Former researches provide evidence that invasive species could alter ecosystem’s components, threaten native species and cause economic losses in southern forest lands. The objective of the project is to explore significant driving factors and develop geospatial models for monitoring, predicting and mapping the extent and conditions of major invasive species. In the study area, 16 invasive species were classified into four groups: regionally spreading species, regionally establishing species, locally spreading species and regionally colonizing species by population size and spatial characteristics. According to local Moran’s I, spatial autocorrelation existed in 16 invasive species. Autologistic model and simultaneous autoregressive model were employed to explore the relationships between spatial distribution and a set of indentified variables for Chinese privet, kudzu, Nepalese browntop and tallow tree at plot and county levels. The project showed that human-caused disturbances and forest types were significantly related to the spatial distribution of four invasive species in different scales.

La cartographie du couvert végétal est un outil essentiel au suivi et à la gestion et des milieux « naturels ». Des cartes caractérisant les essences forestières à l'échelle régionale sont nécessaires pour la gestion des milieux forestiers. Les séries temporelles d'images satellitaires optiques à moyenne résolution spatiale, peuvent permettre de satisfaire ce besoin. L'objectif de cette thèse est d'améliorer la classification supervisée d'une série temporelle afin de produire des cartes à l'échelle régionale détaillant la composition en essences de la végétation forestière. Nous avons d'abord évalué l'apport de la stratification du site d'étude pour améliorer les résultats de la classification d'une série temporelle d'images MODIS. Le recours à une stratification à partir d'une segmentation orientée objet améliore la classification supervisée, avec une augmentation de la valeur de Kappa et du taux de rejet des pixels à classer. Un seuil minimal et un seuil maximal de la surface de végétation à classer ont été identifiés, correspondant respectivement à un taux de rejet trop élevé et à une absence d'effet de la stratification. Nous avons ensuite évalué l'influence de l'organisation de la série temporelle d'images à moyenne résolution spatiale et du choix de l'algorithme de classification. Cette évaluation a été effectuée pour trois algorithmes (maximum de vraisemblance, Support Vector Machine, Random Forest) en faisant varier les caractéristiques de la série temporelle. On observe un effet de la temporalité et de la radiométrie sur la précision de la classification particulièrement significatif et la supériorité de l'algorithme Random Forest. Sur le plan thématique, des confusions subsistent et certains mélanges d'essences sont mal distingués. Nous avons alors cherché à évaluer l'apport du changement de résolution spatiale des images composant la série temporelle pour améliorer les résultats de classification. Les conclusions effectuées précédemment avec les données MODIS sont confortées, ce qui permet de conclure qu'elles sont indépendantes des données d'entrée et de leur résolution spatiale. Une amélioration significative est apportée par le changement de résolution spatiale, avec une augmentation de l'indice de Kappa de 0,60 à 0,72 obtenue grâce à la diminution de la proportion de pixels mixtes. Quelle que soit la résolution spatiale des images utilisées, les résultats obtenus montrent que la définition d'une procédure optimale améliore sensiblement les résultats de la classification. / Forest cover mapping is an essential tool for forest management. Detailed maps, characterizing forest types at a régional scale, are needed. This need can be fulfilled by médium spatial resolution optical satellite images time sériés. This thesis aims at improving the supervised classification procédure applied to a time sériés, to produce maps detailing forest types at a régional scale. To meet this goal, the improvement of the results obtained by the classification of a MODIS time sériés, performed with a stratification of the study area, was assessed. An improvement of classification accuracy due to stratification built by object-based image analysis was observed, with an increase of the Kappa index value and an increase of the reject fraction rate. These two phenomena are correlated to the classified végétation area. A minimal and a maximal value were identified, respectively related to a too high reject fraction rate and a neutral stratification impact.We carried out a second study, aiming at assessing the influence of the médium spatial resolution time sériés organization and of the algorithm on classification quality. Three distinct classification algorithms (maximum likelihood, Support Vector Machine, Random Forest) and several time sériés were studied. A significant improvement due to temporal and radiométrie effects and the superiority of Random Forest were highlighted by the results. Thematic confusions and low user's and producer's accuracies were still observed for several classes. We finally studied the improvement brought by a spatial resolution change for the images composing the time sériés to discriminate classes of mixed forest species. The conclusions of the former study (MODIS images) were confirmed with DEIMOS images. We can conclude that these effects are independent from input data and their spatial resolution. A significant improvement was also observed with an increase of the Kappa index value from 0,60 with MODIS data to 0,72 with DEIMOS data, due to a decrease of the mixed pixels rate.

