In recent years, global warming awareness has resulted in an increased demand for clean sources of energy such as hydropower. As a consequence, its impact on riparian vegetation must be studied. In this research, I aimed to assess how different functional riparian groups from northern Sweden respond to hydropeaking (i.e. short-term flow regime changes due to differences in the daily energy requirements). I selected seedlings of eight species natural from Swedish riparian ecosystems which can be grouped in three different guilds (forbs, graminoids and woody) according to their habitat and morphological traits.  A seven week greenhouse experiment in which the seedlings were subjected to two watering treatments that simulated prolonged and deep submergence and frequent and short shallow submergence conditions was developed. I measured the root, stem and leaf biomass, followed leaf changes and stem growth over the weeks and evaluated the health status. The study showed how some species and guilds responded differently to the treatments although survival rates were similar. Forbs was the most resilient group unlike the woody guild.  Graminoids grew longer and thinner roots in frequent submergence situations. Small seedlings appeared to be more sensitive to prolonged submergence. No significant differences were found for leaf variables. Collectively, these results suggest that hydropeaking could significantly modify the riparian vegetation. More and longer studies are needed in order to understand the capacity that hydropower has to modify the riparian vegetation and therefore the riverine ecosystems.

With the ever increasing reliance on hydropower in the world, it is important that its effects on riparian vegetation is well understood. This study aimed to explore the effects of different forms of hydropeaking, which results from the rapid and frequent opening and closure of hydroturbines, on different species groups of riparian species native to northern Sweden. Thirteen riparian species belonging to three different species groups, forbs, graminoids and woody species, were subjected to three different watering treatments simulating water stress, rapid fluctuation and prolonged flooding, which often correspond to different hydropeaking conditions. A fourth treatment maintained individuals under controlled conditions and simulated a non-regulated stream. The treatments were carried out over seven weeks. The results showed that the treatments affect the three tested species groups differently. Woody species responded to all treatments with lower germination, higher mortality and lower root elongation, while forbs and graminoids responded to flooding and fluctuation with increased germination but higher mortality and lower root elongation. Fluctuation and flooding had in general positive effects on germination and short term growth but also increased mortality, particularly fluctuation. There were no significant results regarding change in number of leaves for any treatment or species group. As this study only covered seven weeks it can only account for short-term trends, but it still provides interesting information about general expected trends for a wide variety of species native to northern Sweden and belonging to the main three species groups. Such information is key for river management.

The dissertation presented in this manuscript contributes to river science by providing a detailed overview on the state of the art on the interaction between riparian vegetation and hydrogeomorphological processes, by devising a novel model encompassing most of such processes and by proposing a field methodology aimed at providing means for improving the modelling of such interactions. The state of the art is summarized in an extensive review describing riparian vegetation and hydrogeomorphological processes mutual feedbacks. Such review did not simply seek to describe these feedbacks but, compiling from a large array of results from field, laboratory and modelling studies, provides a set of physical thresholds that trigger system changes. Therefore, processes are not only described terms but also explained with a quantitative approach. Processes description provided the conceptual foundation for the development of the novel simulation model while model parameterization was based on the quantitative information collected in the review. Such novel model, encompasses the main relationships entwining riparian woody vegetation and hydrogeomorphological processes and is able of replicating long term riparian landscape dynamics considering disturbance events, environmental stressor and riparian woody vegetation establishment from seeds and large wood. The manuscript presents the model structure and its conceptual validation by means of hydrological scenarios aimed at testing the coherence of the simulation results with expected system behaviour. Examples of such coherences are vegetation growth rate in response to hydrological regime, entrainment and establishment of large wood in an unconfined river system and vegetation effect on erosion and deposition patterns. Analysis of sedimentation patterns from the modelled results suggested that vegetation flow resistance should be modelled with greater detail. These conclusions pointed the dissertation research towards the testing of a novel class of vegetation flow resistance equations, proposed by different authors, able of describing woody vegetation flow resistance on a physical basis. These equations have the advantage of considering flow stage, plants foliation level and species-specific flexibility. However, the use of such equations is limited by the difficulty of measuring the vegetation properties required as equation-inputs. In order to test if these equations could effectively improve sediment dynamics predictions, a field method was formulated and tested. The field method allows to sample vegetation properties that can be used with these novel class of flow resistance equations. In the manuscript, such method is applied and the resulting vegetation properties used in several modelling scenarios. Such scenario proved that hydraulic variables modelled with these novel flow resistance approaches are more realistic and thus that the model developed during the dissertation could benefit from inclusion of such flow resistance equations in its source code.

