This thessis deals with assessment of the actual state of an herb layer and regeneration in nature preservations Coufavá and Slunná. Both of nature preservations were affected large-scale wind disturbances. The survey was conducted in June and July 2013. There were totally performed 384 phytosocenological releves (162 in nature preserve Coufavá and 222 in nature preserve Slunná). Research was carried out on a regular grid 15 × 15 meters. There were 254 juvenile trees which were founded in nature preserve Coufavá and also 1891 juvenile trees were founded in nature preserve Slunná. In both of preservations browse didn't rise up 10 % from all regeneration. Most abundant tree species in nature preserve Coufavá were Fagus sylvatica. In nature preserve Slunná were most abundant Tilia sp. Strong clonal species dominated in disturbed desktop non-wood undergrowth in both of studied areas. In closed stand dominated typical forest species. Impatiens parviflora has abundant occurrence in both of biotopes. For both nature preserves was estimated future development and propose of solutions. Result of work should serve as a basis for optimizing the management plan for the said reservation.

Summary: Floristic, Ecological and Phytosociological, Study of the Forests of the Mbiye Island at Kisangani, DR Congo The Mbiye Island is situated on the Congo River, in the Eastern part of Kisangani. It is located upstream of the Wagenia Falls, between latitude 0º31’ North and longitude 25º11’ East, with 376 m of altitude. It adjoins the town of Kisangani, and it is 14 km long and 4 km wide. All around Kisangani, the Mbiye Island is the only ecosystem which has a dense forest that is relatively well preserved. This Island has an area of 1,400 ha, and it comprises three types of forest: dry land forest, periodically flooded forest and swampy forest. The main purpose of this study has been to make a qualitative and quantitative analysis of the physiognomy and structure of these three types of forest. I relied on two methods in data collecting: the Phytosociological plotting method with a sampling equal to 100 individuals for the higher strata and 100 m² grass strata, associated with dbh measure for trees whose diameter is ≥ 10 cm. In total, out of the 80 plottings, 67 multi strata and structurals each of which has been decomposed in 4 sub-plottings fit together. Based on the presence-absence criterion, the other 13 multi strata and structurals have been made into Chablis, and all the data have been globally analyzed. An inventory of 33991 individuals has been made in this study. After their identification, it has been found out that there 470 species belonging to 297 types and 90 families. Rubiaceae, Caesalpiniaceae and euphorbiaceae families have been found out to be the most important in Genus and species. In the raw spectra of biological types, the phanerophyts (83.0%) are the most predominant, and within which the mesophanerophyts (trees 10 to 30 m tall in their old age) are the most predominant (39.0%). The Phytogeographical spectra have more omni-guineo-congolese (35%) elements; and amongst the diaspore types, the sarcochores (69%) are the most predominant. 58 endemic species (12.3%) of the central forest sector have been found out at the Mbiye Island. After the classification and ordination of the 80 plottings by Twinspan and DCA, three Phytosociological groupings have been found out in the dry land forest: (1) grouping of Funtumia elastica and Albizia ealaensis, (2) grouping of Nesogordonia leplaei and Antiaris toxicaria, (3) prouping of Olyra latifolia and Campylospermum elongatum; two others for floodable forests (1) grouping of Rinorea oblongifolia and Byrsocarpus coccineus, (2) grouping of Pachystela seretii, Dracaena kindtiana and Culcasia yangambiensis, and three others for swampy forests: (1) grouping of Strombosia grandifolia, Calamus deerratus, Culcasia angolensis and Elaeis guineensis, (2) grouping of Cleistopholis patens and Pterygota bequaertii, (3) grouping of Diospyros bipendensis, Strychnos icaja and Palisota barteri and then two groupings for the Chablis plottings (1) grouping of Palisota schweinfurthii, Eremospatha haullevilleana, Thomandersia hensii and Pycnocoma insularis and (2) grouping of Nephrolepis biserrata and Scaphopetalum thonneri. The coelocaryon botryoides has shown a uniform spatial distribution in the in the forest periodically flooded, whereas Gilbertiodendron dewevrei has shown an aggregated distribution. Comparison of richness in different taxa within the sampling has given high values as far as the numbers of species is concerned and genus of the same family in the phytosociological sampling. Out of all the species listed, 189 have been considered to be common species with regard to the three forest types, other 144 belonged to two types. 