- Ziele der Arbeit - Ziel vorliegender Arbeit ist, zu analysieren und zu beurteilen, in welchem Umfang autogene Moorregeneration nach anthropogenen Störungen auftritt, welche Voraussetzungen sie erfordert und welche Prozesse von Bedeutung sind. Forstliche und naturschutzfachliche Relevanz der Erkenntnisse sind zu prüfen. - Versuchsanlage und Methoden - Die Versuchsanlage umfasst verschiedene räumliche Skalenebenen und Zeitabschnitte. Im Untersuchungsraum sächsisches Erzgebirge befinden sich fünf Untersuchungsgebiete. Geschichtliche Aspekte werden auf Basis von Recherchen, der aktuelle Moorzustand anhand von Geländeerhebungen zu Vegetation und Standort analysiert. Langzeitbeobachtungen dienen dem Nachweis von Sukzessionsprozessen und den sie beeinflussenden Umweltfaktoren. Regenerationsprozesse werden anhand von Regenerationsmerkmalen, die eigens für diese Zwecke erarbeitet wurden, lokalisiert und im Kontext mit dem aktuellen Moorzustand und den Erfordernissen von Naturschutz und Forstwirtschaft interpretiert. - Ergebnisse und Schlussfolgerungen - - Das Aufnahmematerial lässt die Differenzierung von 28 Vegetationstypen zu, die anhand von Artengruppen und Zeigerwerten charakterisiert und als Kartiereinheiten zur Bewertung der Moore verwendet werden. Die Moorfläche im Untersuchungsraum wird auf 8.500 ha geschätzt. Jedes untersuchte Moor weist eine spezifische Kombination abiotischer sowie biotischer Merkmale auf und ist deshalb individuell zu bewerten. Entwässerungen und Torfstiche führen zu starker Degeneration, Dominanz von Wald, Seltenheit moortypischer Pflanzenarten, Offengesellschaften, nässegeprägter hydromorphologischer Strukturen sowie ökosystemrelevanter Schlüsselarten. Hinzu kommt eine Schädigung der Baum- und Moosschicht durch SO2-Immissionen bis in die 1990er Jahre und eine nachfolgende Regeneration. - Wiedervernässungen durch Graben- und Torfstichverlandung sind anhand von Zeitreihen und aktuellen Regenerationsmerkmalen nachweisbar. Autogene Moorregeneration kommt selbst in stark degenerierten Mooren vor. Regenerationsbereiche sind häufig, erreichen aber nur einen geringen Anteil an der Moorfläche (Untersuchungsraum: 1 %, Untersuchungsgebiete: 12 %). Sie haben meist mesotrophen Charakter. Eine erhöhte Regenerationsdisposition tritt u. a. bei konvergenten Wasserströmen und geringen Neigungen auf. Lokale Prozesse wie die Bildung von Fließhindernissen beeinflussen das Unwirksamwerden von Gräben. Die Etablierung torfbildender Vegetation nimmt eine Schlüsselstellung ein. Verschiedene Stufen eines diskontinuierlichen Regenerationsfortschritts sind zu unterscheiden und in bisher bekannte Prozessabläufe einzuordnen. Häufigkeit und Flächenverhältnis initialer und fortgeschrittener Regeration legen nahe, dass Regenerationsprozesse auf größerer Fläche ablaufen, jedoch nicht erkannt werden. Regeneration führt zu Standortsdrift und Ausbreitung moortypischer Arten. Das Regenerationspotenzial ist abiotisch vorgegeben. Irreversible Veränderungen der hydromorphologischen Struktur durch Entwässerung oder Torfabbau setzen der Regeneration Grenzen. Anthropogene Stoffeinträge und Mangel an Schlüsselarten bewirken weitere Einschränkungen. - Soweit eine rentable Holzproduktion Ziel ist, stellen autogen regenerierende Moore schwer bis nicht bewirtschaftbare Standorte dar, die aufgrund von Standortsdrift und teils Gehölzfeindlichkeit ein hohes Produktionsrisiko bergen. Naturschutzfachlich sind Regenerationsprozesse eine Chance und ein bedeutendes Schutzgut, da sie zu höherwertigeren Moorlebensräumen führen. Bei fortgeschrittener Regeneration erübrigen sich kostenintensive Eingriffe. Ein statischer Schutz wird dem Prozesscharakter nicht gerecht. Initialstadien und Bereiche, die noch nicht regenerieren, aber ein hohes Potenzial aufweisen, lassen sich effizient fördern. Die starke Degeneration der Moore im Untersuchungsraum begründet bei geringem Anteil aktueller Regenerationsbereiche einen hohen Handlungsbedarf, einschließlich eines abschirmenden Schutzes sowie effizienter Kontrollmechanismen. - Geeignete Datengrundlagen zur Lokalisierung von Flächen mit einem hohen Regenerationspotenzial fehlen weitgehend. Eine Behebung dieses Defizits ist möglich. / - Objectives - The objective of this work is to analyse and to assess to which extent autogenous peatland regeneration occurs after anthropogenic disturbances, what preconditions it requires and what processes are important. Relevance of the findings for forestry and nature conservation is to be examined. - Experimental set-up and methods - The experimental set-up comprises various spatial scale levels and time periods. Five study sites are located in the study area Saxon Erzgebirge Mountains. Historical aspects are analysed based on investigations, the actual condition of the peatland is determined by ground surveys focusing on the vegetation and the site. Long-term observations serve to prove succession processes and the environmental factors influencing them. Regeneration processes are localized by means of regeneration characteristics, which were elaborated just for this purpose, localized, and interpreted in the context of the actual peatland condition and of the requirements of nature conservation and forestry. - Results and conclusions - - The material under survey allows distinguishing between 28 vegetation types which are characterised by means of species groups and indicator values and which are used as mapping units for evaluating the peatlands. The peatland area in the study area comprises an estimated 8500 ha. Each investigated peatland has a specific combination of abiotic and biotic characteristics and therefore needs to be evaluated individually. Drainage and peat cuttings lead to strong degeneration, dominance of forest, rarity of plants species typical of peatland, open-land associations, wetness-coined hydromorphological structures as well as ecosystem-relevant key species. In addition, the tree and moss layer had been damaged by SO2-pollution up to the 1990ies and a succeeding regeneration. - Recurring wetting due to filling-up of ditches and peat cuttings by sedimentation are verifiable by time sequences and actual regeneration characteristics. Autogenous peatland regeneration occurs even in heavily degenerated peatlands. Regeneration areas are frequent, accounting, however, only for a small share at the peatland area (study area: 1 %, study sites: 12 %). Often they are of mesotrophic nature. An increased disposition to regeneration is found, among others, in case of convergent water flows and slight inclinations. Local processes like the formation of obstacles for the flowing may cause ditches to become ineffective. Establishing of peat-forming vegetation is a crucial phenomenon. Various stages of a discontinuous regeneration progress can be differentiated and allocated to the processes that are known so far. Frequency and the area ratio of initial and progressed regeneration suggest that regeneration processes take place on a larger area, but that they are not recognised. Regeneration leads to site drifting and the distribution of peatland species. The regeneration potential is given by abiotic conditions. Irreversible changes of the hydro-morphological structure due to drainage or peat cutting are limiting factors for the regeneration. Anthropogenic input of matter and lack of key species bring about other restrictions. - If a profitable timber production is aimed at, autogenously regenerated peatlands are sites difficult to manage or that cannot be managed at all, involving a high production risk due to site drifting and partly inadequacy for woody plants. For nature conservation, regeneration processes are a chance and an important asset worth of protection, as they lead to high-grade peatland habitats. In the case of advanced regeneration cost-intensive operations are not necessary. A protection of static nature is inadequate for the process character. Initial stages and zones not yet under regeneration, implying, however, a high potential, can be promoted efficiently. Heavy degeneration of the peatlands in the study area, given a low proportion of actual regeneration zones, justifies a strong call for action, including a shielding protection as well as efficient control mechanisms. - An appropriate data basis for localisation of areas, having a high regeneration potential, is largely missing. Remedying this deficit is possible.