With the increase in human population, water resources have become more and more precious. A comprehensive study of water yield characteristics is imperative, especially in water-limited semiarid regions. The objective of this study is to examine spring flow and vegetation cover in a first-order watershed and investigate the herbaceous community structure of upland riparian zones. This study consists of two major components: (1) the effects of environmental factors and vegetation cover on spring flow at Pedernales River upland catchments, and (2) the ecological responses of vegetation to altered flow regimes that result from brush management at the upland riparian zones. The study finds that an average of 3.67% of the monthly water budget of first-order catchments in central Texas is made up of spring flow. The influence of woody plant cover on streamflow was evaluated by comparing spring sites with different percentages of woody cover three times during 2003 and 2004. Our findings indicate that changes in woody plant cover had no influence on the amounts of streamflow from these catchments, and the surface catchment area had only a minor influence. This suggests that the real spring catchment area might be different from the surface watershed boundaries that have been delineated by topography. Plant species richness and diversity gradually decreased with increasing lateral distances from the stream bank. Herbaceous richness and diversity declined with increasing Ashe juniper cover in the riparian zone. Ashe juniper canopy cover had a larger effect on the understory composition than the cover of other woody species. Herbaceous diversity and production was greater in areas with sparse tree density than in areas with no trees, but was lowest at high tree densities. The complete removal of Ashe juniper in the riparian zones is not recommended because of the potential loss of grass cover. The recommended management would be to leave a sparse cover of canopy trees to maintain understory plants.

River regulation cause unnatural water flow patterns which disrupt succession, survival and dispersal of riverine plant communities. Riparian zones in regulated rivers are generally more species poor and have a lower vegetation cover compared to free-flowing rivers. Tributary mouths within the impoundment however, are not only affected by processes in the main channel, but also affected by processes such as sediment dynamics and flooding regime from the tributary which may more reflect a natural regime and may therefore not be affected by hydropeaking to the same extent. Hence, tributary sites and the accompanied tributary mouths may represent hotspots for high diversity plant communities in regulated rivers. The riparian habitat is unique in its interaction with adjacent ecological systems and is therefore considered important for the riverine ecosystem’s ecological functioning. The aim of the study was to evaluate the significance of tributary sites and accompanied tributary mouths as a hotspot for diverse plant communities in regulated rivers. Additionally, geomorphological features were analyzed to access the impact of hydropeaking in sheltered and non-sheltered tributary sites. Data was acquired from sampling sites within the Umeå and Luleå rivers, representing both tributary mouths and non-tributary mouth reaches. Inventories included presence of riparian and aquatic vascular plant species, vegetation cover and soil composition. The results indicate a less extensive impact of hydropeaking in tributary mouths compared to non-tributary mouth reaches. The tributary mouths had a higher species richness, diversity and vegetation cover when compared to that of the non-tributary mouth reaches. This supports the concept of tributary mouths being hotspots for plant diversity. Moreover, the sheltered tributary mouths had a higher species richness than the non-sheltered tributary sites, suggesting tributary shelter as a contributing counter of hydropeaking effects.

The purpose of the paper was to evaluate the effects of the restoration efforts in Juktån on the plant species richness and composition, vegetation zonation and soil composition. The restoration involved implementation of a dynamic flow, in addition to hydro-geomorphological alterations. Three sites were inventoried in the bypass reach, representing one reach affected only by flow restoration and two reaches affected both by flow and morphological restoration, along with an upstream reference reach unaffected by regulation. Inventories were conducted along an elevational gradient perpendicular to the water and involved occurring species, plant cover, bare ground and soil composition. The results showed a change in species composition after restoration. Riparian associated species were more common after restoration and the proportions of herbs and graminoids increased. Higher flooding caused a change in zonation, increasing the graminoid, riparian forest and amphibious zones along the riparian habitat. Species richness, as well as soil composition remained largely the same. The effects of the hydro-geomorphological alterations were hard to interpret due to lack of data. The results indicate a positive effect of the new flow on the plant species community in the bypass reach, whilst also highlighting the hydro-geomorphologically altered locations as biodiversity hotspots. Time-delay in ecological response is acknowledged as driving factor for the indifference in species richness and soil composition. The hydrological restoration is concluded to be an effective way of simulating natural flow regimes, improving ecological integrity of riparian communities and structural vegetational patterns in the riparian zone of bypass reaches.

No description available.