90 species have been listed only on solid land, 27 on periodically flooded forest 20 on swampy forest. Euphorbiaceae, Rubiaceae and Caesalpiniaceae families were the best represented in species and genus. With regard to richness in each forest type, the forest of solid soil was the only one to have high values both in species and genus, and the swampy forest was less represented. With reference to richness according to strata, the herbaceous and under-shrub stratum (H) was the richest whereas the stratum of dominant trees (E+A1) was the poorest. Mbiye island forest is the nearest one of Kisangani for which it plays the most important role with regards to supplies in different forest products and non-ligneous forest products. In total, 12.3% endemic species of Central forest sector have been listed there, but due to frequent use by man this time, all these endemic species may disappear from this island. It is therefore urgent that particular attention be paid on it. Actions of conservation and lasting exploitation must be started in order to block their extinction. Key words: DRCongo, Flora, Ecology, Phytosociology, insularity, flooding, spatial distribution, biodiversity, synusie, Mbiye Island, Kisangani./Résumé : Etude floristique, écologique et phytosociologique des forêts de l'île Mbiye, Kisangani, RDCongo. L'île Mbiye est une île du fleuve Congo située dans la partie Est de la ville de Kisangani, en amont des chutes Wagenia, entre 0° 31' de latitude Nord et 25° 11' de longitude Est, avec une altitude de 376 m. Elle jouxte la ville de Kisangani et sa longueur maximale est de 14 km sur 4 km de largeur. Elle est le seul écosystème à proximité de la ville, possédant encore une forêt dense relativement bien conservée et formant un îlot de 1400 ha comprenant trois types forestiers : la forêt de terre ferme, la forêt périodiquement inondée et la forêt marécageuse. L’objectif principal de cette étude a été de faire une analyse quantitative et qualitative de la physionomie et la structure de ces trois types forestiers. Deux méthodes ont permis la récolte de données ; la méthode de relevés phytosociologiques à effort d’échantillonnage égal à 100 individus pour les strates supérieures et 100 m² pour la strate herbacée, associée à celle de mesure de D130, pour les arbres à diamètre ≥ 10 cm. Au total, 80 relevés dont 67 multistrates et structurels, décomposés chacun en 4 sous-relevés emboîtés, ont été mis en place. 13 autres ont été faits dans les chablis avec l’appréciation basée sur le critère de présence-absence et toutes les données ont été analysées globalement. A l’issue de cette étude, 33991 individus ont été recensés. Après leur identification, au total 470 espèces appartenant à 297 genres et 90 familles ont été obtenues. Les familles des Rubiaceae, des Caesalpiniaceae et des Euphorbiaceae sont les plus importantes, tant en genres qu’en espèces. Dans les spectres bruts de types biologiques, les phanérophytes (83,0 %) sont les plus abondants et au sein desquels les mésophanérophytes (arbres de 10 à 30 m au stade adulte) sont les plus dominants (39,0 %). Les spectres phytogéographiques ont montré une prédominance d’éléments omni-guinéo-congolais (35 %) alors que parmi les types de diaspores, les sarcochores (69 %) sont les plus dominants. 58 espèces endémiques du Secteur forestier central (soit 12,3 %) ont été recensées à l’île Mbiye. Après classification et ordination de ces 80 relevés par Twinspan et DCA, trois groupements phytosociologiques ont été obtenus dans la forêt de terre ferme: (1) groupement à Funtumia elastica et Albizia ealaensis, (2) groupement à Nesogordonia leplaei et Antiaris toxicaria, (3) groupement à Olyra latifolia et Campylospermum elongatum ; deux autres pour les forêts inondables (1) groupement à Rinorea oblongifolia et Byrsocarpus coccineus, (2) groupement à Pachystela seretii, Dracaena kindtiana et Culcasia yangambiensis ; trois autres encore pour les forêts marécageuses : (1) groupement à Strombosia grandifolia, Calamus deerratus, Culcasia angolensis et Elaeis guineensis, (2) groupement à Cleistopholis patens et Pterygota bequaertii, (3) groupement à Diospyros bipendensis, Strychnos icaja et Palisota barteri, et enfin, deux groupements pour les relevés de chablis (1) groupement à Palisota