- Ziele der Arbeit - Ziel vorliegender Arbeit ist, zu analysieren und zu beurteilen, in welchem Umfang autogene Moorregeneration nach anthropogenen Störungen auftritt, welche Voraussetzungen sie erfordert und welche Prozesse von Bedeutung sind. Forstliche und naturschutzfachliche Relevanz der Erkenntnisse sind zu prüfen. - Versuchsanlage und Methoden - Die Versuchsanlage umfasst verschiedene räumliche Skalenebenen und Zeitabschnitte. Im Untersuchungsraum sächsisches Erzgebirge befinden sich fünf Untersuchungsgebiete. Geschichtliche Aspekte werden auf Basis von Recherchen, der aktuelle Moorzustand anhand von Geländeerhebungen zu Vegetation und Standort analysiert. Langzeitbeobachtungen dienen dem Nachweis von Sukzessionsprozessen und den sie beeinflussenden Umweltfaktoren. Regenerationsprozesse werden anhand von Regenerationsmerkmalen, die eigens für diese Zwecke erarbeitet wurden, lokalisiert und im Kontext mit dem aktuellen Moorzustand und den Erfordernissen von Naturschutz und Forstwirtschaft interpretiert. - Ergebnisse und Schlussfolgerungen - - Das Aufnahmematerial lässt die Differenzierung von 28 Vegetationstypen zu, die anhand von Artengruppen und Zeigerwerten charakterisiert und als Kartiereinheiten zur Bewertung der Moore verwendet werden. Die Moorfläche im Untersuchungsraum wird auf 8.500 ha geschätzt. Jedes untersuchte Moor weist eine spezifische Kombination abiotischer sowie biotischer Merkmale auf und ist deshalb individuell zu bewerten. Entwässerungen und Torfstiche führen zu starker Degeneration, Dominanz von Wald, Seltenheit moortypischer Pflanzenarten, Offengesellschaften, nässegeprägter hydromorphologischer Strukturen sowie ökosystemrelevanter Schlüsselarten. Hinzu kommt eine Schädigung der Baum- und Moosschicht durch SO2-Immissionen bis in die 1990er Jahre und eine nachfolgende Regeneration. - Wiedervernässungen durch Graben- und Torfstichverlandung sind anhand von Zeitreihen und aktuellen Regenerationsmerkmalen nachweisbar. Autogene Moorregeneration kommt selbst in stark degenerierten Mooren vor. Regenerationsbereiche sind häufig, erreichen aber nur einen geringen Anteil an der Moorfläche (Untersuchungsraum: 1 %, Untersuchungsgebiete: 12 %). Sie haben meist mesotrophen Charakter. Eine erhöhte Regenerationsdisposition tritt u. a. bei konvergenten Wasserströmen und geringen Neigungen auf. Lokale Prozesse wie die Bildung von Fließhindernissen beeinflussen das Unwirksamwerden von Gräben. Die Etablierung torfbildender Vegetation nimmt eine Schlüsselstellung ein. Verschiedene Stufen eines diskontinuierlichen Regenerationsfortschritts sind zu unterscheiden und in bisher bekannte Prozessabläufe einzuordnen. Häufigkeit und Flächenverhältnis initialer und fortgeschrittener Regeration legen nahe, dass Regenerationsprozesse auf größerer Fläche ablaufen, jedoch nicht erkannt werden. Regeneration führt zu Standortsdrift und Ausbreitung moortypischer Arten. Das Regenerationspotenzial ist abiotisch vorgegeben. Irreversible Veränderungen der hydromorphologischen Struktur durch Entwässerung oder Torfabbau setzen der Regeneration Grenzen. Anthropogene Stoffeinträge und Mangel an Schlüsselarten bewirken weitere Einschränkungen. - Soweit eine rentable Holzproduktion Ziel ist, stellen autogen regenerierende Moore schwer bis nicht bewirtschaftbare Standorte dar, die aufgrund von Standortsdrift und teils Gehölzfeindlichkeit ein hohes Produktionsrisiko bergen. Naturschutzfachlich sind Regenerationsprozesse eine Chance und ein bedeutendes Schutzgut, da sie zu höherwertigeren Moorlebensräumen führen. Bei fortgeschrittener Regeneration erübrigen sich kostenintensive Eingriffe. Ein statischer Schutz wird dem Prozesscharakter nicht gerecht. Initialstadien und Bereiche, die noch nicht regenerieren, aber ein hohes Potenzial aufweisen, lassen sich effizient fördern. Die starke Degeneration der Moore im Untersuchungsraum begründet bei geringem Anteil aktueller Regenerationsbereiche einen hohen Handlungsbedarf, einschließlich eines abschirmenden Schutzes sowie effizienter Kontrollmechanismen. - Geeignete Datengrundlagen zur Lokalisierung von Flächen mit einem hohen Regenerationspotenzial fehlen weitgehend. Eine Behebung dieses Defizits ist möglich. / - Objectives - The objective of this work is to analyse and to assess to which extent autogenous peatland regeneration occurs after anthropogenic disturbances, what preconditions it requires and what processes are important. Relevance of the findings for forestry and nature conservation is to be examined. - Experimental set-up and methods - The experimental set-up comprises various spatial scale levels and time periods. Five study sites are located in the study area Saxon Erzgebirge Mountains. Historical aspects are analysed based on investigations, the actual condition of the peatland is determined by ground surveys focusing on the vegetation and the site. Long-term observations serve to prove succession processes and the environmental factors influencing them. Regeneration processes are localized by means of regeneration characteristics, which were elaborated just for this purpose, localized, and interpreted in the context of the actual peatland condition and of the requirements of nature conservation and forestry. - Results and conclusions - - The material under survey allows distinguishing between 28 vegetation types which are characterised by means of species groups and indicator values and which are used as mapping units for evaluating the peatlands. The peatland area in the study area comprises an estimated 8500 ha. Each investigated peatland has a specific combination of abiotic and biotic characteristics and therefore needs to be evaluated individually. Drainage and peat cuttings lead to strong degeneration, dominance of forest, rarity of plants species typical of peatland, open-land associations, wetness-coined hydromorphological structures as well as ecosystem-relevant key species. In addition, the tree and moss layer had been damaged by SO2-pollution up to the 1990ies and a succeeding regeneration. - Recurring wetting due to filling-up of ditches and peat cuttings by sedimentation are verifiable by time sequences and actual regeneration characteristics. Autogenous peatland regeneration occurs even in heavily degenerated peatlands. Regeneration areas are frequent, accounting, however, only for a small share at the peatland area (study area: 1 %, study sites: 12 %). Often they are of mesotrophic nature. An increased disposition to regeneration is found, among others, in case of convergent water flows and slight inclinations. Local processes like the formation of obstacles for the flowing may cause ditches to become ineffective. Establishing of peat-forming vegetation is a crucial phenomenon. Various stages of a discontinuous regeneration progress can be differentiated and allocated to the processes that are known so far. Frequency and the area ratio of initial and progressed regeneration suggest that regeneration processes take place on a larger area, but that they are not recognised. Regeneration leads to site drifting and the distribution of peatland species. The regeneration potential is given by abiotic conditions. Irreversible changes of the hydro-morphological structure due to drainage or peat cutting are limiting factors for the regeneration. Anthropogenic input of matter and lack of key species bring about other restrictions. - If a profitable timber production is aimed at, autogenously regenerated peatlands are sites difficult to manage or that cannot be managed at all, involving a high production risk due to site drifting and partly inadequacy for woody plants. For nature conservation, regeneration processes are a chance and an important asset worth of protection, as they lead to high-grade peatland habitats. In the case of advanced regeneration cost-intensive operations are not necessary. A protection of static nature is inadequate for the process character. Initial stages and zones not yet under regeneration, implying, however, a high potential, can be promoted efficiently. Heavy degeneration of the peatlands in the study area, given a low proportion of actual regeneration zones, justifies a strong call for action, including a shielding protection as well as efficient control mechanisms. - An appropriate data basis for localisation of areas, having a high regeneration potential, is largely missing. Remedying this deficit is possible.

