) comparisons with established chronologies from northern Utah indicated that the Tabernacle chronology extends from 1702–1862. Cutting dates ranged from 1836–1863, with most in 1862 or 1863 and a smaller cluster around 1855. The broad range of cutting dates suggests that some of the timbers were used in previous structures, and that some trees were dead before they were cut. This study provides valuable information for the preservation of historical materials, and increases the sample depth of existing chronologies during the 18th and 19th Centuries.

No description available.

The 1994 Beaver Mountain fire ignited the canopies of subalpine fir, Abies lasiocarpa, and spread ground fire into adjacent Douglas-fir forests, Pseudotsuga menziesii var. glauca. Despite shorter flight seasons due to lower annual temperatures and persistent snow, the Douglas-fir bark beetle, Dendroctonus pseudotsugae Hopkins, attacked a range of moderately fire-injured host conifers. Logistic regression models illustrated that in 1995 associated bark beetles selected large diameter Douglas-fir with 60-80% bole char, 60-80% crown volume scorch, and 50-70% probability of mortality due to fire. In 1996 beetle preference shifted to smaller diameter trees with lighter fire injury. Tree size was less significant for predicted attack in 1996 because most large fire-damaged conifers were colonized by beetles in 1995. Beetle populations did not reach outbreak proportions outside the fire boundary, but 53 green trees were also infested in 1997 along the burn perimeter. Log linear tests conducted to quantify beetle emergence supported conclusions that beetles were not only attracted to mature, moderately fire-weakened conifers, but also produced greater brood numbers with up to 60-80 emergence holes/ 1800 cm2. Fire-defoliated trees provided bark beetles with sufficient phloem and limited resistance, allowing beetles to aggregate on areas of viable stem tissue regardless of overall bole char extent.

Tree-ring studies have demonstrated that conifer latewood measurements contain information on long-term North American monsoon (NAM) variability, a hydroclimatic feature of great importance to plants, animals, and human society in the US Southwest. This paper explores data-treatment options for developing latewood chronologies aimed at NAM reconstruction. Archived wood samples for five Douglas-fir (Pseudotsuga menziesii, Mirb. Franco) sites in southeastern Arizona are augmented with new collections. The combined dataset is analyzed along with time series of regionally averaged observed precipitation to quantify the strength of regional precipitation signal in latewood time series and to identify ways of increasing the signal strength. Analysis addresses the signal strength influences of including or excluding ‘‘false’’ latewood bands in the nominal ‘‘latewood’’ portion of the ring, the necessary adjustment of latewood width for statistical dependence on antecedent earlywood width, and tree age. Results suggest that adjusted latewood width chronologies from individual sites can explain around 30% of the variance of regional summer (July–August) precipitation—increasing to more than 50% with use of multiple chronologies. This assessment is fairly insensitive to the treatment of false latewood bands (in intra-annual width and 𝛿¹³C variables), and to whether latewood-width is adjusted for dependence on earlywood-width at the core or site level. Considerations for operational chronology development in future studies are (1) large tree-to-tree differences in moisture signal, (2) occasional nonlinearity in EW-LW dependence, and (3) extremely narrow and invariant latewood width in outer portions of some cores. A protocol for chronology development addressing these considerations is suggested.

La capitalisation sur pied et la substitution d'essences sont deux stratégies sylvicoles analysées dans cette thèse, chacune pressentie pour être favorable au stockage du carbone, enjeu majeur de ce XXIème siècle pour la lutte contre le réchauffement climatique. La première correspond à un ralentissement des récoltes, qui devrait générer un puits de carbone additionnel par l'augmentation des stocks moyens de biomasse sur pied. La substitution d'essences à croissance lente comme le Chêne sessile ou le Hêtre par des essences résineuses à croissance rapide comme le Pin laricio ou le Douglas pourrait entraîner une absorption plus rapide du CO2. L'impact sur le carbone de ces deux stratégies est d'autant plus intéressant à étudier que les gestionnaires forestiers les pratiquent déjà. La construction d'une chaîne de modèles prenant en compte l'ensemble des compartiments dans lesquels le carbone est impliqué, à savoir la biomasse forestière, le sol, mais aussi les produits forestiers issus de l'exploitation, permet d'effectuer des simulations donnant des résultats quantitatifs sur ces pratiques. Nous montrons qu'une capitalisation sur pied engendrerait un puits de carbone additionnel dépendant de l'état initial des peuplements, de la fertilité et de la nouvelle sylviculture pratiquée. Il pourrait s'élever jusqu'à 142 tC/ha après obtention du régime permanent dans le cas le plus favorable. La substitution de feuillus à croissance lente par des résineux à croissance rapide serait également un puits de carbone important (jusqu'à 2 tC/ha/an) pendant plusieurs décennies, mais s'avérerait d'un stock de carbone moyen inférieur sur le long terme.