schweinfurthii, Eremospatha haullevilleana, Thomandersia hensii et Pycnocoma insularum et (2) groupement à Nephrolepis biserrata et Scaphopetalum thonneri. L’espèce Coelocaryon botryoides a montré une répartition spatiale uniforme dans la forêt périodiquement inondée alors que Gilbertiodendron dewevrei y a montré une répartition agrégée. La comparaison de la richesse en différents taxa au sein de relevés et de placettes a donné de valeurs supérieures en nombre d’espèces, de genres et même de familles, dans les relevés phytociologiques. Sur le total des espèces recensées, 189 ont été considérées comme espèces communes aux trois types forestiers, 144 autres appartenaient aux deux types. 90 espèces n’ont été recensées qu’en forêt de terre ferme, 27 en forêt périodiquement inondée et 20 dans la forêt marécageuse. Les familles des Euphorbiaceae, des Rubiaceae et celle des Caesalpiniaceae ont été les mieux représentées en espèces et en genres. En ce qui concerne la richesse dans chaque type forestier, la forêt de terre ferme était la seule à avoir des valeurs supérieures tant en espèces qu’en genres et la forêt marécageuse était la moins représentée. Quant à la richesse spécifique selon les strates, la strate herbacée et sous-arbustive (H) était la plus riche alors que la strate des arbres dominants (E+A1) était la plus pauvre. La forêt de l’île Mbiye est la seule forêt proche de la ville de Kisangani pour laquelle elle joue un rôle de premier plan en ce qui concerne le ravitaillement en différents produits forestiers et forestiers non ligneux. Au total, 12,3% d’espèces endémiques du Secteur forestier Central y ont été recensées, mais vue l’action anthropique qu’elle connaît ce dernier temps, toutes ces espèces endémiques risquent de disparaître de cette île. Il est alors urgent qu’une attention particulière, soit tournée vers elle. Des actions de conservation et d’exploitation durable doivent être amorcées en vue contrecarrer l’extinction des celles-ci. Mots clés : RDCongo, flore, écologie, phytosociologie, insularité, inondation, répartition spatiale, biodiversité, synusie, île Mbiye, Kisangani.

Floristic, Ecological and Phytosociological, Study of the Forests of the Mbiye Island at Kisangani, DR Congo The Mbiye Island is situated on the Congo River, in the Eastern part of Kisangani. It is located upstream of the Wagenia Falls, between latitude 0º31’ North and longitude 25º11’ East, with 376 m of altitude. It adjoins the town of Kisangani, and it is 14 km long and 4 km wide. All around Kisangani, the Mbiye Island is the only ecosystem which has a dense forest that is relatively well preserved. This Island has an area of 1,400 ha, and it comprises three types of forest: dry land forest, periodically flooded forest and swampy forest. The main purpose of this study has been to make a qualitative and quantitative analysis of the physiognomy and structure of these three types of forest. I relied on two methods in data collecting: the Phytosociological plotting method with a sampling equal to 100 individuals for the higher strata and 100 m² grass strata, associated with dbh measure for trees whose diameter is ≥ 10 cm. In total, out of the 80 plottings, 67 multi strata and structurals each of which has been decomposed in 4 sub-plottings fit together. Based on the presence-absence criterion, the other 13 multi strata and structurals have been made into Chablis, and all the data have been globally analyzed. An inventory of 33991 individuals has been made in this study. After their identification, it has been found out that there 470 species belonging to 297 types and 90 families. Rubiaceae, Caesalpiniaceae and euphorbiaceae families have been found out to be the most important in Genus and species. In the raw spectra of biological types, the phanerophyts (83.0%) are the most predominant, and within which the mesophanerophyts (trees 10 to 30 m tall in their old age) are the most predominant (39.0%). The Phytogeographical spectra have more omni-guineo-congolese (35%) elements; and amongst the diaspore types, the sarcochores (69%) are the most predominant. 