I. An original methodological discussion is proposed on the problem of the typology of tropical rain forest’s plant communities, based on the study of forest types across gradients of continentality and elevation, within Atlantic central Africa. These investigations were based on the statement that the main problems in forest typology are related to the non-zonal or zonal character of the different vegetation types and to non considering the relations and differences between forest strata. II. Field data consisted in phytosociological homogeneous sample plots localized within different recognized phytogeographical entities, in a region of tropical Africa where these entities are known to be well conserved. A total of 37 such plots were inventoried in the region extending from the littoral forests of Ndoté, Equatorial Guinea, which are wet evergreen forests, to the continental forests of the Dja, Cameroon, known as evergreen seasonal forests. The studied region also included the oriental Atlantic forests of Equatorial Guinea, known as moist evergreen forests or caesalp forests. In various parts of this continentality gradient, some plots were localized within climax non-zonal formations, namely the submontane rain forests. The emphasis was put on the vegetation of the Monte Alén National Park. The sampling methodology was willing to be as "complete ", including all strata, "quantitative ", enumerating all individuals, and "representative ", within each stratum, as possible. These multi-layers plots were realised using nested sub-plots, with a sampling size of 100 individuals for every ligneous stratum recognized (dominant trees, dominated trees and shrubs) and a sampling size of 200m² for the herbaceous and suffrutex stratum. Forest types were defined independently for each stratum and the differences were analysed. A method was proposed for the simultaneous analysis of all floristic data, converting and standardizing the values from ligneous strata, on the one hand, and from understorey strata, on the other hand. III. Ten forest types were described using IndVal and discussed in the general context of the guineo-congolian region, from a syntaxonomic view point (agglomerative classification) and from a phytogeographical view point (divisive classification). Homologies between these two approaches are described. The proposed phytogeographical system is based on an "open " conception of hierarchical classifications, combining advantages of agglomerative and divisive classifications. In concrete terms, the non-zonal criteria, for example the submontane variants, are categorised separately and in analogy with the zonal criteria, related to the usual phytochoria. Analysis of ecological relationships for the 10 communities showed that the main variables related to the floristic variability in our mainland rain forests are elevation, rainfall, hygrometry (estimated using bryophytes cover levels) and distance to the ocean. The two extremes on the vertical microclimatic gradient, dominant trees stratum and herbaceous stratum, give similar typologies, however canonical analysis showed that for the herbaceous layer, non-zonal variables (hygrometry and elevation) were gaining more importance when the influence of the two zonal variables was attenuated. In every case, spatial autocorrelation was less important than the environment in explaining floristic variability but its role increased in the spatial arrangement of understorey species, whose dispersal capacity is generally lower than canopy trees. The phytosociological, phytogeographical and ecological description of forest types is accompanied by a physiognomical description using biological types spectrum, as well as architectural models, leaf sizes, etc. With regard to diversity, we have demonstrated that species richness was higher from upper to lower strata because of the accumulation in lower strata of species from various strata. On the other hand, the proper stratum diversity, i.e. the structural set, decreased from dominant trees to shrubs. The proper diversity of the herb layer showed relatively high figures mainly due to the higher individual density in relation to the existence of microstrata. Within the 37 sample plots, 1,050 taxa have been identified to species or morpho-species levels, for a total of 25,750 individuals. These taxa represent 442 genus among 104 families. The richest forest type is found on the foothills of the Niefang range, on the windward side. This forest type is also characterised by a high number of oligotypic genus and by species belonging to functional types indicators of glacial refuges. These functional types are defined on the basis of the dispersal capacity and on kind of stand needed for effective germination. We formulated the hypothesis that this kind of "foothills refuge ", characterised by his zonal nature, could have been one of the rare refuges for species from mainland rain forests, while montane and fluvial refuges would mainly have preserved species from non-zonal forest types: (sub)montane and riverine. Based on indicator species of submontane forests, a potential distribution map of this forest type has been realised at the Atlantic central African scale. More than 400 submontane forest localities have been mapped. These forests begin at 400m of altitude near the ocean, and progressively at higher altitude for increasing distance to the ocean. Many lowland localities also comprised submontane species, which could indicate the existence of ecological transgressions. These transgressions would allow migratory tracks for submontane species between isolated mountain ranges, not only during glacial periods, through heights at the northern and southern borders of the congo basin, but also contemporarily through the lowland riverine forest network, in the centre of this basin. Finally, a special attention has been attributed to littoral forests and to some cases of choroecological transgressions, coupled to the ecological equalization phenomenon.