Thesis (PhD)--Stellenbosch University, 2013. / ENGLISH ABSTRACT: Riparian vegetation communities that occur along perennial rivers are structured in lateral zones that run parallel to river flow. This dissertation investigated the structure of South African riparian vegetation communities along perennial, single-thread headwater streams. The central assumption was that lateral zones result from differential species’ responses to changing abiotic factors along a lateral gradient up the river bank. It was first necessary to establish the pattern of zones and whether this pattern occurs repetitively and predictably on different rivers in different biomes. Since the flow regime is considered to be the master variable that controls the occurrence of lateral zones, the link between flow as the major abiotic driver and the distribution of plants in zones was determined. Predictions were made with respect to how variable flow may influence phenological traits, particularly with respect to seed dispersal, and physiological tolerances to drying out and were tested. The existence of lateral zones at reference sites in the Western Cape of South Africa was explored and their vegetation characteristics were described. Plant distribution was related to bank slope, as defined by elevation and distance from the wetted channel edge during summer (dry season) low flow, indicating a direct link to river bank hydraulics. Whether or not the same zonation patterns occur in riparian communities in other parts of South Africa was explored next. The four zones described for Fynbos Riparian Vegetation were evident at all of the other rivers tested, despite major differences in geographic location, vegetation community type, climate and patterns of seasonal flow. The four lateral zones could be separated from each other using a combination of flood recurrence and inundation duration. Functional differences were investigated between three tree species that occur in Fynbos Riparian Vegetation. Functional differences were apparent with respect to timing of seed dispersal, growth in branch length versus girth and three physiological measures of tolerance to drying out; specific leaf area (cm2.g-1), wood density (g.cm-3) and levels of carbon isotopes (δ13C). In order to determine the impact of invasive alien plants and to monitor recovery after clearing, the physical rules devised to help delineate zones were used to locate lateral zones that had been obliterated after invasion and subsequent clearing. At the sites invaded by A. mearnsii plants, the zone delineations showed that invasion started in the lower dynamic zone, where adult and sapling A. mearnsii were most abundant. In un-invaded systems, this zone was the least densely vegetated of the four zones, the most varied in terms of inundation duration and the frequency of inter- and intra-annual floods, and was an area of active recruitment comprised mainly of recruiting seedlings and saplings. An understanding of the functional differences between lateral zones was a common thread at each riparian community that was linked to the annual frequency of inundation and the period, when inundated. / AFRIKAANSE OPSOMMING: Oewer plantegroei gemeenskappe wat langs standhoudende riviere voorkom is gestruktureer in laterale sones parallel met die rivier vloei. Hierdie verhandeling ondersoek die struktuur van Suid-Afrikaanse oewer plantegroei gemeenskappe langs standhoudende, enkelloop hoof strome. Die sentrale aanname was dat laterale sones vorm as gevolg van verskillende spesies se reaksie teenoor die verandering van abiotiese faktore teen 'n laterale gradiënt met die rivierbank op. Dit was eers nodig om die patroon van die gebiede vas te stel en uit te vind of hierdie patroon herhaaldelik en voorspelbaar binne verskillende riviere in verskillende biome voorkom. Aangesien die vloeiwyse beskou word as die hoof veranderlike wat die teenwoordigheid van laterale sones beheer, is die skakel tussen die vloei, as die belangrikste abiotiese bestuurder, en die verspreiding van plante in sones bepaal. Voorspellings is gemaak met betrekking tot hoe veranderlike vloei fenologiese eienskappe kan beïnvloed, veral met betrekking tot die saad verspreiding, en fisiologiese toleransie teen uitdroog, en is getoets. Die bestaan van laterale sones binne verwysings studie terreine in die Wes-Kaap van Suid- Afrika is ondersoek en hul plantegroei eienskappe is beskryf. Plant verspreiding was verwant aan bank helling, soos gedefinieer deur hoogte en afstand vanaf die nat kanaal rand gedurende somer (droë seisoen) lae vloei, en dui dus op 'n direkte skakel met die rivier bank hidroulika. Of dieselfde sonering patrone voorkom in oewer gemeenskappe in ander dele van Suid-Afrika is volgende verken. Die vier sones beskryf vir fynbos oewer plantegroei was duidelik by al die ander riviere wat ondersoek is, ten spyte van groot verskille in geografiese ligging, plantegroei gemeenskap tipe, klimaat en patrone van seisoenale vloei. Die vier laterale sones kan onderskei word van mekaar deur middel van 'n kombinasie van vloed herhaling en oorstroomde toestand duur. Funksionele verskille is ondersoek tussen drie boom spesies wat voorkom in Fynbos Oewer Plantegroei. Funksionele verskille was duidelik met betrekking tot tydsberekening van saad verspreiding, groei in tak lengte tenoor omtrek, en drie fisiologiese maatstawwe van verdraagsaamheid teenoor uitdroging; spesifieke blaar area (cm2.g-1), hout digtheid (g.cm-3) en vlakke van koolstof isotope (δ13C). Ten einde die impak van indringerplante te bepaal en die herstel na ontbossing te monitor is die fisiese reëls voorheen vasgestel om sones te help baken gebruik om laterale sones, wat vernietig is na indringing en die daaropvolgende ontbossing, te vind. Op die terreine wat deur A. mearnsii indringerplante binnegeval is, het die indeling van sones getoon dat die indringing begin het in die laer dinamiese sone, waar volwasse en klein A. mearnsii bome die volopste was. In stelsels wat nie binnegeval is deur indringerplante was hierdie sone die minste dig begroei van die vier sones, die mees verskillend in terme van oorstroomde toestand duur en die frekwensie van inter-en intra-jaarlikse vloede, en was 'n gebied van aktiewe werwing hoofsaaklik bestaande uit rekruut saailinge en boompies. 'n Begrip van die funksionele verskille tussen laterale sones was 'n algemene verskynsel by elke oewer gemeenskap wat gekoppel was aan die jaarlikse frekwensie van oorstroming en die oorstroomde toestand duur.