58 endemic species (12.3%) of the central forest sector have been found out at the Mbiye Island. After the classification and ordination of the 80 plottings by Twinspan and DCA, three Phytosociological groupings have been found out in the dry land forest: (1) grouping of Funtumia elastica and Albizia ealaensis, (2) grouping of Nesogordonia leplaei and Antiaris toxicaria, (3) prouping of Olyra latifolia and Campylospermum elongatum; two others for floodable forests (1) grouping of Rinorea oblongifolia and Byrsocarpus coccineus, (2) grouping of Pachystela seretii, Dracaena kindtiana and Culcasia yangambiensis, and three others for swampy forests: (1) grouping of Strombosia grandifolia, Calamus deerratus, Culcasia angolensis and Elaeis guineensis, (2) grouping of Cleistopholis patens and Pterygota bequaertii, (3) grouping of Diospyros bipendensis, Strychnos icaja and Palisota barteri and then two groupings for the Chablis plottings (1) grouping of Palisota schweinfurthii, Eremospatha haullevilleana, Thomandersia hensii and Pycnocoma insularis and (2) grouping of Nephrolepis biserrata and Scaphopetalum thonneri. The coelocaryon botryoides has shown a uniform spatial distribution in the in the forest periodically flooded, whereas Gilbertiodendron dewevrei has shown an aggregated distribution. Comparison of richness in different taxa within the sampling has given high values as far as the numbers of species is concerned and genus of the same family in the phytosociological sampling. Out of all the species listed, 189 have been considered to be common species with regard to the three forest types, other 144 belonged to two types. 90 species have been listed only on solid land, 27 on periodically flooded forest 20 on swampy forest. Euphorbiaceae, Rubiaceae and Caesalpiniaceae families were the best represented in species and genus. With regard to richness in each forest type, the forest of solid soil was the only one to have high values both in species<p>and genus, and the swampy forest was less represented. With reference to richness according to strata, the herbaceous and under-shrub stratum (H) was the richest whereas the stratum of dominant trees (E+A1) was the poorest. Mbiye island forest is the nearest one of Kisangani for which it plays the most important role with regards to supplies in different forest products and non-ligneous forest products. In total, 12.3% endemic species of Central forest sector have been listed there, but due to frequent use by man this time, all these endemic species may disappear from this island. It is therefore urgent that particular attention be paid on it. Actions of conservation and lasting exploitation must be started in order to block their extinction. Key words: DRCongo, Flora, Ecology, Phytosociology, insularity, flooding, spatial distribution, biodiversity, synusie, Mbiye Island, Kisangani./Etude floristique, écologique et phytosociologique des forêts de l'île Mbiye, Kisangani, RDCongo. L'île Mbiye est une île du fleuve Congo située dans la partie Est de la ville de Kisangani, en amont des chutes Wagenia, entre 0° 31' de latitude Nord et 25° 11' de longitude Est, avec une altitude de 376 m. Elle jouxte la ville de Kisangani et sa longueur maximale est de 14 km sur 4 km de largeur. Elle est le seul écosystème à proximité de la ville, possédant encore une forêt dense relativement bien conservée et formant un îlot de 1400 ha comprenant trois types forestiers :la forêt de terre ferme, la forêt périodiquement inondée et la forêt marécageuse. L’objectif principal de cette étude a été de faire une analyse quantitative et qualitative de la physionomie et la structure de ces trois types forestiers. Deux méthodes ont permis la récolte de données ;la méthode de relevés phytosociologiques à effort d’échantillonnage égal à 100 individus pour les strates supérieures et 100 m² pour la strate herbacée, associée à celle de mesure de D130, pour les arbres à diamètre ≥ 10 cm. Au total, 80 relevés dont 67 multistrates et structurels, décomposés chacun en 4 sous-relevés emboîtés, ont été mis en place. 13 autres ont été faits dans les chablis avec l’appréciation basée sur le critère de présence-absence et toutes les données ont été analysées globalement. A l’issue de cette étude, 33991 individus ont été recensés. Après leur identification, au total 470 espèces appartenant à 297 genres et 90 familles ont été obtenues. Les familles des Rubiaceae, des Caesalpiniaceae et des Euphorbiaceae sont les plus importantes, tant en genres qu’en espèces. Dans les spectres bruts de types biologiques, les phanérophytes (83,0 %) sont les plus abondants et au sein desquels les mésophanérophytes (arbres de 10 à 30 m au stade adulte) sont les plus dominants (39,0 %). Les spectres phytogéographiques ont montré une prédominance d’éléments omni-guinéo-congolais (35 %) alors que parmi les types de diaspores, les sarcochores (69 %) sont les plus dominants. 