I. An original methodological discussion is proposed on the problem of the typology of tropical rain forest’s plant communities, based on the study of forest types across gradients of continentality and elevation, within Atlantic central Africa. These investigations were based on the statement that the main problems in forest typology are related to the non-zonal or zonal character of the different vegetation types and to non considering the relations and differences between forest strata.<p><p>II. Field data consisted in phytosociological homogeneous sample plots localized within different recognized phytogeographical entities, in a region of tropical Africa where these entities are known to be well conserved. A total of 37 such plots were inventoried in the region extending from the littoral forests of Ndoté, Equatorial Guinea, which are wet evergreen forests, to the continental forests of the Dja, Cameroon, known as evergreen seasonal forests. The studied region also included the oriental Atlantic forests of Equatorial Guinea, known as moist evergreen forests or caesalp forests. In various parts of this continentality gradient, some plots were localized within climax non-zonal formations, namely the submontane rain forests. The emphasis was put on the vegetation of the Monte Alén National Park.<p><p>The sampling methodology was willing to be as "complete ", including all strata, "quantitative ", enumerating all individuals, and "representative ", within each stratum, as possible. These multi-layers plots were realised using nested sub-plots, with a sampling size of 100 individuals for every ligneous stratum recognized (dominant trees, dominated trees and shrubs) and a sampling size of 200m² for the herbaceous and suffrutex stratum.<p><p>Forest types were defined independently for each stratum and the differences were analysed. A method was proposed for the simultaneous analysis of all floristic data, converting and standardizing the values from ligneous strata, on the one hand, and from understorey strata, on the other hand.<p><p>III. Ten forest types were described using IndVal and discussed in the general context of the guineo-congolian region, from a syntaxonomic view point (agglomerative classification) and from a phytogeographical view point (divisive classification). Homologies between these two approaches are described. The proposed phytogeographical system is based on an "open " conception of hierarchical classifications, combining advantages of agglomerative and divisive classifications. In concrete terms, the non-zonal criteria, for example the submontane variants, are categorised separately and in analogy with the zonal criteria, related to the usual phytochoria.<p><p>Analysis of ecological relationships for the 10 communities showed that the main variables related to the floristic variability in our mainland rain forests are elevation, rainfall, hygrometry (estimated using bryophytes cover levels) and distance to the ocean. The two extremes on the vertical microclimatic gradient, dominant trees stratum and herbaceous stratum, give similar typologies, however canonical analysis showed that for the herbaceous layer, non-zonal variables (hygrometry and elevation) were gaining more importance when the influence of the two zonal variables was attenuated. In every case, spatial autocorrelation was less important than the environment in explaining floristic variability but its role increased in the spatial arrangement of understorey species, whose dispersal capacity is generally lower than canopy trees. The phytosociological, phytogeographical and ecological description of forest types is accompanied by a physiognomical description using biological types spectrum, as well as architectural models, leaf sizes, etc.<p><p>With regard to diversity, we have demonstrated that species richness was higher from upper to lower strata because of the accumulation in lower strata of species from various strata. On the other hand, the proper stratum diversity, i.e. the structural set, decreased from dominant trees to shrubs. The proper diversity of the herb layer showed relatively high figures mainly due to the higher individual density in relation to the existence of microstrata. Within the 37 sample plots, 1,050 taxa have been identified to species or morpho-species levels, for a total of 25,750 individuals. These taxa represent 442 genus among 104 families. The richest forest type is found on the foothills of the Niefang range, on the windward side. This forest type is also characterised by a high number of oligotypic genus and by species belonging to functional types indicators of glacial refuges. These functional types are defined on the basis of the dispersal capacity and on kind of stand needed for effective germination. We formulated the hypothesis that this kind of "foothills refuge ", characterised by his zonal nature, could have been one of the rare refuges for species from mainland rain forests, while montane and fluvial refuges would mainly have preserved species from non-zonal forest types: (sub)montane and riverine.<p><p>Based on indicator species of submontane forests, a potential distribution map of this forest type has been realised at the Atlantic central African scale. More than 400 submontane forest localities have been mapped. These forests begin at 400m of altitude near the ocean, and progressively at higher altitude for increasing distance to the ocean. Many lowland localities also comprised submontane species, which could indicate the existence of ecological transgressions. These transgressions would allow migratory tracks for submontane species between isolated mountain ranges, not only during glacial periods, through heights at the northern and southern borders of the congo basin, but also contemporarily through the lowland riverine forest network, in the centre of this basin. Finally, a special attention has been attributed to littoral forests and to some cases of choroecological transgressions, coupled to the ecological equalization phenomenon.<p> / Doctorat en sciences agronomiques et ingénierie biologique / info:eu-repo/semantics/nonPublished