Freshwater systems have, during human history, endured large-scale impacts. According to the water framework directive, measures must be developed to create a better environment for species in, and connected to, freshwater systems. Hydropower plants has caused loss of natural disturbance (e.g., floodings). Using already collected data from vegetation inventory in the riparian zone, probable distribution curves were created to find flooding requirements of different plant species, and linear regression analyses were run to see if hours of flooding and elevation above sea level had an effect on species richness and vegetation cover in the riparian zone. A model of Bredforsen 3 was conducted by using modelled values obtained from Vattenfall AB, with the aim to answer the following questions: How is riparian vegetation in mixed alluvial forests in reaches affected by static minimum flow levels structured according to flow dynamics? What would the zonation of riparian vegetation look like in an unregulated situation? How can minimum flow in Bredforsen be designed to better match the flooding regime similar to a natural riparian vegetation? Species richness showed to be dependent on elevation. Furthermore, significant results for species richness and vegetation cover both showed to be dependent on hours of flooding in the riparian zone for two of the three inventoried areas. In the alluvial forest, Picea abies, was not affected by hours of flooding. This might be caused by a low number of replicates available in Bredforsen of P. abies. In contrast, Quercus robur did show a significance towards hours of flooding and a vague negative trend for trees with high inundation distributed on lower elevations, though, it seems more parameters are affecting its distribution. The probable species distribution curves revealed the riparian zone in Bredforsen lack the clear vegetative zones visible along unregulated rivers. However, the model created for Bredforsen 3, following the assumptions of Ström et al. (2012) showed that today two vegetative zones, are apparent in Bredforsen 3 (amphibian zone and upland vegetation). With the modelled flow, a third zone (riparian forest) could appear. Because Bredforsen is a Natura 2000 reserve, the minimum flow needs alteration to mimic a relatively natural flow, which could lead to all vegetative zones to develop (amphibian zone, graminoids, willow shrubs, riparian forest, and upland vegetation). This could be achieved by using the spill water from Söderfors and time the release of spill to natural flow events. The model made from Bredforsen 3 indicates that this could cause positive changes in the riparian zone where a more natural distribution of vegetative zones is in place, meaning that species distribution returns to previous distribution patterns and would help disturbance dependent species.

Riparian vegetation is one of the most complex and abundant ecosystems in the world and it provides important ecosystem services. These services are affected by electricity production from hydropower dams. Hydropower accounts for 16% of the global electricity production and almost 50% in Sweden. One effect of hydropower is sub-daily fluctuations of water level caused by the turbines being turned on and off according to electricity demand. This is referred to as hydropeaking and has largely unknown effects on the fluvial ecosystem, and especially on the riparian vegetation. No studies have been made on the effects of hydropeaking on riparian vegetation. In this study, three native plants (Carex acuta, Betula pubescens and Salix phylicifolia x myrsinifolia) and one non-native plant (Helianthus annuus) were used as indicators (i.e., phytometers) for the effects of hydropeaking along two rivers from northern Sweden; one used for hydropower production and the other free flowing. From each of the four species, seedlings of two sizes and seeds were transplanted into five different river reaches and bank elevations along a hydropeaking gradient from none to high hydropeaking intensity. C. acuta and S. phylicifolia x myrsinifolia showed significant positive relationships to the hydropeaking gradient, likely due to their natural high tolerance to frequent inundation events. Therefore, they are suitable for restoration of river shores along reaches affected by hydropeaking. In contrast, B. pubescens was negatively related to the hydropeaking gradient, losing leaves and biomass with increasing hydropeaking intensities. It turned out to be the most sensitive species among the ones used in the experiment making it suitable as an indicator. H. annuus showed no response and therefore did not serve as impact indicator or for restoration. Germination for all native species was significantly lower along the reaches affected by hydropeaking which indicates a strong connection between hydropeaking and germination. These findings showed that recruitment becomes a bottleneck in riparian communities’ conservation along rivers affected by hydropeaking, and highlight the importance of mitigation actions focused on favoring riparian species seeds’ germination.