58 espèces endémiques du Secteur forestier central (soit 12,3 %) ont été recensées à l’île Mbiye. Après classification et ordination de ces 80 relevés par Twinspan et DCA, trois groupements phytosociologiques ont été obtenus dans la forêt de terre ferme: (1) groupement à Funtumia elastica et Albizia ealaensis, (2) groupement à Nesogordonia leplaei et Antiaris toxicaria, (3) groupement à Olyra latifolia et Campylospermum elongatum ;deux autres pour les forêts inondables (1) groupement à Rinorea oblongifolia et Byrsocarpus coccineus, (2) groupement à Pachystela seretii, Dracaena kindtiana et Culcasia yangambiensis ;trois autres encore pour les forêts marécageuses :(1) groupement à Strombosia grandifolia, Calamus deerratus, Culcasia angolensis et Elaeis guineensis, (2) groupement à Cleistopholis patens et Pterygota bequaertii, (3) groupement à Diospyros bipendensis, Strychnos icaja et Palisota barteri, et enfin, deux groupements pour les relevés de chablis (1) groupement à Palisota schweinfurthii, Eremospatha haullevilleana, Thomandersia hensii et Pycnocoma insularum et (2) groupement à Nephrolepis biserrata et Scaphopetalum thonneri. L’espèce Coelocaryon botryoides a montré une répartition spatiale uniforme dans la forêt périodiquement inondée alors que Gilbertiodendron dewevrei y a montré une répartition agrégée. La comparaison de la richesse en différents taxa au sein de relevés et de placettes a donné de valeurs supérieures en nombre d’espèces, de genres et même de familles, dans les relevés phytociologiques. Sur le total des espèces recensées, 189 ont été considérées comme espèces communes aux trois types forestiers, 144 autres appartenaient aux deux types. 90 espèces<p>n’ont été recensées qu’en forêt de terre ferme, 27 en forêt périodiquement inondée et 20 dans la forêt marécageuse. Les familles des Euphorbiaceae, des Rubiaceae et celle des Caesalpiniaceae ont été les mieux représentées en espèces et en genres. En ce qui concerne la richesse dans chaque type forestier, la forêt de terre ferme était la seule à avoir des valeurs supérieures tant en espèces qu’en genres et la forêt marécageuse était la moins représentée. Quant à la richesse spécifique selon les strates, la strate herbacée et sous-arbustive (H) était la plus riche alors que la strate des arbres dominants (E+A1) était la plus pauvre. La forêt de l’île Mbiye est la seule forêt proche de la ville de Kisangani pour laquelle elle joue un rôle de premier plan en ce qui concerne le ravitaillement en différents produits forestiers et forestiers non ligneux. Au total, 12,3% d’espèces endémiques du Secteur forestier Central y ont été recensées, mais vue l’action anthropique qu’elle connaît ce dernier temps, toutes ces espèces endémiques risquent de disparaître de cette île. Il est alors urgent qu’une attention particulière, soit tournée vers elle. Des actions de conservation et d’exploitation durable doivent être amorcées en vue contrecarrer l’extinction des celles-ci. Mots clés :RDCongo, flore, écologie, phytosociologie, insularité, inondation, répartition spatiale, biodiversité, synusie, île Mbiye, Kisangani. / Doctorat en Sciences / info:eu-repo/semantics/nonPublished

I. An original methodological discussion is proposed on the problem of the typology of tropical rain forest’s plant communities, based on the study of forest types across gradients of continentality and elevation, within Atlantic central Africa. These investigations were based on the statement that the main problems in forest typology are related to the non-zonal or zonal character of the different vegetation types and to non considering the relations and differences between forest strata. II. Field data consisted in phytosociological homogeneous sample plots localized within different recognized phytogeographical entities, in a region of tropical Africa where these entities are known to be well conserved. A total of 37 such plots were inventoried in the region extending from the littoral forests of Ndoté, Equatorial Guinea, which are wet evergreen forests, to the continental forests of the Dja, Cameroon, known as evergreen seasonal forests. The studied region also included the oriental Atlantic forests of Equatorial Guinea, known as moist evergreen forests or caesalp forests. In various parts of this continentality gradient, some plots were localized within climax non-zonal formations, namely the submontane rain forests. The emphasis was put on the vegetation of the Monte Alén National Park. The sampling methodology was willing to be as "complete ", including all strata, "quantitative ", enumerating all individuals, and "representative ", within each stratum, as possible. These multi-layers plots were realised using nested sub-plots, with a sampling size of 100 individuals for every ligneous stratum recognized (dominant trees, dominated trees and shrubs) and a sampling size of 200m² for the herbaceous and suffrutex stratum. Forest types were defined independently for each stratum and the differences were analysed. A method was proposed for the simultaneous analysis of all floristic data, converting and standardizing the values from ligneous strata, on the one hand, and from understorey strata, on the other hand. III. Ten forest types were described using IndVal and discussed in the general context of the guineo-congolian region, from a syntaxonomic view point (agglomerative classification) and from a phytogeographical view point (divisive classification). Homologies between these two approaches are described. The proposed phytogeographical system is based on an "open " conception of hierarchical classifications, combining advantages of agglomerative and divisive classifications. In concrete terms, the non-zonal criteria, for example the submontane variants, are categorised separately and in analogy with the zonal criteria, related to the usual phytochoria. Analysis of ecological relationships for the 10 communities showed that the main variables related to the floristic variability in our mainland rain forests are elevation, rainfall, hygrometry (estimated using bryophytes cover levels) and distance to the ocean. The two extremes on the vertical microclimatic gradient, dominant trees stratum and herbaceous stratum, give similar typologies, however canonical analysis showed that for the herbaceous layer, non-zonal variables (hygrometry and elevation) were gaining more importance when the influence of the two zonal variables was attenuated. In every case, spatial autocorrelation was less important than the environment in explaining floristic variability but its role increased in the spatial arrangement of understorey species, whose dispersal capacity is generally lower than canopy trees. The phytosociological, phytogeographical and ecological description of forest types is accompanied by a physiognomical description using biological types spectrum, as well as architectural models, leaf sizes, etc. With regard to diversity, we have demonstrated that species richness was higher from upper to lower strata because of the accumulation in lower strata of species from various strata. On the other hand, the proper stratum diversity, i.e. the structural set, decreased from dominant trees to shrubs. The proper diversity of the herb layer showed relatively high figures mainly due to the higher individual density in relation to the existence of microstrata. Within the 37 sample plots, 1,050 taxa have been identified to species or morpho-species levels, for a total of 25,750 individuals. These taxa represent 442 genus among 104 families. The richest forest type is found on the foothills of the Niefang range, on the windward side. This forest type is also characterised by a high number of oligotypic genus and by species belonging to functional types indicators of glacial refuges. These functional types are defined on the basis of the dispersal capacity and on kind of stand needed for effective germination. We formulated the hypothesis that this kind of "foothills refuge ", characterised by his zonal nature, could have been one of the rare refuges for species from mainland rain forests, while montane and fluvial refuges would mainly have preserved species from non-zonal forest types: (sub)montane and riverine. Based on indicator species of submontane forests, a potential distribution map of this forest type has been realised at the Atlantic central African scale. More than 400 submontane forest localities have been mapped. These forests begin at 400m of altitude near the ocean, and progressively at higher altitude for increasing distance to the ocean. Many lowland localities also comprised submontane species, which could indicate the existence of ecological transgressions. These transgressions would allow migratory tracks for submontane species between isolated mountain ranges, not only during glacial periods, through heights at the northern and southern borders of the congo basin, but also contemporarily through the lowland riverine forest network, in the centre of this basin. Finally, a special attention has been attributed to littoral forests and to some cases of choroecological transgressions, coupled to the ecological equalization phenomenon.

I. An original methodological discussion is proposed on the problem of the typology of tropical rain forest’s plant communities, based on the study of forest types across gradients of continentality and elevation, within Atlantic central Africa. These investigations were based on the statement that the main problems in forest typology are related to the non-zonal or zonal character of the different vegetation types and to non considering the relations and differences between forest strata.<p><p>II. Field data consisted in phytosociological homogeneous sample plots localized within different recognized phytogeographical entities, in a region of tropical Africa where these entities are known to be well conserved. A total of 37 such plots were inventoried in the region extending from the littoral forests of Ndoté, Equatorial Guinea, which are wet evergreen forests, to the continental forests of the Dja, Cameroon, known as evergreen seasonal forests. The studied region also included the oriental Atlantic forests of Equatorial Guinea, known as moist evergreen forests or caesalp forests. In various parts of this continentality gradient, some plots were localized within climax non-zonal formations, namely the submontane rain forests. The emphasis was put on the vegetation of the Monte Alén National Park.<p><p>The sampling methodology was willing to be as "complete ", including all strata, "quantitative ", enumerating all individuals, and "representative ", within each stratum, as possible. These multi-layers plots were realised using nested sub-plots, with a sampling size of 100 individuals for every ligneous stratum recognized (dominant trees, dominated trees and shrubs) and a sampling size of 200m² for the herbaceous and suffrutex stratum.<p><p>Forest types were defined independently for each stratum and the differences were analysed. A method was proposed for the simultaneous analysis of all floristic data, converting and standardizing the values from ligneous strata, on the one hand, and from understorey strata, on the other hand.<p><p>III. Ten forest types were described using IndVal and discussed in the general context of the guineo-congolian region, from a syntaxonomic view point (agglomerative classification) and from a phytogeographical view point (divisive classification). Homologies between these two approaches are described. The proposed phytogeographical system is based on an "open " conception of hierarchical classifications, combining advantages of agglomerative and divisive classifications. In concrete terms, the non-zonal criteria, for example the submontane variants, are categorised separately and in analogy with the zonal criteria, related to the usual phytochoria.<p><p>Analysis of ecological relationships for the 10 communities showed that the main variables related to the floristic variability in our mainland rain forests are elevation, rainfall, hygrometry (estimated using bryophytes cover levels) and distance to the ocean. The two extremes on the vertical microclimatic gradient, dominant trees stratum and herbaceous stratum, give similar typologies, however canonical analysis showed that for the herbaceous layer, non-zonal variables (hygrometry and elevation) were gaining more importance when the influence of the two zonal variables was attenuated. In every case, spatial autocorrelation was less important than the environment in explaining floristic variability but its role increased in the spatial arrangement of understorey species, whose dispersal capacity is generally lower than canopy trees. The phytosociological, phytogeographical and ecological description of forest types is accompanied by a physiognomical description using biological types spectrum, as well as architectural models, leaf sizes, etc.<p><p>With regard to diversity, we have demonstrated that species richness was higher from upper to lower strata because of the accumulation in lower strata of species from various strata. On the other hand, the proper stratum diversity, i.e. the structural set, decreased from dominant trees to shrubs. The proper diversity of the herb layer showed relatively high figures mainly due to the higher individual density in relation to the existence of microstrata. Within the 37 sample plots, 1,050 taxa have been identified to species or morpho-species levels, for a total of 25,750 individuals. These taxa represent 442 genus among 104 families. The richest forest type is found on the foothills of the Niefang range, on the windward side. This forest type is also characterised by a high number of oligotypic genus and by species belonging to functional types indicators of glacial refuges. These functional types are defined on the basis of the dispersal capacity and on kind of stand needed for effective germination. We formulated the hypothesis that this kind of "foothills refuge ", characterised by his zonal nature, could have been one of the rare refuges for species from mainland rain forests, while montane and fluvial refuges would mainly have preserved species from non-zonal forest types: (sub)montane and riverine.<p><p>Based on indicator species of submontane forests, a potential distribution map of this forest type has been realised at the Atlantic central African scale. More than 400 submontane forest localities have been mapped. These forests begin at 400m of altitude near the ocean, and progressively at higher altitude for increasing distance to the ocean. Many lowland localities also comprised submontane species, which could indicate the existence of ecological transgressions. These transgressions would allow migratory tracks for submontane species between isolated mountain ranges, not only during glacial periods, through heights at the northern and southern borders of the congo basin, but also contemporarily through the lowland riverine forest network, in the centre of this basin. Finally, a special attention has been attributed to littoral forests and to some cases of choroecological transgressions, coupled to the ecological equalization phenomenon.<p> / Doctorat en sciences agronomiques et ingénierie biologique / info:eu-repo/semantics/nonPublished